A new early Cretaceous clupeomorph fish from the Arratia Valley, Basque Country, Spain
Identifieur interne : 009039 ( Main/Exploration ); précédent : 009038; suivant : 009040A new early Cretaceous clupeomorph fish from the Arratia Valley, Basque Country, Spain
Auteurs : F. J. Poyato-Ariza [Espagne] ; M. A. L Pez-Horgue [Espagne] ; F. Garc Amp X0301 A-Garmilla [Espagne]Source :
- Cretaceous Research [ 0195-6671 ] ; 2000.
Descripteurs français
- Wicri :
- geographic : Espagne.
- topic : Pays basque.
English descriptors
- KwdEn :
- Abdominal scutes, Actinopterygii, American museum, Ammonium silicone, Anal, Arratia, Arratia valley, Articular, Articular head, Basque, Basque Country, Basque country, Bilbao, Black shales, Brackish water, Camera lucida, Carbonate, Carmeni, Caudal, Centrum, Chang, Chang grande, Ciencias, Ciencias naturales, Clupeomorph, Clupeomorpha, Clupeomorphs, Cretaceous, Diplomystus, Diplomystus dentatus, Doctoral thesis, Dorsal, Dorsal scutes, Early cretaceous, Early cretaceous clupeomorph, Ellimmichthys, Epineurals, Epural, Epurals, Ezkutuberezi, Ezkutuberezi carmeni, Facies, Fauna, Fossil, Grande, Herriko unibertsitatea, Holotype, Hypural, Hypurals, Lacustrine, Last dorsal scute, Lateral, Lateral view, Lower cretaceous, Lowest point, Main body, Maisey, Major sequences, Maximum body height, Mcna, Mesozoic, Museo civico, Natural history, Neural spine, Original specimen, Other characters, Paraclupea, Paratype upvlp, Patterson rosen, Positive cycles, Posterior dorsal scute, Posterior supramaxilla, Preural, Preural centrum, Primitive clupeomorphs, Proximal, Pujalte, Redrawn, Right side, Sandstone, Scute, Second ural centrum, Second uroneural, Shale, Shes, Silicone, Spain, Standard length, Storia naturale, Supramaxilla, Taverne, Taxon, Teleost, Third epural, Third hypural, Universidad, Upvlp, Ural, Ural centra, Ural centrum, Uroneural, Uyeno yabumoto, Ventral, Ventral border, Ventral scutes, Vertebral centra, Villaro, Villaro formation, Villaro member, Yabumoto, Younger specimens, black shales, early Cretaceous, lacustrine system.
- Teeft :
- Abdominal scutes, American museum, Ammonium silicone, Anal, Arratia, Arratia valley, Articular, Articular head, Basque, Basque country, Bilbao, Black shales, Brackish water, Camera lucida, Carbonate, Carmeni, Caudal, Centrum, Chang, Chang grande, Ciencias, Ciencias naturales, Clupeomorph, Clupeomorpha, Clupeomorphs, Cretaceous, Diplomystus, Diplomystus dentatus, Doctoral thesis, Dorsal, Dorsal scutes, Early cretaceous, Early cretaceous clupeomorph, Ellimmichthys, Epineurals, Epural, Epurals, Ezkutuberezi, Ezkutuberezi carmeni, Facies, Fauna, Fossil, Grande, Herriko unibertsitatea, Holotype, Hypural, Hypurals, Lacustrine, Last dorsal scute, Lateral, Lateral view, Lower cretaceous, Lowest point, Main body, Maisey, Major sequences, Maximum body height, Mcna, Mesozoic, Museo civico, Natural history, Neural spine, Original specimen, Other characters, Paraclupea, Paratype upvlp, Patterson rosen, Positive cycles, Posterior dorsal scute, Posterior supramaxilla, Preural, Preural centrum, Primitive clupeomorphs, Proximal, Pujalte, Redrawn, Right side, Sandstone, Scute, Second ural centrum, Second uroneural, Shale, Shes, Silicone, Standard length, Storia naturale, Supramaxilla, Taverne, Taxon, Teleost, Third epural, Third hypural, Universidad, Upvlp, Ural, Ural centra, Ural centrum, Uroneural, Uyeno yabumoto, Ventral, Ventral border, Ventral scutes, Vertebral centra, Villaro, Villaro formation, Villaro member, Yabumoto, Younger specimens.
Abstract
Abstract: Fish-bearing beds of early Cretaceous age crop out in the Arratia Valley, province of Biscay, along the nucleus of the Bilbao Anticlinorium in the Basque-Cantabrian Basin, northern Spain. Teleost fossils have been found in several stratigraphic levels, dated as Valanginian–Barremian, in the Villaro Member of the Villaro Formation. The Villaro Member is a 1100-m-thick succession of black shales, sandstones and scattered limestones that represent deposition in a shallow lacustrine system periodically fed by delta prograding bars and channels. Deposition took place in a restricted freshwater–brackish environment that was occasionally affected by marine incursions. The first new taxon from the Arratia Valley, described here, isEzkutuberezi carmeni gen. nov., sp. nov., a deep-bodied clupeomorph whose autapomorphies include: a posterior supramaxilla with a long anterior process forming an angle of 90° with the main body of the bone; a hypertrophied, claw-shaped last dorsal scute; only two epurals; and hypural 3 with a submedian longitudinal crest. The morphology of the dorsal scutes supports its assignment to the paraclupeid subfamily Paraclupeinae within the order Ellimmichthyiformes. The new taxon is the oldest European clupeomorph, and the first articulated vertebrate fossil to be described from the Basque-Cantabrian Basin.
Url:
DOI: 10.1006/cres.2000.0212
Affiliations:
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J." last="Poyato-Ariza">F. J. Poyato-Ariza</name><affiliation wicri:level="3"><country wicri:rule="url">Espagne</country><wicri:regionArea>Unidad de Paleontologı́a, Departamento de Biologı́a, Facultad de Ciencias, Universidad Autónoma, Cantoblanco, 28049, Madrid</wicri:regionArea><placeName><settlement type="city">Madrid</settlement><region nuts="2" type="region">Communauté de Madrid</region></placeName></affiliation></author><author><name sortKey="L Pez Horgue, M A" sort="L Pez Horgue, M A" uniqKey="L Pez Horgue M" first="M. A." last="L Pez-Horgue">M. A. 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Garc Amp X0301 A-Garmilla</name><affiliation wicri:level="1"><country xml:lang="fr">Espagne</country><wicri:regionArea>Departamento de Mineralogı́a y Petrologı́a, Facultad de Ciencias, UPV/EHU, 48080, Bilbao</wicri:regionArea><wicri:noRegion>Bilbao</wicri:noRegion></affiliation></author></analytic><monogr></monogr><series><title level="j">Cretaceous Research</title><title level="j" type="abbrev">YCRES</title><idno type="ISSN">0195-6671</idno><imprint><publisher>ELSEVIER</publisher><date type="published" when="2000">2000</date><biblScope unit="volume">21</biblScope><biblScope unit="issue">4</biblScope><biblScope unit="page" from="571">571</biblScope><biblScope unit="page" to="585">585</biblScope></imprint><idno type="ISSN">0195-6671</idno></series></biblStruct></sourceDesc><seriesStmt><idno type="ISSN">0195-6671</idno></seriesStmt></fileDesc><profileDesc><textClass><keywords scheme="KwdEn" xml:lang="en"><term>Abdominal scutes</term><term>Actinopterygii</term><term>American museum</term><term>Ammonium silicone</term><term>Anal</term><term>Arratia</term><term>Arratia valley</term><term>Articular</term><term>Articular head</term><term>Basque</term><term>Basque Country</term><term>Basque country</term><term>Bilbao</term><term>Black shales</term><term>Brackish water</term><term>Camera lucida</term><term>Carbonate</term><term>Carmeni</term><term>Caudal</term><term>Centrum</term><term>Chang</term><term>Chang grande</term><term>Ciencias</term><term>Ciencias naturales</term><term>Clupeomorph</term><term>Clupeomorpha</term><term>Clupeomorphs</term><term>Cretaceous</term><term>Diplomystus</term><term>Diplomystus dentatus</term><term>Doctoral thesis</term><term>Dorsal</term><term>Dorsal scutes</term><term>Early cretaceous</term><term>Early cretaceous clupeomorph</term><term>Ellimmichthys</term><term>Epineurals</term><term>Epural</term><term>Epurals</term><term>Ezkutuberezi</term><term>Ezkutuberezi carmeni</term><term>Facies</term><term>Fauna</term><term>Fossil</term><term>Grande</term><term>Herriko unibertsitatea</term><term>Holotype</term><term>Hypural</term><term>Hypurals</term><term>Lacustrine</term><term>Last dorsal scute</term><term>Lateral</term><term>Lateral view</term><term>Lower cretaceous</term><term>Lowest point</term><term>Main body</term><term>Maisey</term><term>Major sequences</term><term>Maximum body height</term><term>Mcna</term><term>Mesozoic</term><term>Museo civico</term><term>Natural history</term><term>Neural spine</term><term>Original specimen</term><term>Other characters</term><term>Paraclupea</term><term>Paratype upvlp</term><term>Patterson rosen</term><term>Positive cycles</term><term>Posterior dorsal scute</term><term>Posterior supramaxilla</term><term>Preural</term><term>Preural centrum</term><term>Primitive clupeomorphs</term><term>Proximal</term><term>Pujalte</term><term>Redrawn</term><term>Right side</term><term>Sandstone</term><term>Scute</term><term>Second ural centrum</term><term>Second uroneural</term><term>Shale</term><term>Shes</term><term>Silicone</term><term>Spain</term><term>Standard length</term><term>Storia naturale</term><term>Supramaxilla</term><term>Taverne</term><term>Taxon</term><term>Teleost</term><term>Third epural</term><term>Third hypural</term><term>Universidad</term><term>Upvlp</term><term>Ural</term><term>Ural centra</term><term>Ural centrum</term><term>Uroneural</term><term>Uyeno yabumoto</term><term>Ventral</term><term>Ventral border</term><term>Ventral scutes</term><term>Vertebral centra</term><term>Villaro</term><term>Villaro formation</term><term>Villaro member</term><term>Yabumoto</term><term>Younger specimens</term><term>black shales</term><term>early Cretaceous</term><term>lacustrine system</term></keywords><keywords scheme="Teeft" xml:lang="en"><term>Abdominal scutes</term><term>American museum</term><term>Ammonium silicone</term><term>Anal</term><term>Arratia</term><term>Arratia valley</term><term>Articular</term><term>Articular head</term><term>Basque</term><term>Basque country</term><term>Bilbao</term><term>Black shales</term><term>Brackish water</term><term>Camera lucida</term><term>Carbonate</term><term>Carmeni</term><term>Caudal</term><term>Centrum</term><term>Chang</term><term>Chang grande</term><term>Ciencias</term><term>Ciencias naturales</term><term>Clupeomorph</term><term>Clupeomorpha</term><term>Clupeomorphs</term><term>Cretaceous</term><term>Diplomystus</term><term>Diplomystus dentatus</term><term>Doctoral thesis</term><term>Dorsal</term><term>Dorsal scutes</term><term>Early cretaceous</term><term>Early cretaceous clupeomorph</term><term>Ellimmichthys</term><term>Epineurals</term><term>Epural</term><term>Epurals</term><term>Ezkutuberezi</term><term>Ezkutuberezi carmeni</term><term>Facies</term><term>Fauna</term><term>Fossil</term><term>Grande</term><term>Herriko unibertsitatea</term><term>Holotype</term><term>Hypural</term><term>Hypurals</term><term>Lacustrine</term><term>Last dorsal scute</term><term>Lateral</term><term>Lateral view</term><term>Lower cretaceous</term><term>Lowest point</term><term>Main body</term><term>Maisey</term><term>Major sequences</term><term>Maximum body height</term><term>Mcna</term><term>Mesozoic</term><term>Museo civico</term><term>Natural history</term><term>Neural spine</term><term>Original specimen</term><term>Other characters</term><term>Paraclupea</term><term>Paratype upvlp</term><term>Patterson rosen</term><term>Positive cycles</term><term>Posterior dorsal scute</term><term>Posterior supramaxilla</term><term>Preural</term><term>Preural centrum</term><term>Primitive clupeomorphs</term><term>Proximal</term><term>Pujalte</term><term>Redrawn</term><term>Right side</term><term>Sandstone</term><term>Scute</term><term>Second ural centrum</term><term>Second uroneural</term><term>Shale</term><term>Shes</term><term>Silicone</term><term>Standard length</term><term>Storia naturale</term><term>Supramaxilla</term><term>Taverne</term><term>Taxon</term><term>Teleost</term><term>Third epural</term><term>Third hypural</term><term>Universidad</term><term>Upvlp</term><term>Ural</term><term>Ural centra</term><term>Ural centrum</term><term>Uroneural</term><term>Uyeno yabumoto</term><term>Ventral</term><term>Ventral border</term><term>Ventral scutes</term><term>Vertebral centra</term><term>Villaro</term><term>Villaro formation</term><term>Villaro member</term><term>Yabumoto</term><term>Younger specimens</term></keywords><keywords scheme="Wicri" type="geographic" xml:lang="fr"><term>Espagne</term></keywords><keywords scheme="Wicri" type="topic" xml:lang="fr"><term>Pays basque</term></keywords></textClass><langUsage><language ident="en">en</language></langUsage></profileDesc></teiHeader><front><div type="abstract" xml:lang="en">Abstract: Fish-bearing beds of early Cretaceous age crop out in the Arratia Valley, province of Biscay, along the nucleus of the Bilbao Anticlinorium in the Basque-Cantabrian Basin, northern Spain. Teleost fossils have been found in several stratigraphic levels, dated as Valanginian–Barremian, in the Villaro Member of the Villaro Formation. The Villaro Member is a 1100-m-thick succession of black shales, sandstones and scattered limestones that represent deposition in a shallow lacustrine system periodically fed by delta prograding bars and channels. Deposition took place in a restricted freshwater–brackish environment that was occasionally affected by marine incursions. The first new taxon from the Arratia Valley, described here, isEzkutuberezi carmeni gen. nov., sp. nov., a deep-bodied clupeomorph whose autapomorphies include: a posterior supramaxilla with a long anterior process forming an angle of 90° with the main body of the bone; a hypertrophied, claw-shaped last dorsal scute; only two epurals; and hypural 3 with a submedian longitudinal crest. The morphology of the dorsal scutes supports its assignment to the paraclupeid subfamily Paraclupeinae within the order Ellimmichthyiformes. The new taxon is the oldest European clupeomorph, and the first articulated vertebrate fossil to be described from the Basque-Cantabrian Basin.</div></front></TEI><affiliations><list><country><li>Espagne</li></country><region><li>Communauté de Madrid</li></region><settlement><li>Madrid</li></settlement></list><tree><country name="Espagne"><region name="Communauté de Madrid"><name sortKey="Poyato Ariza, F J" sort="Poyato Ariza, F J" uniqKey="Poyato Ariza F" first="F. J." last="Poyato-Ariza">F. J. Poyato-Ariza</name></region><name sortKey="Garc Amp X0301 A Garmilla, F" sort="Garc Amp X0301 A Garmilla, F" uniqKey="Garc Amp X0301 A Garmilla F" first="F." last="Garc Amp X0301 A-Garmilla">F. Garc Amp X0301 A-Garmilla</name><name sortKey="L Pez Horgue, M A" sort="L Pez Horgue, M A" uniqKey="L Pez Horgue M" first="M. A." last="L Pez-Horgue">M. A. L Pez-Horgue</name></country></tree></affiliations></record>Pour manipuler ce document sous Unix (Dilib)
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