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<record><TEI><teiHeader><fileDesc><titleStmt><title xml:lang="en">A new species of <italic>Aeneator</italic>
Finlay, 1926 (Mollusca, Gastropoda, Buccinidae) from northern Chile, with comments on the genus and a key to the Chilean species</title>
<author><name sortKey="Araya, Juan Francisco" sort="Araya, Juan Francisco" uniqKey="Araya J" first="Juan Francisco" last="Araya">Juan Francisco Araya</name>
<affiliation><nlm:aff id="A1">Universidad de Chile, Facultad de Ciencias, Las Palmeras 3425. Ñuñoa. Santiago. Chile</nlm:aff>
</affiliation>
</author>
</titleStmt>
<publicationStmt><idno type="wicri:source">PMC</idno>
<idno type="pmid">23653495</idno>
<idno type="pmc">3591740</idno>
<idno type="url">http://www.ncbi.nlm.nih.gov/pmc/articles/PMC3591740</idno>
<idno type="RBID">PMC:3591740</idno>
<idno type="doi">10.3897/zookeys.257.4446</idno>
<date when="2013">2013</date>
<idno type="wicri:Area/Pmc/Corpus">000441</idno>
</publicationStmt>
<sourceDesc><biblStruct><analytic><title xml:lang="en" level="a" type="main">A new species of <italic>Aeneator</italic>
Finlay, 1926 (Mollusca, Gastropoda, Buccinidae) from northern Chile, with comments on the genus and a key to the Chilean species</title>
<author><name sortKey="Araya, Juan Francisco" sort="Araya, Juan Francisco" uniqKey="Araya J" first="Juan Francisco" last="Araya">Juan Francisco Araya</name>
<affiliation><nlm:aff id="A1">Universidad de Chile, Facultad de Ciencias, Las Palmeras 3425. Ñuñoa. Santiago. Chile</nlm:aff>
</affiliation>
</author>
</analytic>
<series><title level="j">ZooKeys</title>
<idno type="ISSN">1313-2989</idno>
<idno type="eISSN">1313-2970</idno>
<imprint><date when="2013">2013</date>
</imprint>
</series>
</biblStruct>
</sourceDesc>
</fileDesc>
<profileDesc><textClass></textClass>
</profileDesc>
</teiHeader>
<front><div type="abstract" xml:lang="en"><label>Abstract</label>
<p>A new species of the genus <italic><named-content content-type="taxon-name">Aeneator</named-content>
</italic>
Finlay, 1926 is described from off the coast of Caldera (27°S), northern Chile. <italic><named-content content-type="taxon-name">Aeneator martae</named-content>
</italic>
sp. n. has a small, broad, stout, angulated shell with more prominent axial ribs and a more obviously keeled periphery than all previously named Chilean species. Comparisons are provided with all other South American named species of <italic><named-content content-type="taxon-name">Aeneator</named-content>
</italic>
.</p>
</div>
</front>
<back><div1 type="bibliography"><listBibl><biblStruct><analytic><author><name sortKey="Beu, Ag" uniqKey="Beu A">AG Beu</name>
</author>
<author><name sortKey="Marshall, Ba" uniqKey="Marshall B">BA Marshall</name>
</author>
</analytic>
</biblStruct>
<biblStruct><analytic><author><name sortKey="Beu, Ag" uniqKey="Beu A">AG Beu</name>
</author>
<author><name sortKey="Marshall, Ba" uniqKey="Marshall B">BA Marshall</name>
</author>
</analytic>
</biblStruct>
<biblStruct><analytic><author><name sortKey="Beu, Ag" uniqKey="Beu A">AG Beu</name>
</author>
<author><name sortKey="Maxwell, Pa" uniqKey="Maxwell P">PA Maxwell</name>
</author>
</analytic>
</biblStruct>
<biblStruct><analytic><author><name sortKey="Beu, Ag" uniqKey="Beu A">AG Beu</name>
</author>
</analytic>
</biblStruct>
<biblStruct><analytic><author><name sortKey="Chiu, Yw" uniqKey="Chiu Y">YW Chiu</name>
</author>
<author><name sortKey="Chen, Hc" uniqKey="Chen H">HC Chen</name>
</author>
<author><name sortKey="Lee, Sc" uniqKey="Lee S">SC Lee</name>
</author>
<author><name sortKey="Chen, Ca" uniqKey="Chen C">CA Chen</name>
</author>
</analytic>
</biblStruct>
<biblStruct><analytic><author><name sortKey="Dawson, Ew" uniqKey="Dawson E">EW Dawson</name>
</author>
</analytic>
</biblStruct>
<biblStruct><analytic><author><name sortKey="Depraz, A" uniqKey="Depraz A">A Dépraz</name>
</author>
<author><name sortKey="Hausser, J" uniqKey="Hausser J">J Hausser</name>
</author>
<author><name sortKey="Pfenninger, M" uniqKey="Pfenninger M">M Pfenninger</name>
</author>
</analytic>
</biblStruct>
<biblStruct><analytic><author><name sortKey="Finlay, Hj" uniqKey="Finlay H">HJ Finlay</name>
</author>
</analytic>
</biblStruct>
<biblStruct><analytic><author><name sortKey="Fraussen, K" uniqKey="Fraussen K">K Fraussen</name>
</author>
<author><name sortKey="Sellanes, J" uniqKey="Sellanes J">J Sellanes</name>
</author>
</analytic>
</biblStruct>
<biblStruct><analytic><author><name sortKey="Guzman, N" uniqKey="Guzman N">N Guzmán</name>
</author>
<author><name sortKey="Saa, S" uniqKey="Saa S">S Saá</name>
</author>
<author><name sortKey="Ortlieb, L" uniqKey="Ortlieb L">L Ortlieb</name>
</author>
</analytic>
</biblStruct>
<biblStruct><analytic><author><name sortKey="Laudien, J" uniqKey="Laudien J">J Laudien</name>
</author>
<author><name sortKey="Rojo, Me" uniqKey="Rojo M">ME Rojo</name>
</author>
<author><name sortKey="Oliva, Me" uniqKey="Oliva M">ME Oliva</name>
</author>
<author><name sortKey="Arntz, We" uniqKey="Arntz W">WE Arntz</name>
</author>
<author><name sortKey="Thatje, S" uniqKey="Thatje S">S Thatje</name>
</author>
</analytic>
</biblStruct>
<biblStruct><analytic><author><name sortKey="Marwick, J" uniqKey="Marwick J">J Marwick</name>
</author>
</analytic>
</biblStruct>
<biblStruct><analytic><author><name sortKey="Mclean, Jh" uniqKey="Mclean J">JH McLean</name>
</author>
<author><name sortKey="Andrade, H" uniqKey="Andrade H">H Andrade</name>
</author>
</analytic>
</biblStruct>
<biblStruct><analytic><author><name sortKey="Nielsen, S" uniqKey="Nielsen S">S Nielsen</name>
</author>
</analytic>
</biblStruct>
<biblStruct><analytic><author><name sortKey="Osorio, C" uniqKey="Osorio C">C Osorio</name>
</author>
<author><name sortKey="Pe A, R" uniqKey="Pe A R">R Peña</name>
</author>
<author><name sortKey="Ramajo, L" uniqKey="Ramajo L">L Ramajo</name>
</author>
<author><name sortKey="Garcelon, N" uniqKey="Garcelon N">N Garcelon</name>
</author>
</analytic>
</biblStruct>
<biblStruct><analytic><author><name sortKey="Parraga, D" uniqKey="Parraga D">D Párraga</name>
</author>
<author><name sortKey="Wiff, R" uniqKey="Wiff R">R Wiff</name>
</author>
<author><name sortKey="Quiroz, Jc" uniqKey="Quiroz J">JC Quiroz</name>
</author>
<author><name sortKey="Zilleruelo, M" uniqKey="Zilleruelo M">M Zilleruelo</name>
</author>
<author><name sortKey="Bernal, C" uniqKey="Bernal C">C Bernal</name>
</author>
<author><name sortKey="Az Car, J" uniqKey="Az Car J">J Azócar</name>
</author>
</analytic>
</biblStruct>
<biblStruct><analytic><author><name sortKey="Powell, Awb" uniqKey="Powell A">AWB Powell</name>
</author>
</analytic>
</biblStruct>
<biblStruct><analytic><author><name sortKey="Queirolo, D" uniqKey="Queirolo D">D Queirolo</name>
</author>
<author><name sortKey="Erzini, K" uniqKey="Erzini K">K Erzini</name>
</author>
<author><name sortKey="Hurtado, C" uniqKey="Hurtado C">C Hurtado</name>
</author>
<author><name sortKey="Gaete, E" uniqKey="Gaete E">E Gaete</name>
</author>
<author><name sortKey="Soriguer, M" uniqKey="Soriguer M">M Soriguer</name>
</author>
</analytic>
</biblStruct>
<biblStruct><analytic><author><name sortKey="Rehder, Ha" uniqKey="Rehder H">HA Rehder</name>
</author>
</analytic>
</biblStruct>
</listBibl>
</div1>
</back>
</TEI>
<pmc article-type="research-article"><pmc-dir>properties open_access</pmc-dir>
<front><journal-meta><journal-id journal-id-type="nlm-ta">Zookeys</journal-id>
<journal-id journal-id-type="iso-abbrev">Zookeys</journal-id>
<journal-id journal-id-type="publisher-id">ZooKeys</journal-id>
<journal-title-group><journal-title>ZooKeys</journal-title>
</journal-title-group>
<issn pub-type="ppub">1313-2989</issn>
<issn pub-type="epub">1313-2970</issn>
<publisher><publisher-name>Pensoft Publishers</publisher-name>
</publisher>
</journal-meta>
<article-meta><article-id pub-id-type="pmid">23653495</article-id>
<article-id pub-id-type="pmc">3591740</article-id>
<article-id pub-id-type="doi">10.3897/zookeys.257.4446</article-id>
<article-categories><subj-group subj-group-type="heading"><subject>Article</subject>
</subj-group>
</article-categories>
<title-group><article-title>A new species of <italic>Aeneator</italic>
Finlay, 1926 (Mollusca, Gastropoda, Buccinidae) from northern Chile, with comments on the genus and a key to the Chilean species</article-title>
</title-group>
<contrib-group><contrib contrib-type="author"><name><surname>Araya</surname>
<given-names>Juan Francisco</given-names>
</name>
<xref ref-type="aff" rid="A1">1</xref>
<uri content-type="lsid" xlink:type="simple">urn:lsid:zoobank.org:author:443B4F42-FB13-42A6-B92B-1B0F835698A9</uri>
</contrib>
</contrib-group>
<aff id="A1"><label>1</label>
Universidad de Chile, Facultad de Ciencias, Las Palmeras 3425. Ñuñoa. Santiago. Chile</aff>
<author-notes><corresp>Corresponding author: Juan Francisco Araya (<email xlink:type="simple">jfaraya@u.uchile.cl</email>
)</corresp>
<fn fn-type="edited-by"><p>Academic editor: Nathalie Yonow</p>
</fn>
</author-notes>
<pub-date pub-type="collection"><year>2013</year>
</pub-date>
<pub-date pub-type="epub"><day>14</day>
<month>1</month>
<year>2013</year>
</pub-date>
<issue>257</issue>
<fpage>89</fpage>
<lpage>101</lpage>
<history><date date-type="received"><day>3</day>
<month>12</month>
<year>2012</year>
</date>
<date date-type="accepted"><day>27</day>
<month>12</month>
<year>2012</year>
</date>
</history>
<permissions><copyright-statement>Juan Francisco Araya</copyright-statement>
<license license-type="creative-commons-attribution" xlink:href="http://creativecommons.org/licenses/by/3.0"><license-p>This is an open access article distributed under the terms of the Creative Commons Attribution License 3.0 (CC-BY), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.</license-p>
</license>
</permissions>
<self-uri content-type="lsid" xlink:type="simple">urn:lsid:zoobank.org:pub:13316CEA-0B85-4883-BBC6-503B99831EF8</self-uri>
<abstract><label>Abstract</label>
<p>A new species of the genus <italic><named-content content-type="taxon-name">Aeneator</named-content>
</italic>
Finlay, 1926 is described from off the coast of Caldera (27°S), northern Chile. <italic><named-content content-type="taxon-name">Aeneator martae</named-content>
</italic>
sp. n. has a small, broad, stout, angulated shell with more prominent axial ribs and a more obviously keeled periphery than all previously named Chilean species. Comparisons are provided with all other South American named species of <italic><named-content content-type="taxon-name">Aeneator</named-content>
</italic>
.</p>
</abstract>
<kwd-group><label>Keywords</label>
<kwd>New taxa</kwd>
<kwd>East Pacific</kwd>
<kwd>deep water</kwd>
</kwd-group>
</article-meta>
</front>
<body><sec><title>Introduction</title>
<p>The genus <italic><named-content content-type="taxon-name">Aeneator</named-content>
</italic>
Finlay, 1926 comprises a group of deep-water gastropods of moderate size, distributed in the South Pacific Ocean around New Zealand (<xref ref-type="bibr" rid="B17">Powell 1979</xref>
, <xref ref-type="bibr" rid="B4">Beu 1979</xref>
) and Chile (<xref ref-type="bibr" rid="B19">Rehder 1971</xref>
, <xref ref-type="bibr" rid="B13">McLean and Andrade 1982</xref>
, <xref ref-type="bibr" rid="B9">Fraussen and Sellanes 2008</xref>
). Almost all the species have offshore distributions, and they are common on the sea floor (<xref ref-type="bibr" rid="B6">Dawson 1965</xref>
, <xref ref-type="bibr" rid="B17">Powell 1979</xref>
, <xref ref-type="bibr" rid="B3">Beu and Maxwell 1990</xref>
). Their elongate fusiform shells have rounded whorls with a subsutural concavity, a lip with a broad shallow sinus below the suture, and a sculpture of strong axial ribs overridden by spiral cords (<xref ref-type="bibr" rid="B13">McLean and Andrade 1982</xref>
).</p>
<pmc-comment>PageBreak</pmc-comment>
<p>In the southeastern Pacific the genus encompasses five extant species: <italic><named-content content-type="taxon-name">Aeneator castillai</named-content>
</italic>
McLean & Andrade, 1982, <italic><named-content content-type="taxon-name">Aeneator fontainei</named-content>
</italic>
(d’Orbigny, 1839), <italic><named-content content-type="taxon-name">Aeneator (Ellicea) loisae</named-content>
</italic>
Rehder, 1971, <italic><named-content content-type="taxon-name">Aeneator portentosus</named-content>
</italic>
Fraussen & Sellanes, 2008 and <italic><named-content content-type="taxon-name">Aeneator prognaviter</named-content>
</italic>
Fraussen & Sellanes, 2008. The distribution of these species ranges from Bahía Independencia (14°S), in the south of Peru to Canal Moraleda 45°22'S, southern Chile (<xref ref-type="bibr" rid="B15">Osorio et al. 2006</xref>
). Their bathymetric range is from 10 m depth for <italic><named-content content-type="taxon-name">Aeneator fontainei</named-content>
</italic>
collected at Mejillones Bay, in the north of Chile (<xref ref-type="bibr" rid="B10">Guzmán et al. 1998</xref>
, <xref ref-type="bibr" rid="B11">Laudien et al. 2007</xref>
) to 800 m depth for <italic><named-content content-type="taxon-name">Aeneator portentosus</named-content>
</italic>
, collected off Iquique (<xref ref-type="bibr" rid="B9">Fraussen and Sellanes 2008</xref>
). Most of the species have been recovered in the trawls of the local shrimp industry (<xref ref-type="bibr" rid="B13">McLean and Andrade 1982</xref>
, <xref ref-type="bibr" rid="B19">Rehder 1971</xref>
, Párraga 2012, <xref ref-type="bibr" rid="B18">Queirolo et al. 2011</xref>
), and very little is known of their population biology, ecology and conservation status.</p>
<p>The present work describes a new species of <italic><named-content content-type="taxon-name">Aeneator</named-content>
</italic>
from northern Chile based on shell morphological features. Criteria were shell shape, number of primary spiral cords, development of secondary spirals, and axial sculpture. An identification key, based on shell characters, is given for all the extant Chilean <italic><named-content content-type="taxon-name">Aeneator</named-content>
</italic>
species.</p>
</sec>
<sec sec-type="materials|methods"><title>Material and methods</title>
<p>Material examined: <italic><named-content content-type="taxon-name">Aeneator martae</named-content>
</italic>
sp. n. types, Chile, Region of Atacama, Caldera, holotype MZUC 37890, paratype 1 MZUC 37891, paratype 2 MZUC 37892, paratype 3 MG 200105.</p>
<p>Examination was made of shell only specimens; all measurements were made with vernier callipers (± 0.1 mm). For the measure of length of aperture and angle of the spire, the methodology of <xref ref-type="bibr" rid="B7">Dépraz et al. (2009)</xref>
and <xref ref-type="bibr" rid="B5">Chiu et al. (2002)</xref>
was used.</p>
<p>Abbreviations: KF; Private collection of Mr Koen Fraussen, Aarschot, Belgium, MG: private collection of the author, section marine Gastropoda, MZUC; Museo de Zoología de la Universidad de Concepción, Concepción, Chile, RC Coll; private collection of Mr Ricardo Catalán, Servicio Nacional de Pesca, Chile.</p>
</sec>
<sec sec-type="Results"><title>Results</title>
<sec sec-type="Systematics"><title>Systematics</title>
<p>Class: <named-content content-type="taxon-name">Gastropoda</named-content>
Cuvier, 1797</p>
<p>Order: <named-content content-type="taxon-name">Neogastropoda</named-content>
Wenz, 1938</p>
<p>Superfamily: <named-content content-type="taxon-name">Buccinoidea</named-content>
Rafinesque, 1815</p>
<p>Family: <named-content content-type="taxon-name">Buccinidae</named-content>
Rafinesque, 1815</p>
<p><bold>Genus: <italic><named-content content-type="taxon-name">Aeneator</named-content>
</italic>
</bold>
<bold>Finlay, 1926:414</bold>
</p>
<p><bold>Type species.</bold>
<italic><named-content content-type="taxon-name">Verconella marshalli</named-content>
</italic>
Murdoch 1924 (by original designation), Pleistocene and recent, New Zealand.</p>
<pmc-comment>PageBreak</pmc-comment>
<sec sec-type="taxon-treatment"><title><named-content content-type="taxon-name"><named-content content-type="genus">Aeneator</named-content>
<named-content content-type="species">martae</named-content>
</named-content>
<named-content content-type="taxon-status">sp. n.</named-content>
</title>
<p>urn:lsid:zoobank.org:act:73AC9156-214E-4941-BFF6-0F94F8E17381</p>
<p>http://species-id.net/wiki/Aeneator_martae</p>
<p><xref ref-type="fig" rid="F1">Figs 1</xref>
<xref ref-type="fig" rid="F2">–14</xref>
<xref ref-type="fig" rid="F4">, 18</xref>
<xref ref-type="table" rid="T1">, Tables 1</xref>
<xref ref-type="table" rid="T2">, 2</xref>
<xref ref-type="table" rid="T3">, 3</xref>
</p>
<sec sec-type="treatment-Type material"><title>Type material.</title>
<p>Holotype (MZUC 37890), 47.9 mm. Chile, off Caldera (<named-content content-type="dwc:verbatimCoordinates">27°04'S, 70°50'W</named-content>
), 550–600 m depth, live collected on shrimp trawl nets, January 2001, S. Castillo leg. Paratype 1 (MZUC 37891), length 44.0 mm. Paratype 2 (MZUC 37892), 41.7 mm, Paratype 3 (MG 200105), length 40.2 mm. All the paratypes with same locality as the holotype.</p>
</sec>
<sec sec-type="treatment-Distribution"><title>Distribution.</title>
<p>Known only from the type locality; Chile, Region de Atacama, Caldera (<named-content content-type="dwc:verbatimCoordinates">27°04'S, 70°50'W</named-content>
), 550–600 m depth.</p>
</sec>
<sec sec-type="treatment-Diagnosis"><title>Diagnosis.</title>
<p>A small species of <italic><named-content content-type="taxon-name">Aeneator</named-content>
</italic>
, height up to 47.9 mm, shell stout, inside of aperture pale orange, exterior sculptured by well-defined axial ribs, spiral cords, and a conspicuous stepped shoulder.</p>
</sec>
<sec sec-type="treatment-Description"><title>Description.</title>
<p>Shell small for genus (height up to 47.9 mm, <xref ref-type="table" rid="T1">Table 1</xref>
), thick, solid, fusiform, chalky white to pale brownish, inside of aperture pale orange. Shape broad, angulate, length of aperture and canal more than half length of shell, width/height ratio 0.53 <pmc-comment>PageBreak</pmc-comment>
to 0.56, whorls convex apart from slightly concave sutural ramp, suture shallow but impressed. Spire angle 63° to 68°. Protoconch and upper teleoconch whorls missing, remaining whorls about 4.5, last 3 with sculpture intact with 7–9 primary spiral cords, interspaces each occupied by one narrow, well defined secondary cord. Last whorl with 16–18 spiral cords, more prominent at periphery of shell than elsewhere, forming a distinct keel. Spire whorls with 24–28 pronounced axial ribs, interspaces deep, each almost equal to a rib in width. Last whorl with 14–15 such ribs. Ribs more pronounced towards the anterior end of shell. Aperture ovate. Parietal and columellar area well-defined, glazed; outer lip thin, slightly crenulated, without lirae or teeth. Siphonal canal short, open, directed slightly to left. Operculum large, thin, dark brown, elongate, nucleus terminal, tip sharp.</p>
<table-wrap id="T1" orientation="portrait" position="float"><label>Table 1.</label>
<caption><p><italic><named-content content-type="taxon-name">Aeneator martae</named-content>
</italic>
sp. n. measurements of specimens. (%) means percentage compared to the total length of the shell.</p>
</caption>
<table frame="hsides" rules="groups"><tbody><tr><th rowspan="1" colspan="1"></th>
<th rowspan="1" colspan="1">Maximum length (mm)</th>
<th rowspan="1" colspan="1">Maximum width (mm)</th>
<th rowspan="1" colspan="1">Length of aperture</th>
<th rowspan="1" colspan="1">Width/Length</th>
</tr>
<tr><td rowspan="1" colspan="1">Holotype</td>
<td rowspan="1" colspan="1">47.9</td>
<td rowspan="1" colspan="1">25.6</td>
<td rowspan="1" colspan="1">26.8 (56 %)</td>
<td rowspan="1" colspan="1">0.53</td>
</tr>
<tr><td rowspan="1" colspan="1">Paratype 1</td>
<td rowspan="1" colspan="1">44.0</td>
<td rowspan="1" colspan="1">23.6</td>
<td rowspan="1" colspan="1">27.9 (57 %)</td>
<td rowspan="1" colspan="1">0.54</td>
</tr>
<tr><td rowspan="1" colspan="1">Paratype 2</td>
<td rowspan="1" colspan="1">41.7</td>
<td rowspan="1" colspan="1">23.4</td>
<td rowspan="1" colspan="1">23.0 (57 %)</td>
<td rowspan="1" colspan="1">0.56</td>
</tr>
<tr><td rowspan="1" colspan="1">Paratype 3</td>
<td rowspan="1" colspan="1">40.2</td>
<td rowspan="1" colspan="1">22.1</td>
<td rowspan="1" colspan="1">22.6 (56 %)</td>
<td rowspan="1" colspan="1">0.55</td>
</tr>
<tr><td rowspan="1" colspan="1">Average</td>
<td rowspan="1" colspan="1">43.4</td>
<td rowspan="1" colspan="1">23.6</td>
<td rowspan="1" colspan="1">25.1 (56 %)</td>
<td rowspan="1" colspan="1">0.54</td>
</tr>
</tbody>
</table>
</table-wrap>
<fig id="F1" orientation="portrait" position="float"><label>Figures 1–5.</label>
<caption><p><italic><named-content content-type="taxon-name">Aeneator martae</named-content>
</italic>
sp. n. shell, Holotype 47.9 mm, Chile, Off Caldera, <named-content content-type="dwc:verbatimCoordinates">27°04'S, 70°50'W</named-content>
. 550–600 m. MZUC 37890.</p>
</caption>
<graphic xlink:href="ZooKeys-257-089-g001"></graphic>
</fig>
<fig id="F2" orientation="portrait" position="float"><label>Figures 6–14.</label>
<caption><p><italic><named-content content-type="taxon-name">Aeneator martae</named-content>
</italic>
sp. n. shells. <bold>6–8</bold>
Paratype 1 (MZUC 37891), 44.0 mm height <bold>9–11</bold>
Paratype 2 (MZUC 37892), 41.7 mm height <bold>12–14</bold>
Paratype 3 (MG 200105), 40.2 mm height.</p>
</caption>
<graphic xlink:href="ZooKeys-257-089-g002"></graphic>
</fig>
<fig id="F3" orientation="portrait" position="float"><label>Figures 15–20.</label>
<caption><p>Details of shell sculpture of Chilean<italic><named-content content-type="taxon-name">Aeneator</named-content>
</italic>
species. <bold>15</bold>
<italic><named-content content-type="taxon-name">Aeneator castillai</named-content>
</italic>
(RC Coll.), 85.7 mm <bold>16</bold>
<italic><named-content content-type="taxon-name">Aeneator fontainei</named-content>
</italic>
(RC Coll.), 48.0 mm <bold>17</bold>
<italic><named-content content-type="taxon-name">Aeneator loisae</named-content>
</italic>
(MG 200003), 78 mm <bold>18</bold>
<italic><named-content content-type="taxon-name">Aeneator martae</named-content>
</italic>
sp. n. paraype 3 (MG 200105), 40.2 mm <bold>19</bold>
<italic><named-content content-type="taxon-name">Aeneator portentosus</named-content>
</italic>
, Paratype KF-0338, 45.5 mm <bold>20</bold>
<italic><named-content content-type="taxon-name">Aeneator prognaviter</named-content>
</italic>
(MG 200124), 33,0 mm.</p>
</caption>
<graphic xlink:href="ZooKeys-257-089-g003"></graphic>
</fig>
</sec>
<sec sec-type="treatment-Etymology"><title>Etymology.</title>
<p>Named in honour of Mrs Marta Araya,Caldera, Chile, who presented the specimens to the author.</p>
</sec>
<sec sec-type="treatment-Remarks"><title>Remarks.</title>
<p>In Chile the genus <italic><named-content content-type="taxon-name">Aeneator</named-content>
</italic>
encompasses five extant species: <italic><named-content content-type="taxon-name">Aeneator castillai</named-content>
</italic>
, found from Coquimbo (29°55'S) to Punta Peñablanca (33°22'S) in 200–450 m (<xref ref-type="bibr" rid="B13">McLean and Andrade 1982</xref>
), <italic><named-content content-type="taxon-name">Aeneator fontainei</named-content>
</italic>
, the most common species, with records from Bahía Independencia (14°S) in the south of Peru (<xref ref-type="bibr" rid="B13">McLean and Andrade 1982</xref>
) to Estero Elefantes, 46°05'S (<xref ref-type="bibr" rid="B15">Osorio et al. 2006</xref>
) and with a bathymetric range of 10 m near Mejillones (<xref ref-type="bibr" rid="B10">Guzmán et al. 1998</xref>
, <xref ref-type="bibr" rid="B11">Laudien et al. 2007</xref>
) to 421 m for a specimen collected off Coquimbo (<xref ref-type="fig" rid="F5">Figs 27–30</xref>
), <italic><named-content content-type="taxon-name">Aeneator (Ellicea) loisae</named-content>
</italic>
, distributed from Caldera (27°04'S), for material examined in this work (<xref ref-type="fig" rid="F5">Figs 27–30</xref>
), to Canal Moraleda (45°22'S), in the fjords area (<xref ref-type="bibr" rid="B15">Osorio et al. 2006</xref>
) with a bathymetric range of 200 m (McLean & Andrade 1982) to 465 m, <italic><named-content content-type="taxon-name">Aeneator portentosus</named-content>
</italic>
reported only form the original locality off Iquique (21°19'S) in 605 m and off Coquimbo at 800 m and <italic><named-content content-type="taxon-name">Aeneator prognaviter</named-content>
</italic>
, distributed off Antofagasta (22°51'S) in 318 m (<xref ref-type="bibr" rid="B9">Fraussen and Sellanes 2008</xref>
) and in 748 m off Iquique for material examined in this work (<xref ref-type="fig" rid="F6">Fig. 37</xref>
). Data on the localities of Chilean species of <italic><named-content content-type="taxon-name">Aeneator</named-content>
</italic>
is provided in <xref ref-type="table" rid="T3">Table 3</xref>
.</p>
<table-wrap id="T2" orientation="portrait" position="float"><label>Table 2.</label>
<caption><p>Synthesis of characters of the Chilean species of <italic><named-content content-type="taxon-name">Aeneator</named-content>
</italic>
Finlay, 1926 based on <xref ref-type="bibr" rid="B19">Rehder (1971)</xref>
, <xref ref-type="bibr" rid="B13">McLean and Andrade (1982)</xref>
, <xref ref-type="bibr" rid="B9">Fraussen and Sellanes (2008)</xref>
and material examined in this work.</p>
</caption>
<table frame="hsides" rules="groups"><tbody><tr><th rowspan="1" colspan="1"></th>
<th rowspan="1" colspan="1"><italic><named-content content-type="taxon-name">Aeneator castillai</named-content>
</italic>
</th>
<th rowspan="1" colspan="1"><italic><named-content content-type="taxon-name">Aeneator fontainei</named-content>
</italic>
</th>
<th rowspan="1" colspan="1"><italic><named-content content-type="taxon-name">Aeneator loisae</named-content>
</italic>
</th>
<th rowspan="1" colspan="1"><italic><named-content content-type="taxon-name">Aeneator portentosus</named-content>
</italic>
</th>
<th rowspan="1" colspan="1"><italic><named-content content-type="taxon-name">Aeneator prognaviter</named-content>
</italic>
</th>
<th rowspan="1" colspan="1"><italic><named-content content-type="taxon-name">Aeneator martae</named-content>
</italic>
sp. n.</th>
</tr>
<tr><td rowspan="1" colspan="1">Length</td>
<td rowspan="1" colspan="1">85.7</td>
<td rowspan="1" colspan="1">85.8</td>
<td rowspan="1" colspan="1">104</td>
<td rowspan="1" colspan="1">45.5</td>
<td rowspan="1" colspan="1">32.2</td>
<td rowspan="1" colspan="1">47.9</td>
</tr>
<tr><td rowspan="1" colspan="1">Width / Length</td>
<td rowspan="1" colspan="1">0.50–0.51</td>
<td rowspan="1" colspan="1">0.48–0.54</td>
<td rowspan="1" colspan="1">0.43–0.48</td>
<td rowspan="1" colspan="1">0.49–0.59</td>
<td rowspan="1" colspan="1">0.55 – 0.60</td>
<td rowspan="1" colspan="1">0.54–0.56</td>
</tr>
<tr><td rowspan="1" colspan="1">Aperture length/ total length</td>
<td rowspan="1" colspan="1">0.55–0.63</td>
<td rowspan="1" colspan="1">0.55–0.56</td>
<td rowspan="1" colspan="1">0.54 – 0.56</td>
<td rowspan="1" colspan="1">0.43–0.52</td>
<td rowspan="1" colspan="1">0.49 – 0.53</td>
<td rowspan="1" colspan="1">0.55–0.56</td>
</tr>
<tr><td rowspan="1" colspan="1">Spire angle</td>
<td rowspan="1" colspan="1">50°</td>
<td rowspan="1" colspan="1">51°–57°</td>
<td rowspan="1" colspan="1">44°–46°</td>
<td rowspan="1" colspan="1">44°–51°</td>
<td rowspan="1" colspan="1">60°</td>
<td rowspan="1" colspan="1">63°–68°</td>
</tr>
<tr><td rowspan="1" colspan="1">Axial ribs on last whorl</td>
<td rowspan="1" colspan="1">16, absent in subsutural area</td>
<td rowspan="1" colspan="1">12–15</td>
<td rowspan="1" colspan="1">Faint, absent</td>
<td rowspan="1" colspan="1">Faint, absent</td>
<td rowspan="1" colspan="1">22, bent</td>
<td rowspan="1" colspan="1">14–15, straight</td>
</tr>
<tr><td rowspan="1" colspan="1">Spiral cords in last whorl</td>
<td rowspan="1" colspan="1">12 – 15 brown primary, 1–3 secondary in each interspace</td>
<td rowspan="1" colspan="1">12–15 brown primary, 3–5 secondary in each interspace</td>
<td rowspan="1" colspan="1">9–10 primary, many secondary</td>
<td rowspan="1" colspan="1">20</td>
<td rowspan="1" colspan="1">20–24</td>
<td rowspan="1" colspan="1">16–18 primary, 7–9 secondary</td>
</tr>
<tr><td rowspan="1" colspan="1">Siphonal canal</td>
<td rowspan="1" colspan="1">short, twisted</td>
<td rowspan="1" colspan="1">medium to long, straight</td>
<td rowspan="1" colspan="1">long</td>
<td rowspan="1" colspan="1">short, broad, slightly bent</td>
<td rowspan="1" colspan="1">short, broad</td>
<td rowspan="1" colspan="1">short, slightly curved to left</td>
</tr>
<tr><td rowspan="1" colspan="1">Aperture</td>
<td rowspan="1" colspan="1">ovate</td>
<td rowspan="1" colspan="1">ovate</td>
<td rowspan="1" colspan="1">elongate ovate</td>
<td rowspan="1" colspan="1">round</td>
<td rowspan="1" colspan="1">oval</td>
<td rowspan="1" colspan="1">oval</td>
</tr>
<tr><td rowspan="1" colspan="1">Shell color</td>
<td rowspan="1" colspan="1">brown</td>
<td rowspan="1" colspan="1">white - yellowish</td>
<td rowspan="1" colspan="1">white</td>
<td rowspan="1" colspan="1">snow white</td>
<td rowspan="1" colspan="1">snow white</td>
<td rowspan="1" colspan="1">white, pale brownish</td>
</tr>
<tr><td rowspan="1" colspan="1">Distribution</td>
<td rowspan="1" colspan="1">29°55'S to 39.1°S</td>
<td rowspan="1" colspan="1">14°13'S to 46°S</td>
<td rowspan="1" colspan="1">27°04'S to 53.7°S</td>
<td rowspan="1" colspan="1">21.19°S and 29.95°S</td>
<td rowspan="1" colspan="1">21°19'S and 22°51'S</td>
<td rowspan="1" colspan="1">27°04'S</td>
</tr>
</tbody>
</table>
</table-wrap>
<table-wrap id="T3" orientation="portrait" position="float"><label>Table 3.</label>
<caption><p>Table of localities of Chilean species of <italic><named-content content-type="taxon-name">Aeneator</named-content>
</italic>
Finlay, 1926 based on <xref ref-type="bibr" rid="B19">Rehder (1971)</xref>
, <xref ref-type="bibr" rid="B13">McLean and Andrade (1982)</xref>
, <xref ref-type="bibr" rid="B9">Fraussen and Sellanes (2008)</xref>
, and material examined in this work.</p>
</caption>
<table frame="hsides" rules="groups"><tbody><tr><th rowspan="1" colspan="1">Species</th>
<th rowspan="1" colspan="1">Latitude</th>
<th rowspan="1" colspan="1">Longitude</th>
<th rowspan="1" colspan="1">Depth (m)</th>
</tr>
<tr><td rowspan="1" colspan="1"><italic><named-content content-type="taxon-name">Aeneator castillai</named-content>
</italic>
</td>
<td rowspan="1" colspan="1">29°55'S to 33°22'S</td>
<td rowspan="1" colspan="1">71°53'W to 71°20'W</td>
<td rowspan="1" colspan="1">200–450</td>
</tr>
<tr><td rowspan="1" colspan="1"><italic><named-content content-type="taxon-name">Aeneator fontainei</named-content>
</italic>
</td>
<td rowspan="1" colspan="1">14°14'S to 46°05'S</td>
<td rowspan="1" colspan="1">76°11'W to 73°41'W </td>
<td rowspan="1" colspan="1">10–421</td>
</tr>
<tr><td rowspan="1" colspan="1"><italic><named-content content-type="taxon-name">Aeneator loisae</named-content>
</italic>
</td>
<td rowspan="1" colspan="1">27°04'S to 45°22'S</td>
<td rowspan="1" colspan="1">73°21'W to 70°50'W </td>
<td rowspan="1" colspan="1">200–465</td>
</tr>
<tr><td rowspan="1" colspan="1"><italic><named-content content-type="taxon-name">Aeneator martae</named-content>
</italic>
</td>
<td rowspan="1" colspan="1">27°04'S</td>
<td rowspan="1" colspan="1">70°50'W</td>
<td rowspan="1" colspan="1">550–600</td>
</tr>
<tr><td rowspan="1" colspan="1"><italic><named-content content-type="taxon-name">Aeneator portentosus</named-content>
</italic>
</td>
<td rowspan="1" colspan="1">21°19'S to 29°55'S</td>
<td rowspan="1" colspan="1">71°20'W to 70°09'W</td>
<td rowspan="1" colspan="1">800</td>
</tr>
<tr><td rowspan="1" colspan="1"><italic><named-content content-type="taxon-name">Aeneator prognaviter</named-content>
</italic>
</td>
<td rowspan="1" colspan="1">21°19'S to 22°51'S</td>
<td rowspan="1" colspan="1">70°24'W to 70°09'W</td>
<td rowspan="1" colspan="1">600–748</td>
</tr>
</tbody>
</table>
</table-wrap>
<fig id="F4" orientation="portrait" position="float"><label>Figures 21–26.</label>
<caption><p><italic><named-content content-type="taxon-name">Aeneator fontainei</named-content>
</italic>
varieties and details of shell sculpture. <bold>21–22</bold>
Off Coquimbo, Chile, trawled 421 m (RC Coll), 58 mm <bold>23–24</bold>
Washed ashore, Calderilla beach, Caldera, Chile (MG 200011), 28.5 mm <bold>25–26</bold>
Dredged 20 m depth off Loreto beach, Caldera, Chile (MG 200012), 52.8 mm.</p>
</caption>
<graphic xlink:href="ZooKeys-257-089-g004"></graphic>
</fig>
<fig id="F5" orientation="portrait" position="float"><label>Figures 27–32.</label>
<caption><p><italic><named-content content-type="taxon-name">Aeneator loisae</named-content>
</italic>
varieties and details of shell sculpture <bold>27–28</bold>
Off Caldera, Chile, 450–500 m depth (MG 200003), 78 mm <bold>29–30</bold>
Off Caldera, Chile, 420 m depth (MG 200007) 71.9 mm <bold>31–32</bold>
Off Coquimbo, Chile, 400 m depth (RC Coll.), 104 mm</p>
</caption>
<graphic xlink:href="ZooKeys-257-089-g005"></graphic>
</fig>
<p>In size, the shell of <italic><named-content content-type="taxon-name">Aeneator martae</named-content>
</italic>
sp. n. is similar to <italic><named-content content-type="taxon-name">Aeneator prognaviter</named-content>
</italic>
(<xref ref-type="fig" rid="F3">Figs 20</xref>
<xref ref-type="fig" rid="F6">, 37, 38</xref>
) and <italic><named-content content-type="taxon-name">Aeneator portentosus</named-content>
</italic>
(<xref ref-type="fig" rid="F6">Figs 35, 36</xref>
). However, the former of these two can be clearly differentiated from the new species by its wider and shorter siphonal canal, less numerous and more curved axial ribs and a thinner, snow white shell (<xref ref-type="bibr" rid="B9">Fraussen and Sellanes 2008</xref>
). From <italic><named-content content-type="taxon-name">Aeneator portentosus</named-content>
</italic>
the new species differs by having a much wider, thicker shell with a shorter spire, a more elongate aperture, dominant axial sculpture and less rounded whorls. Moreover <italic><named-content content-type="taxon-name">Aeneator portentosus</named-content>
</italic>
exhibit a very distinctively sculptured peri<pmc-comment>PageBreak</pmc-comment>
<pmc-comment>PageBreak</pmc-comment>
ostracum (<xref ref-type="fig" rid="F3">Fig. 19</xref>
), with low axial ridges, very different from all the other Chilean <italic><named-content content-type="taxon-name">Aeneator</named-content>
</italic>
species. A periostracum is absent in the examined specimens of <italic><named-content content-type="taxon-name">Aeneator martae</named-content>
</italic>
sp. n.</p>
<fig id="F6" orientation="portrait" position="float"><label>Figures 33–38.</label>
<caption><p><italic><named-content content-type="taxon-name">Aeneator</named-content>
</italic>
species and details of shell sculpture. <bold>33–34</bold>
<italic><named-content content-type="taxon-name">Aeneator castillai</named-content>
</italic>
, off Coquimbo, Chile, 380 m depth (RC Coll.), 85.7 mm <bold>35–36</bold>
<italic><named-content content-type="taxon-name">Aeneator portentosus</named-content>
</italic>
, Paratype KF-0338, 45.5 mm <bold>37–38</bold>
<italic><named-content content-type="taxon-name">Aeneator prognaviter</named-content>
</italic>
, off Iquique, Chile, 748 m depth (MG 200124), 33,0 mm.</p>
</caption>
<graphic xlink:href="ZooKeys-257-089-g006"></graphic>
</fig>
<p><italic><named-content content-type="taxon-name">Aeneator castillai</named-content>
</italic>
(<xref ref-type="fig" rid="F6">Figs 33, 34</xref>
), and <italic><named-content content-type="taxon-name">Aeneator fontainei</named-content>
</italic>
(<xref ref-type="fig" rid="F4">Figs 21–26</xref>
) differ markedly from the new species by their much larger shells, reaching up to 85.8 mm, more fusiform shells, with a much less stepped or indistinct shoulder, lower and fewer axial ribs, <pmc-comment>PageBreak</pmc-comment>
brown primary spiral cords (<xref ref-type="fig" rid="F3">Figs 15, 16</xref>
) and lip lirated within. The spiral sculpture is quite different; <italic><named-content content-type="taxon-name">Aeneator fontainei</named-content>
</italic>
has 13 to 16 dark brown major cords, with interspaces filled with five secondary cords separated by fine grooves or by secondary and tertiary cords. <italic><named-content content-type="taxon-name">Aeneator castillai</named-content>
</italic>
has brown primary cords with 3 to 5 fine secondary cords fill<pmc-comment>PageBreak</pmc-comment>
<pmc-comment>PageBreak</pmc-comment>
ing the interspaces and exhibits a longer, twisted, siphonal canal. In contrast <italic><named-content content-type="taxon-name">Aeneator martae</named-content>
</italic>
sp. n. lacks any brown coloration, shows a sculpture of alternated single major and minor spiral cords defined mostly in the posterior part of the whorls, and has a conspicuous stepped shoulder, forming a keel at the periphery.</p>
<p><italic><named-content content-type="taxon-name">Aeneator loisae</named-content>
</italic>
(<xref ref-type="fig" rid="F5">Figs 27–32</xref>
) differs from the new species in having a larger, up to 104 mm, white to snow white shell (different from the white to light brown shell of <italic><named-content content-type="taxon-name">Aeneator martae</named-content>
</italic>
sp. n.), more inflated last whorl, with a much longer siphonal canal, a higher number of primary and secondary spiral cords, more prominent spiral sculpture, and fewer, more tenuous, axial ribs.</p>
<p>The new species is tentatively assigned, given the generic uncertainties within the Chilean species, to the genus <italic><named-content content-type="taxon-name">Aeneator</named-content>
</italic>
<xref ref-type="bibr" rid="B8">Finlay 1926</xref>
, typified by the species <italic><named-content content-type="taxon-name">Aeneator marshalli marshalli</named-content>
</italic>
(Murdoch, 1924) recorded from Castlecliff (as fossils) and, as a recent species (= <italic><named-content content-type="taxon-name">Aeneator marshalli separabilis</named-content>
</italic>
Dell, 1956), from Wanganui and Ohope beach, Whakatane, New Zealand. Similar to the type species, <italic><named-content content-type="taxon-name">Aeneator martae</named-content>
</italic>
sp. n. has a fusiform shell with moderately tall spire, shallow sinus in outer lip and a spiral sculpture of cords crossed by axial costae (<xref ref-type="bibr" rid="B3">Beu and Maxwell 1990</xref>
). The new species differs from <italic><named-content content-type="taxon-name">Aeneator marshalli</named-content>
</italic>
in its smaller shell, shorter anterior canal, the absence of nodules along the columellar lip, less inflated whorls and by the presence of a distinct keel at the periphery. From the genus <italic><named-content content-type="taxon-name">Austrofusus</named-content>
</italic>
Kobelt, 1879, with the type species <italic><named-content content-type="taxon-name">Austrofusus glans</named-content>
</italic>
(Röding, 1798), the new species differs in the smaller size, its thicker shell, more prominent sculpture, the more prominent ridges over the periphery, and the pale orange colour of the aperture, which is white in <italic><named-content content-type="taxon-name">Aeneator glans</named-content>
</italic>
(Beu & Marshall 2010). Comparative characters in the Chilean species of <italic><named-content content-type="taxon-name">Aeneator</named-content>
</italic>
are compared in <xref ref-type="table" rid="T2">table 2</xref>
.</p>
<p>In a recent revision of the fossil fauna of Mejillones, north of Chile (<xref ref-type="bibr" rid="B14">Nielsen 2012</xref>
), the species <italic><named-content content-type="taxon-name">Aeneator loisae</named-content>
</italic>
was synonymized with the fossil species <italic><named-content content-type="taxon-name">Fusus steinmanni</named-content>
</italic>
Möricke, 1896 into <italic><named-content content-type="taxon-name">Austrofusus</named-content>
</italic>
. However, this was based partly on the incorrect conclusion by <xref ref-type="bibr" rid="B1">Beu and Marshall (2010)</xref>
that <italic><named-content content-type="taxon-name">Aeneator fontainei</named-content>
</italic>
is the type species of <italic><named-content content-type="taxon-name">Austrofusus</named-content>
</italic>
; this was later corrected by <xref ref-type="bibr" rid="B2">Beu and Marshall (2011)</xref>
. On morphological grounds, the author concurs with <xref ref-type="bibr" rid="B13">McLean and Andrade (1982)</xref>
and considers that <italic><named-content content-type="taxon-name">Aeneator (Ellicea) loisae</named-content>
</italic>
does belong to the genus <italic><named-content content-type="taxon-name">Aeneator</named-content>
</italic>
and the sub-genus <italic><named-content content-type="taxon-name">Ellicea</named-content>
</italic>
Finlay in Marwick, 1928. However the generic placement of the species <italic><named-content content-type="taxon-name">Aeneator fontainei</named-content>
</italic>
, <italic><named-content content-type="taxon-name">Aeneator castillai</named-content>
</italic>
, and possibly the new species described here, should be further investigated or even be ascribed to a new genus.</p>
<p>Further study of radular characters, comparative anatomy and DNA will improve the taxonomic placement of the Chilean species. Fossil studies would also give a general insight into the development of the genus and their relationships with the South Pacific related fauna, especially those from New Zealand and adjacent waters.</p>
</sec>
<sec sec-type="treatment-Comparative material examined"><title>Comparative material examined:</title>
<p><italic><named-content content-type="taxon-name">Aeneator castillai</named-content>
</italic>
, Chile, Region of Coquimbo, Coquimbo, 2 specimens RC Coll. <italic><named-content content-type="taxon-name">Aeneator fontainei</named-content>
</italic>
, Chile, Region of Atacama, Caldera, 3 specimens MG 200011–200013, 5 specimens RC Coll. <italic><named-content content-type="taxon-name">Aeneator loisae</named-content>
</italic>
, Chile, Region of Atacama, Chile, 4 specimens MG 200003–200006, 1 specimen RC Coll, <italic><named-content content-type="taxon-name">Aeneator prognaviter</named-content>
</italic>
, 2 specimens MG 200124–200125, <italic><named-content content-type="taxon-name">Aeneator portentosus</named-content>
</italic>
, 1 specimen (examined from images), KF-0338.</p>
<pmc-comment>PageBreak</pmc-comment>
<pmc-comment>PageBreak</pmc-comment>
</sec>
</sec>
</sec>
<sec sec-type="Key for the identification of fully-grown Chilean species of Aeneator based on shell characters"><title>Key for the identification of fully-grown Chilean species of <italic><named-content content-type="taxon-name">Aeneator</named-content>
</italic>
based on shell characters</title>
<table-wrap content-type="key" orientation="portrait" id="d35e1187" position="anchor"><table frame="hsides" rules="groups"><tbody><tr><td rowspan="1" colspan="1">1</td>
<td rowspan="1" colspan="1">Aperture ovate-elongate</td>
<td rowspan="1" colspan="1">2</td>
</tr>
<tr><td rowspan="1" colspan="1">–</td>
<td rowspan="1" colspan="1">Aperture rounded, shell pagodoid, periostracum sculptured</td>
<td rowspan="1" colspan="1"><italic><named-content content-type="taxon-name">Aeneator portentosus</named-content>
</italic>
Fraussen & Sellanes, 2008</td>
</tr>
<tr><td rowspan="1" colspan="1">2(1)</td>
<td rowspan="1" colspan="1">Siphonal canal short</td>
<td rowspan="1" colspan="1">3</td>
</tr>
<tr><td rowspan="1" colspan="1">–</td>
<td rowspan="1" colspan="1">Siphonal canal long, outer lip reflexed, shell elongated</td>
<td rowspan="1" colspan="1"><italic><named-content content-type="taxon-name">Aeneator loisae</named-content>
</italic>
Rehder, 1971</td>
</tr>
<tr><td rowspan="1" colspan="1">3(2)</td>
<td rowspan="1" colspan="1">Spiral cords brown</td>
<td rowspan="1" colspan="1">5</td>
</tr>
<tr><td rowspan="1" colspan="1">–</td>
<td rowspan="1" colspan="1">Spiral cords white, axial ribs thick, shell length up to 49 mm</td>
<td rowspan="1" colspan="1">4</td>
</tr>
<tr><td rowspan="1" colspan="1">4(3)</td>
<td rowspan="1" colspan="1">Siphonal canal broad, axial ribs strongly curved</td>
<td rowspan="1" colspan="1"><italic><named-content content-type="taxon-name">Aeneator prognaviter</named-content>
</italic>
Fraussen & Sellanes, 2008</td>
</tr>
<tr><td rowspan="1" colspan="1">–</td>
<td rowspan="1" colspan="1">Shell with a distinct keel, aperture almost subquadrate</td>
<td rowspan="1" colspan="1"><italic><named-content content-type="taxon-name">Aeneator martae</named-content>
</italic>
sp. n.</td>
</tr>
<tr><td rowspan="1" colspan="1">5(3)</td>
<td rowspan="1" colspan="1">Axial ribs on subsutural area</td>
<td rowspan="1" colspan="1"><italic><named-content content-type="taxon-name">Aeneator fontanei</named-content>
</italic>
(d’Orbigny, 1841)</td>
</tr>
<tr><td rowspan="1" colspan="1">–</td>
<td rowspan="1" colspan="1">Sculpture absent on subsutural area, siphonal canal twisted</td>
<td rowspan="1" colspan="1"><italic><named-content content-type="taxon-name">Aeneator castillai</named-content>
</italic>
MacLean & Andrade, 1982</td>
</tr>
</tbody>
</table>
</table-wrap>
</sec>
</sec>
<sec sec-type="supplementary-material"><title>Supplementary Material</title>
<supplementary-material id="zookeys.257.4446-treatment1" content-type="local-data"><caption><title>XML Treatment for
<named-content content-type="genus">Aeneator</named-content>
<named-content content-type="species">martae</named-content>
</title>
</caption>
<media xlink:href="zookeys.257.4446-treatment1.xml" mimetype="text" mime-subtype="xml"></media>
</supplementary-material>
</sec>
</body>
<back><ack><title>Acknowledgements</title>
<p>The author wishes to thank Mrs Sandra Castillo for collecting the specimens <italic>in situ</italic>
, Mr Koen Fraussen (Aarschot, Belgium) for sending bibliography and images of the paratype of <italic><named-content content-type="taxon-name">Aeneator portentosus</named-content>
</italic>
, commenting about the genus, and reviewing the manuscript. I am grateful to Dr. Guillermo Guzmán (Museo del Mar, Universidad Arturo Prat, Chile) for helping with the specimens of <italic><named-content content-type="taxon-name">Aeneator prognaviter</named-content>
</italic>
, Sergio Miquel (Museo Argentino de Ciencias Naturales Bernardino Rivadavia, Buenos Aires, Argentina) for his comments on the early manuscript, Dr. Sven Nielsen (Institut für Geowissenschaften, Christian-Albrechts-Universität zu Kiel, Kiel, Germany) for sending useful bibliography, Mr Ricardo Catalán (Servicio Nacional de Pesca, Chile) for loaning <italic><named-content content-type="taxon-name">Aeneator</named-content>
</italic>
specimens for examination from his personal collection, Dr. Alan G. Beu (GNS Science, Lower Hutt, New Zealand) for his comments on the genus, help with essential bibliography, and for his comments and correction of the manuscript. Finally, I would like to thank the reviewers and the associate editor for their very helpful suggestions on the manuscript and the Encyclopedia of Life (EOL) Open Access Support Project (EOASP) for supporting the open access of this paper.</p>
</ack>
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