Prokaryotic Genomes and Diversity in Surface Ocean Waters: Interrogating the Global Ocean Sampling Metagenome▿ †
Identifieur interne : 000202 ( Pmc/Curation ); précédent : 000201; suivant : 000203Prokaryotic Genomes and Diversity in Surface Ocean Waters: Interrogating the Global Ocean Sampling Metagenome▿ †
Auteurs : Erin J. Biers ; Shulei Sun ; Erinn C. HowardSource :
- Applied and Environmental Microbiology [ 0099-2240 ] ; 2009.
Abstract
The
Url:
DOI: 10.1128/AEM.02118-08
PubMed: 19201952
PubMed Central: 2663191
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<record><TEI><teiHeader><fileDesc><titleStmt><title xml:lang="en">Prokaryotic Genomes and Diversity in Surface Ocean Waters: Interrogating the Global Ocean Sampling Metagenome<xref ref-type="fn" rid="fn3">▿</xref>
<xref ref-type="fn" rid="fn2">†</xref>
</title>
<author><name sortKey="Biers, Erin J" sort="Biers, Erin J" uniqKey="Biers E" first="Erin J." last="Biers">Erin J. Biers</name>
<affiliation><nlm:aff id="aff1"></nlm:aff>
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<author><name sortKey="Sun, Shulei" sort="Sun, Shulei" uniqKey="Sun S" first="Shulei" last="Sun">Shulei Sun</name>
<affiliation><nlm:aff id="aff1"></nlm:aff>
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<author><name sortKey="Howard, Erinn C" sort="Howard, Erinn C" uniqKey="Howard E" first="Erinn C." last="Howard">Erinn C. Howard</name>
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<date when="2009">2009</date>
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<sourceDesc><biblStruct><analytic><title xml:lang="en" level="a" type="main">Prokaryotic Genomes and Diversity in Surface Ocean Waters: Interrogating the Global Ocean Sampling Metagenome<xref ref-type="fn" rid="fn3">▿</xref>
<xref ref-type="fn" rid="fn2">†</xref>
</title>
<author><name sortKey="Biers, Erin J" sort="Biers, Erin J" uniqKey="Biers E" first="Erin J." last="Biers">Erin J. Biers</name>
<affiliation><nlm:aff id="aff1"></nlm:aff>
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<author><name sortKey="Sun, Shulei" sort="Sun, Shulei" uniqKey="Sun S" first="Shulei" last="Sun">Shulei Sun</name>
<affiliation><nlm:aff id="aff1"></nlm:aff>
</affiliation>
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<author><name sortKey="Howard, Erinn C" sort="Howard, Erinn C" uniqKey="Howard E" first="Erinn C." last="Howard">Erinn C. Howard</name>
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</affiliation>
</author>
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<series><title level="j">Applied and Environmental Microbiology</title>
<idno type="ISSN">0099-2240</idno>
<idno type="eISSN">1098-5336</idno>
<imprint><date when="2009">2009</date>
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<front><div type="abstract" xml:lang="en"><p>The <italic>Sorcerer II</italic>
Global Ocean Sampling (GOS) sequencing effort has vastly expanded the landscape of metagenomics, providing an opportunity to study the genetic potential of surface ocean water bacterioplankton on a global scale. Here we describe the habitat-based microbial diversity, both taxon evenness and taxon richness, for each GOS site and estimate genome characteristics of a typical free-living, surface ocean water bacterium. While <italic>Alphaproteobacteria</italic>
and particularly SAR11 dominate the 0.1- to 0.8-μm size fraction of surface ocean water bacteria (43% and 31%, respectively), the proportions of other taxa varied with ocean habitat type. Within each habitat type, lower-bound estimates of phylum richness ranged between 18 and 59 operational taxonomic units (OTUs). However, OTU richness was relatively low in the hypersaline lagoon community at every taxonomic level, and open-ocean communities had much more microdiversity than any other habitat. Based on the abundance of single-copy eubacterial genes from the same data set, we estimate that the genome of an average free-living surface ocean water bacterium (sized between 0.1 and 0.8 μm) contains ∼1,019 genes and 1.8 copies of the 16S rRNA gene, suggesting that these bacteria have relatively streamlined genomes in comparison to those of cultured bacteria and bacteria from other habitats (e.g., soil or acid mine drainage).</p>
</div>
</front>
</TEI>
<pmc article-type="research-article"><pmc-comment>The publisher of this article does not allow downloading of the full text in XML form.</pmc-comment>
<front><journal-meta><journal-id journal-id-type="nlm-ta">Appl Environ Microbiol</journal-id>
<journal-id journal-id-type="publisher-id">aem</journal-id>
<journal-title>Applied and Environmental Microbiology</journal-title>
<issn pub-type="ppub">0099-2240</issn>
<issn pub-type="epub">1098-5336</issn>
<publisher><publisher-name>American Society for Microbiology (ASM)</publisher-name>
</publisher>
</journal-meta>
<article-meta><article-id pub-id-type="pmid">19201952</article-id>
<article-id pub-id-type="pmc">2663191</article-id>
<article-id pub-id-type="publisher-id">2118-08</article-id>
<article-id pub-id-type="doi">10.1128/AEM.02118-08</article-id>
<article-categories><subj-group subj-group-type="heading"><subject>Evolutionary and Genomic Microbiology</subject>
</subj-group>
</article-categories>
<title-group><article-title>Prokaryotic Genomes and Diversity in Surface Ocean Waters: Interrogating the Global Ocean Sampling Metagenome<xref ref-type="fn" rid="fn3">▿</xref>
<xref ref-type="fn" rid="fn2">†</xref>
</article-title>
</title-group>
<contrib-group><contrib contrib-type="author"><name><surname>Biers</surname>
<given-names>Erin J.</given-names>
</name>
<xref ref-type="aff" rid="aff1">1</xref>
<xref ref-type="corresp" rid="cor1">*</xref>
</contrib>
<contrib contrib-type="author"><name><surname>Sun</surname>
<given-names>Shulei</given-names>
</name>
<xref ref-type="aff" rid="aff1">1</xref>
</contrib>
<contrib contrib-type="author"><name><surname>Howard</surname>
<given-names>Erinn C.</given-names>
</name>
<xref ref-type="aff" rid="aff1">2</xref>
<xref ref-type="fn" rid="fn1">§</xref>
</contrib>
</contrib-group>
<aff id="aff1">Department of Marine Sciences, University of Georgia, Athens, Georgia 30602,<label>1</label>
Department of Microbiology, University of Georgia, Athens, Georgia 30602<label>2</label>
</aff>
<author-notes><fn id="cor1"><label>*</label>
<p>Corresponding author. Present address: Department of Environmental Health Sciences, University of South Carolina, Columbia, SC 29208. Phone: (803) 777-4553. Fax: (803) 777-3391. E-mail: <email>ejbiers@mailbox.sc.edu</email>
</p>
</fn>
<fn id="fn1"><label>§</label>
<p>Present address: Department of Marine Sciences, University of Georgia, Athens, GA 30602.</p>
</fn>
</author-notes>
<pub-date pub-type="ppub"><month>4</month>
<year>2009</year>
</pub-date>
<pub-date pub-type="epub"><day>6</day>
<month>2</month>
<year>2009</year>
</pub-date>
<volume>75</volume>
<issue>7</issue>
<fpage>2221</fpage>
<lpage>2229</lpage>
<history><date date-type="received"><day>12</day>
<month>9</month>
<year>2008</year>
</date>
<date date-type="accepted"><day>2</day>
<month>2</month>
<year>2009</year>
</date>
</history>
<permissions><copyright-statement>Copyright © 2009, American Society for Microbiology</copyright-statement>
</permissions>
<self-uri xlink:title="pdf" xlink:href="zam00709002221.pdf"></self-uri>
<abstract><p>The <italic>Sorcerer II</italic>
Global Ocean Sampling (GOS) sequencing effort has vastly expanded the landscape of metagenomics, providing an opportunity to study the genetic potential of surface ocean water bacterioplankton on a global scale. Here we describe the habitat-based microbial diversity, both taxon evenness and taxon richness, for each GOS site and estimate genome characteristics of a typical free-living, surface ocean water bacterium. While <italic>Alphaproteobacteria</italic>
and particularly SAR11 dominate the 0.1- to 0.8-μm size fraction of surface ocean water bacteria (43% and 31%, respectively), the proportions of other taxa varied with ocean habitat type. Within each habitat type, lower-bound estimates of phylum richness ranged between 18 and 59 operational taxonomic units (OTUs). However, OTU richness was relatively low in the hypersaline lagoon community at every taxonomic level, and open-ocean communities had much more microdiversity than any other habitat. Based on the abundance of single-copy eubacterial genes from the same data set, we estimate that the genome of an average free-living surface ocean water bacterium (sized between 0.1 and 0.8 μm) contains ∼1,019 genes and 1.8 copies of the 16S rRNA gene, suggesting that these bacteria have relatively streamlined genomes in comparison to those of cultured bacteria and bacteria from other habitats (e.g., soil or acid mine drainage).</p>
</abstract>
</article-meta>
</front>
<floats-wrap><fig position="float" id="f1"><label>FIG. 1.</label>
<caption><p>Microbial diversity (16S rRNA) for all GOS samples. Dotted lines connect taxonomic data to sample locations (filled circles). Numbers within taxonomic plots indicate the sample identification numbers (preceded by “GS” as described by Rusch et al. [<xref ref-type="bibr" rid="r41">41</xref>
]). The border color surrounding each taxonomic plot indicates the habitat type. The habitats classified as “Other” include the following: 5, embayment; 16, coastal sea; 20, fresh water; 25, fringing reef; 30, warm seep; 31, coastal upwelling; 32, mangrove; 51, coral reef atoll. All samples were collected from the 0.1- to 0.8-μm size fraction, unless indicated otherwise by the following symbols: *, from the 0.22- to 0.8-μm size fraction; †, from the 3- to 20-μm size fraction; and ‡, from the 0.8- to 3-μm size fraction.</p>
</caption>
<graphic xlink:href="zam0070997880001"></graphic>
</fig>
<fig position="float" id="f2"><label>FIG. 2.</label>
<caption><p>Diversity, evenness, and richness indices for GOS habitats, as calculated by DOTUR. (A) Diversity indices were estimated using the Shannon-Weaver index of diversity. (B) Evenness scores were estimated from Shannon-Weaver indices of diversity as well as richness. (C) Lower bounds of richness were calculated using the Chao1 estimator. Error bars indicate 95% confidence intervals.</p>
</caption>
<graphic xlink:href="zam0070997880002"></graphic>
</fig>
<table-wrap position="float" id="t1"><label>TABLE 1.</label>
<caption><p>Ribotype diversity for habitats sampled multiple times by GOS<xref ref-type="table-fn" rid="t1fn1"><italic>a</italic>
</xref>
</p>
</caption>
<table frame="hsides" rules="groups"><thead><tr><th colspan="1" rowspan="2" align="center" valign="middle">Classification</th>
<th colspan="2" rowspan="1" align="center" valign="bottom">Coastal habitat
<hr></hr>
</th>
<th colspan="2" rowspan="1" align="center" valign="bottom">Estuary habitat
<hr></hr>
</th>
<th colspan="2" rowspan="1" align="center" valign="bottom">Open-ocean habitat
<hr></hr>
</th>
<th colspan="2" rowspan="1" align="center" valign="bottom">Entire GOS metagenome
<hr></hr>
</th>
</tr>
<tr><th colspan="1" rowspan="1" align="center" valign="bottom">No. of hits</th>
<th colspan="1" rowspan="1" align="center" valign="bottom">% of hits</th>
<th colspan="1" rowspan="1" align="center" valign="bottom">No. of hits</th>
<th colspan="1" rowspan="1" align="center" valign="bottom">% of hits</th>
<th colspan="1" rowspan="1" align="center" valign="bottom">No. of hits</th>
<th colspan="1" rowspan="1" align="center" valign="bottom">% of hits</th>
<th colspan="1" rowspan="1" align="center" valign="bottom">No. of hits</th>
<th colspan="1" rowspan="1" align="center" valign="bottom">% of hits</th>
</tr>
</thead>
<tbody><tr><td colspan="1" rowspan="1" align="left" valign="top"><italic>Archaea</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">14</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.4</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">8</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.2</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">32</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.3</bold>
</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"><italic>Bacteria</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Actinobacteria</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">230</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>7.3</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">92</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>23.8</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">188</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>5.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">824</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>8.2</bold>
</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> CFB</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">252</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>8.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">28</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>7.3</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">126</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>3.3</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">603</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>6.0</bold>
</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Chlamydiae</italic>
/<italic>Verrucomicrobia</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">12</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.4</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">4</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>1.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">9</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.2</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">41</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.4</bold>
</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Chloroflexi</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.1</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.8</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.1</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.2</bold>
</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> Unclassified</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">2</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">3</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.8</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">15</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.1</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> SAR202</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Cyanobacteria</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">167</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>5.3</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">255</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>6.7</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">532</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>5.3</bold>
</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Deinococcus</italic>
/<italic>Thermus</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">4</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Fibrobacteres</italic>
/<italic>Acidobacteria</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">6</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.2</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">3</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.1</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">33</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.3</bold>
</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Firmicutes</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">4</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.1</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.3</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">5</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.1</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">22</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.2</bold>
</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> OP11</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> OD1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">2</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> SR1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Planctomycetales</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">2</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.5</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">4</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.1</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">19</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.2</bold>
</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Alphaproteobacteria</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>51.3</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>36.8</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>40.8</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>43.1</bold>
</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> Unclassified</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">128</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">4.1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">8</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">2.1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">162</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">4.3</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">410</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">4.1</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Caulobacterales</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">2</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">2</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">7</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.1</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Rhizobiales</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">26</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.8</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">2</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.5</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">13</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.3</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">91</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.9</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Rhodobacterales</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">39</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">1.2</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">35</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.9</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">122</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">1.2</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Rhodobacterales</italic>
, <italic>Roseobacter</italic>
clade</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">118</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">3.8</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.3</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">40</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">1.1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">258</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">2.6</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Rhodospirillales</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">13</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.4</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.3</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">27</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.7</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">54</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.5</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Rickettsiales</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Rickettsiales</italic>
, SAR11 cluster</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">1,149</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">36.7</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">127</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">32.9</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">1,178</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">31.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">3,093</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">30.9</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> SAR116 cluster</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">130</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">4.1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">3</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.8</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">89</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">2.3</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">275</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">2.7</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Sphingomonadales</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">3</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">3</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">10</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.1</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Betaproteobacteria</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.9</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>11.1</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>5.2</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>2.8</bold>
</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> Unclassified</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">8</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.3</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">9</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">2.3</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">23</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.2</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Burkholderiales</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">20</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.6</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">34</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">8.8</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">195</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">5.1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">260</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">2.6</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Deltaproteobacteria</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.2</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.8</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.4</bold>
</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> Unclassified</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">2</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Bdellovibrionales</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">2</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Desulfobacterales</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Desulfovibrionales</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Desulfuromonadales</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Myxococcales</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> SAR324</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">5</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.2</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">27</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.7</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">38</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.4</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Syntrophobacterales</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Epsilonproteobacteria</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.1</bold>
</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> Unclassified</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Campylobacterales</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">4</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.1</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Sulfurovum</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">2</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Gammaproteobacteria</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>14.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>4.9</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>21.2</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>17.3</bold>
</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> Unclassified</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">140</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">4.5</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">2</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.5</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">196</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">5.2</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">562</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">5.6</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Aeromonadales</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">3</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">16</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.4</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">19</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.2</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Alteromonadales</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">17</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.5</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.3</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">249</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">6.6</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">290</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">2.9</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Chromatiales</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">3</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">5</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Enterobacteriales</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">2</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">4</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">6</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.1</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Methylococcales</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">5</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.2</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.3</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">7</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.2</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">15</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.1</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Nitrincola</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">6</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.2</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">11</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.3</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">38</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.4</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Oceanospirillales</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">9</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.3</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">15</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.4</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">47</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.5</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Pasteurellales</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Pseudomonadales</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">18</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.6</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">2</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.5</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">27</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.7</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">67</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.7</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> SAR86 cluster</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">231</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">7.4</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">12</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">3.1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">269</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">7.1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">636</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">6.3</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Thiotrichales</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.3</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">8</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.1</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Vibrionales</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">7</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.2</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">7</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.2</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">40</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.4</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> <italic>Xanthomonadales</italic>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">1</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">5</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0.0</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> SAR406</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">32</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>1.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">66</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>1.7</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">106</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>1.1</bold>
</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> Spirochaetes</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top"> Unclassified</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">20</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.6</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">43</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>1.1</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">67</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.7</bold>
</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top">Eukaryotes</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">9</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.3</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">0</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">15</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.4</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">28</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>0.3</bold>
</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top">Unclassified at >90%</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">303</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>9.7</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">52</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>13.5</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">494</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>13.0</bold>
</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">1,304</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"><bold>13.0</bold>
</td>
</tr>
<tr><td colspan="1" rowspan="1" align="left" valign="top">Total</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">3,133</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">386</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">3,794</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
<td colspan="1" rowspan="1" align="char" char="." valign="top">10,025</td>
<td colspan="1" rowspan="1" align="char" char="." valign="top"></td>
</tr>
</tbody>
</table>
<table-wrap-foot><fn id="t1fn1"><label>a</label>
<p>Coastal habitats were sampled 18 times, estuary habitats 3 times, and open-ocean habitats 14 times. Ribotype diversity for individual samples is provided in Table S3 in the supplemental material. The number of hits listed for each taxonomic bin is the raw, unnormalized count after duplicate reads (paired-end reads) were removed. Numbers in bold are total percentages of hits that were binned into the indicated domain/phylum/clade.</p>
</fn>
</table-wrap-foot>
</table-wrap>
</floats-wrap>
</pmc>
</record>
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