A revision of Aceratherium blanfordi Lydekker, 1884 (Mammalia: Rhinocerotidae) from the Early Miocene of Pakistan: postcranials as a key
Identifieur interne : 001D41 ( Istex/Corpus ); précédent : 001D40; suivant : 001D42A revision of Aceratherium blanfordi Lydekker, 1884 (Mammalia: Rhinocerotidae) from the Early Miocene of Pakistan: postcranials as a key
Auteurs : Pierre-Olivier Antoine ; Kevin F. Downing ; Jean-Yves Crochet ; Francis Duranthon ; Lawrence J. Flynn ; Laurent Marivaux ; Grégoire Métais ; Abdul Rahim Rajpar ; Ghazala RoohiSource :
- Zoological Journal of the Linnean Society [ 0024-4082 ] ; 2010-09.
English descriptors
- KwdEn :
- Aceratheriini, Aceratherium, Albigense, Alicornops, Antecrochet, Anterior, Anterior border, Anterior side, Anterior view, Anteroposterior, Anteroposterior diameter, Antoine, Antoine welcomme, Apophysis, Approximate dimensions, Articular, Articulation, Astragalus, Attened, Balochistan, Bicornis, Blanfordi, Bonis, Brachypotherium, Brachypus, Bracket, Browni, Bugti, Bugti hills, Bula, Calcaneum, Calcaneus, Carpalis, Cheek teeth, Chilotherium, Chitarwata, Cingulum, Clade, Coeval, Constricted, Convex, Corpus mandibulae, Cranial, Crista, Cuboid, Deciduous, Dentition, Dera, Dera bugti, Diaceratherium, Diaphysis, Diceratherium, Dicerorhinus, Diceros, Dihedron, Distal articulation, Distal border, Distal view, Dorsal, Dorsal view, Douvillei, Duvernoy, Earliest miocene, Early miocene, Ectocuneiform, Ectoloph, Ectometaloph, Elasmotheriina, Elasmotheriinae, Entoconid, Extensor carpalis, External groove, Facet, Fauna, Femur, Foramen, Fossil, Gaimersheimense, Gaindatherium, Gand, Gers, Ginsburg, Groove, Gutter, Heissig, Homoplastic, Hooijer, Humerus, Hypocone, Hypolophid, Incisivum, Insertion, Interossei, Kumbi, Labial, Labial cingulum, Lartet, Lateral, Lateral side, Lateral view, Laterally, Lholi, Lingual, Lingual bridge, Lingual cingulum, Lingual groove, Linnaeus, Linnean, Linnean society, Lower cheek teeth, Lower molars, Lower premolars, Lydekker, Mammal, Mammalia, Mandible, Mandibulae, Mandibular, Maxilla, Mcii, Mciii, Medial, Medial side, Median, Medifossette, Mesaceratherium, Mesaceratherium welcommi, Mesocuneiform, Mesostyle, Metacarpal, Metacone, Metaconid, Metaloph, Metapodials, Mhnt, Minutum, Miocene, Miocene rhinocerotids, Mirallesi, Mnhn, Molar, Monophyletic, Mtii, Mtiii, Mtiv, Natural history, Naturelle, Navicular, Node, Oblique, Occlusal, Occlusal view, Oligocene, Osborni, Outgroup, Paracone, Paralophid, Patella, Paulhiacense, Perissodactyla, Phylogenetic, Plesiaceratherium, Pleuroceros, Pleuroceros blanfordi, Pmnh, Postcranial, Postcranials, Posterior, Posterior border, Posterior part, Posterior tuberosity, Posterior valley, Posterolateral, Posteroproximal, Praecursor, Premolar, Processus, Processus zygomaticus, Prosantorhinus, Protaceratherium, Protocone, Protoloph, Prox, Proximal, Proximal border, Proximal view, Reversion, Rhinocerotid, Rhinocerotidae, Rhinocerotids, Rhinocerotina, Rhinocerotinae, Right astragalus, Right cuboid, Right scaphoid, Robust, Ronzotherium, Sagittal, Sagittally, Scaphoid, Semilunate, Sensu, Simorrense, Squamosal, Square brackets, Stratigraphical, Suprageneric, Symphysis, Synapomorphies, Talus, Taxon, Tibia, Transverse diameter, Trapezoid, Trigonias, Trigonid, Trochlea, Tuber, Tubercle, Tuberosity, Ulna, Unciform, Upper member, Upper molars, Upper premolars, Welcomme, Welcommi, Zinda, Zoological, Zoological journal, Zygomatic, Zygomatic arch, Zygomaticus.
- Teeft :
- Aceratheriini, Aceratherium, Albigense, Alicornops, Antecrochet, Anterior, Anterior border, Anterior side, Anterior view, Anteroposterior, Anteroposterior diameter, Antoine, Antoine welcomme, Apophysis, Approximate dimensions, Articular, Articulation, Astragalus, Attened, Balochistan, Bicornis, Blanfordi, Bonis, Brachypotherium, Brachypus, Bracket, Browni, Bugti, Bugti hills, Bula, Calcaneum, Calcaneus, Carpalis, Cheek teeth, Chilotherium, Chitarwata, Cingulum, Clade, Coeval, Constricted, Convex, Corpus mandibulae, Cranial, Crista, Cuboid, Deciduous, Dentition, Dera, Dera bugti, Diaceratherium, Diaphysis, Diceratherium, Dicerorhinus, Diceros, Dihedron, Distal articulation, Distal border, Distal view, Dorsal, Dorsal view, Douvillei, Duvernoy, Earliest miocene, Early miocene, Ectocuneiform, Ectoloph, Ectometaloph, Elasmotheriina, Elasmotheriinae, Entoconid, Extensor carpalis, External groove, Facet, Fauna, Femur, Foramen, Fossil, Gaimersheimense, Gaindatherium, Gand, Gers, Ginsburg, Groove, Gutter, Heissig, Homoplastic, Hooijer, Humerus, Hypocone, Hypolophid, Incisivum, Insertion, Interossei, Kumbi, Labial, Labial cingulum, Lartet, Lateral, Lateral side, Lateral view, Laterally, Lholi, Lingual, Lingual bridge, Lingual cingulum, Lingual groove, Linnaeus, Linnean, Linnean society, Lower cheek teeth, Lower molars, Lower premolars, Lydekker, Mammal, Mammalia, Mandible, Mandibulae, Mandibular, Maxilla, Mcii, Mciii, Medial, Medial side, Median, Medifossette, Mesaceratherium, Mesaceratherium welcommi, Mesocuneiform, Mesostyle, Metacarpal, Metacone, Metaconid, Metaloph, Metapodials, Mhnt, Minutum, Miocene, Miocene rhinocerotids, Mirallesi, Mnhn, Molar, Monophyletic, Mtii, Mtiii, Mtiv, Natural history, Naturelle, Navicular, Node, Oblique, Occlusal, Occlusal view, Oligocene, Osborni, Outgroup, Paracone, Paralophid, Patella, Paulhiacense, Perissodactyla, Phylogenetic, Plesiaceratherium, Pleuroceros, Pleuroceros blanfordi, Pmnh, Postcranial, Postcranials, Posterior, Posterior border, Posterior part, Posterior tuberosity, Posterior valley, Posterolateral, Posteroproximal, Praecursor, Premolar, Processus, Processus zygomaticus, Prosantorhinus, Protaceratherium, Protocone, Protoloph, Prox, Proximal, Proximal border, Proximal view, Reversion, Rhinocerotid, Rhinocerotidae, Rhinocerotids, Rhinocerotina, Rhinocerotinae, Right astragalus, Right cuboid, Right scaphoid, Robust, Ronzotherium, Sagittal, Sagittally, Scaphoid, Semilunate, Sensu, Simorrense, Squamosal, Square brackets, Stratigraphical, Suprageneric, Symphysis, Synapomorphies, Talus, Taxon, Tibia, Transverse diameter, Trapezoid, Trigonias, Trigonid, Trochlea, Tuber, Tubercle, Tuberosity, Ulna, Unciform, Upper member, Upper molars, Upper premolars, Welcomme, Welcommi, Zinda, Zoological, Zoological journal, Zygomatic, Zygomatic arch, Zygomaticus.
Abstract
Rhinocerotids are particularly abundant and diversified in Neogene deposits of the Indian subcontinent, but their systematics is far from being well defined. Based on the revision of old collections and new findings from the Early Miocene of the Bugti Hills and Zinda Pir, Pakistan, ‘Aceratherium blanfordi Lydekker, 1884’ is a chimera, consisting of two dentally convergent but postcranially distinct rhinocerotid taxa: Pleuroceros blanfordi and Mesaceratherium welcommi sp. nov. Postcranial features appear to be much more diagnostic than craniodental morphology in this case. A phylogenetic analysis based on 282 morphological characters scored for 28 taxa (four outgroups and ingroup including both taxa of interest and a ‘branching group’) strengthens this statement and supports Pleuroceros and Mesaceratherium as monophyletic genera within Rhinocerotinae. Both genera are recognized for the first time outside Europe. In the Bugti Hills, P. blanfordi and M. welcommi are part of an exceptionally diversified rhinocerotid fauna, with up to nine species associated in the same locality (Kumbi 4f). This rhinocerotid assemblage confirms the earliest Miocene age (Agenian/Aquitanian) of the upper member of the Chitarwata Formation as a whole. Coeval homotaxic rhinocerotid faunas from Europe (France, Czech Republic) and East Africa (Uganda, Kenya) support broad and sustainable rhinocerotid interchanges amongst South Asia, Europe, and Africa under compatible environmental conditions throughout earliest Miocene times.
Url:
DOI: 10.1111/j.1096-3642.2009.00597.x
Links to Exploration step
ISTEX:3C10EB81E8129E9F8E7C93E4DEC42360A31E54B3Le document en format XML
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Downing</name><affiliation><mods:affiliation>DePaul University, Chicago, IL 60604, USA</mods:affiliation></affiliation></author><author><name sortKey="Crochet, Jean Ves" sort="Crochet, Jean Ves" uniqKey="Crochet J" first="Jean-Yves" last="Crochet">Jean-Yves Crochet</name><affiliation><mods:affiliation>Département de Paléontologie, Institut des Sciences de l'Évolution (CNRS‐UMR 5554) Université Montpellier 2, c.c. 064, Place Eugène Bataillon, F‐34095 Montpellier Cedex 5, France</mods:affiliation></affiliation></author><author><name sortKey="Duranthon, Francis" sort="Duranthon, Francis" uniqKey="Duranthon F" first="Francis" last="Duranthon">Francis Duranthon</name><affiliation><mods:affiliation>Laboratoire d'Étude et de Conservation du Patrimoine, Muséum d'Histoire naturelle de Toulouse, 39 allées Jules Guesde, F‐31000 Toulouse, France</mods:affiliation></affiliation></author><author><name sortKey="Flynn, Lawrence J" sort="Flynn, Lawrence J" uniqKey="Flynn L" first="Lawrence J." last="Flynn">Lawrence J. Flynn</name><affiliation><mods:affiliation>Department of Anthropology and Peabody Museum, Harvard University, Cambridge, MA 02138, USA</mods:affiliation></affiliation></author><author><name sortKey="Marivaux, Laurent" sort="Marivaux, Laurent" uniqKey="Marivaux L" first="Laurent" last="Marivaux">Laurent Marivaux</name><affiliation><mods:affiliation>Département de Paléontologie, Institut des Sciences de l'Évolution (CNRS‐UMR 5554) Université Montpellier 2, c.c. 064, Place Eugène Bataillon, F‐34095 Montpellier Cedex 5, France</mods:affiliation></affiliation></author><author><name sortKey="Metais, Gregoire" sort="Metais, Gregoire" uniqKey="Metais G" first="Grégoire" last="Métais">Grégoire Métais</name><affiliation><mods:affiliation>Laboratoire de Paléontologie, UMR 5143 CNRS, Muséum National d'Histoire Naturelle, 8 rue Buffon, F‐75005 Paris, France</mods:affiliation></affiliation></author><author><name sortKey="Rajpar, Abdul Rahim" sort="Rajpar, Abdul Rahim" uniqKey="Rajpar A" first="Abdul Rahim" last="Rajpar">Abdul Rahim Rajpar</name><affiliation><mods:affiliation>9Earth Sciences Division, Pakistan Museum of Natural History, Garden Avenue, Shakarparian, 44000 Islamabad, Pakistan</mods:affiliation></affiliation></author><author><name sortKey="Roohi, Ghazala" sort="Roohi, Ghazala" uniqKey="Roohi G" first="Ghazala" last="Roohi">Ghazala Roohi</name><affiliation><mods:affiliation>9Earth Sciences Division, Pakistan Museum of Natural History, Garden Avenue, Shakarparian, 44000 Islamabad, Pakistan</mods:affiliation></affiliation></author></analytic><monogr></monogr><series><title level="j" type="main">Zoological Journal of the Linnean Society</title><title level="j" type="alt">ZOOLOGICAL JOURNAL OF THE LINNEAN SOCIETY</title><idno type="ISSN">0024-4082</idno><idno type="eISSN">1096-3642</idno><imprint><biblScope unit="vol">160</biblScope><biblScope unit="issue">1</biblScope><biblScope unit="page" from="139">139</biblScope><biblScope unit="page" to="194">194</biblScope><biblScope unit="page-count">56</biblScope><publisher>Blackwell Publishing Ltd</publisher><pubPlace>Oxford, UK</pubPlace><date type="published" when="2010-09">2010-09</date></imprint><idno type="ISSN">0024-4082</idno></series></biblStruct></sourceDesc><seriesStmt><idno type="ISSN">0024-4082</idno></seriesStmt></fileDesc><profileDesc><textClass><keywords scheme="KwdEn" xml:lang="en"><term>Aceratheriini</term><term>Aceratherium</term><term>Albigense</term><term>Alicornops</term><term>Antecrochet</term><term>Anterior</term><term>Anterior border</term><term>Anterior side</term><term>Anterior view</term><term>Anteroposterior</term><term>Anteroposterior diameter</term><term>Antoine</term><term>Antoine welcomme</term><term>Apophysis</term><term>Approximate dimensions</term><term>Articular</term><term>Articulation</term><term>Astragalus</term><term>Attened</term><term>Balochistan</term><term>Bicornis</term><term>Blanfordi</term><term>Bonis</term><term>Brachypotherium</term><term>Brachypus</term><term>Bracket</term><term>Browni</term><term>Bugti</term><term>Bugti hills</term><term>Bula</term><term>Calcaneum</term><term>Calcaneus</term><term>Carpalis</term><term>Cheek teeth</term><term>Chilotherium</term><term>Chitarwata</term><term>Cingulum</term><term>Clade</term><term>Coeval</term><term>Constricted</term><term>Convex</term><term>Corpus mandibulae</term><term>Cranial</term><term>Crista</term><term>Cuboid</term><term>Deciduous</term><term>Dentition</term><term>Dera</term><term>Dera bugti</term><term>Diaceratherium</term><term>Diaphysis</term><term>Diceratherium</term><term>Dicerorhinus</term><term>Diceros</term><term>Dihedron</term><term>Distal articulation</term><term>Distal border</term><term>Distal view</term><term>Dorsal</term><term>Dorsal view</term><term>Douvillei</term><term>Duvernoy</term><term>Earliest miocene</term><term>Early miocene</term><term>Ectocuneiform</term><term>Ectoloph</term><term>Ectometaloph</term><term>Elasmotheriina</term><term>Elasmotheriinae</term><term>Entoconid</term><term>Extensor carpalis</term><term>External groove</term><term>Facet</term><term>Fauna</term><term>Femur</term><term>Foramen</term><term>Fossil</term><term>Gaimersheimense</term><term>Gaindatherium</term><term>Gand</term><term>Gers</term><term>Ginsburg</term><term>Groove</term><term>Gutter</term><term>Heissig</term><term>Homoplastic</term><term>Hooijer</term><term>Humerus</term><term>Hypocone</term><term>Hypolophid</term><term>Incisivum</term><term>Insertion</term><term>Interossei</term><term>Kumbi</term><term>Labial</term><term>Labial cingulum</term><term>Lartet</term><term>Lateral</term><term>Lateral side</term><term>Lateral view</term><term>Laterally</term><term>Lholi</term><term>Lingual</term><term>Lingual bridge</term><term>Lingual cingulum</term><term>Lingual groove</term><term>Linnaeus</term><term>Linnean</term><term>Linnean society</term><term>Lower cheek teeth</term><term>Lower molars</term><term>Lower premolars</term><term>Lydekker</term><term>Mammal</term><term>Mammalia</term><term>Mandible</term><term>Mandibulae</term><term>Mandibular</term><term>Maxilla</term><term>Mcii</term><term>Mciii</term><term>Medial</term><term>Medial side</term><term>Median</term><term>Medifossette</term><term>Mesaceratherium</term><term>Mesaceratherium welcommi</term><term>Mesocuneiform</term><term>Mesostyle</term><term>Metacarpal</term><term>Metacone</term><term>Metaconid</term><term>Metaloph</term><term>Metapodials</term><term>Mhnt</term><term>Minutum</term><term>Miocene</term><term>Miocene rhinocerotids</term><term>Mirallesi</term><term>Mnhn</term><term>Molar</term><term>Monophyletic</term><term>Mtii</term><term>Mtiii</term><term>Mtiv</term><term>Natural history</term><term>Naturelle</term><term>Navicular</term><term>Node</term><term>Oblique</term><term>Occlusal</term><term>Occlusal view</term><term>Oligocene</term><term>Osborni</term><term>Outgroup</term><term>Paracone</term><term>Paralophid</term><term>Patella</term><term>Paulhiacense</term><term>Perissodactyla</term><term>Phylogenetic</term><term>Plesiaceratherium</term><term>Pleuroceros</term><term>Pleuroceros blanfordi</term><term>Pmnh</term><term>Postcranial</term><term>Postcranials</term><term>Posterior</term><term>Posterior border</term><term>Posterior part</term><term>Posterior tuberosity</term><term>Posterior valley</term><term>Posterolateral</term><term>Posteroproximal</term><term>Praecursor</term><term>Premolar</term><term>Processus</term><term>Processus zygomaticus</term><term>Prosantorhinus</term><term>Protaceratherium</term><term>Protocone</term><term>Protoloph</term><term>Prox</term><term>Proximal</term><term>Proximal border</term><term>Proximal view</term><term>Reversion</term><term>Rhinocerotid</term><term>Rhinocerotidae</term><term>Rhinocerotids</term><term>Rhinocerotina</term><term>Rhinocerotinae</term><term>Right astragalus</term><term>Right cuboid</term><term>Right scaphoid</term><term>Robust</term><term>Ronzotherium</term><term>Sagittal</term><term>Sagittally</term><term>Scaphoid</term><term>Semilunate</term><term>Sensu</term><term>Simorrense</term><term>Squamosal</term><term>Square brackets</term><term>Stratigraphical</term><term>Suprageneric</term><term>Symphysis</term><term>Synapomorphies</term><term>Talus</term><term>Taxon</term><term>Tibia</term><term>Transverse diameter</term><term>Trapezoid</term><term>Trigonias</term><term>Trigonid</term><term>Trochlea</term><term>Tuber</term><term>Tubercle</term><term>Tuberosity</term><term>Ulna</term><term>Unciform</term><term>Upper member</term><term>Upper molars</term><term>Upper premolars</term><term>Welcomme</term><term>Welcommi</term><term>Zinda</term><term>Zoological</term><term>Zoological journal</term><term>Zygomatic</term><term>Zygomatic arch</term><term>Zygomaticus</term></keywords><keywords scheme="Teeft" xml:lang="en"><term>Aceratheriini</term><term>Aceratherium</term><term>Albigense</term><term>Alicornops</term><term>Antecrochet</term><term>Anterior</term><term>Anterior border</term><term>Anterior side</term><term>Anterior view</term><term>Anteroposterior</term><term>Anteroposterior diameter</term><term>Antoine</term><term>Antoine welcomme</term><term>Apophysis</term><term>Approximate dimensions</term><term>Articular</term><term>Articulation</term><term>Astragalus</term><term>Attened</term><term>Balochistan</term><term>Bicornis</term><term>Blanfordi</term><term>Bonis</term><term>Brachypotherium</term><term>Brachypus</term><term>Bracket</term><term>Browni</term><term>Bugti</term><term>Bugti hills</term><term>Bula</term><term>Calcaneum</term><term>Calcaneus</term><term>Carpalis</term><term>Cheek teeth</term><term>Chilotherium</term><term>Chitarwata</term><term>Cingulum</term><term>Clade</term><term>Coeval</term><term>Constricted</term><term>Convex</term><term>Corpus mandibulae</term><term>Cranial</term><term>Crista</term><term>Cuboid</term><term>Deciduous</term><term>Dentition</term><term>Dera</term><term>Dera bugti</term><term>Diaceratherium</term><term>Diaphysis</term><term>Diceratherium</term><term>Dicerorhinus</term><term>Diceros</term><term>Dihedron</term><term>Distal articulation</term><term>Distal border</term><term>Distal view</term><term>Dorsal</term><term>Dorsal view</term><term>Douvillei</term><term>Duvernoy</term><term>Earliest miocene</term><term>Early miocene</term><term>Ectocuneiform</term><term>Ectoloph</term><term>Ectometaloph</term><term>Elasmotheriina</term><term>Elasmotheriinae</term><term>Entoconid</term><term>Extensor carpalis</term><term>External groove</term><term>Facet</term><term>Fauna</term><term>Femur</term><term>Foramen</term><term>Fossil</term><term>Gaimersheimense</term><term>Gaindatherium</term><term>Gand</term><term>Gers</term><term>Ginsburg</term><term>Groove</term><term>Gutter</term><term>Heissig</term><term>Homoplastic</term><term>Hooijer</term><term>Humerus</term><term>Hypocone</term><term>Hypolophid</term><term>Incisivum</term><term>Insertion</term><term>Interossei</term><term>Kumbi</term><term>Labial</term><term>Labial cingulum</term><term>Lartet</term><term>Lateral</term><term>Lateral side</term><term>Lateral view</term><term>Laterally</term><term>Lholi</term><term>Lingual</term><term>Lingual bridge</term><term>Lingual cingulum</term><term>Lingual groove</term><term>Linnaeus</term><term>Linnean</term><term>Linnean society</term><term>Lower cheek teeth</term><term>Lower molars</term><term>Lower premolars</term><term>Lydekker</term><term>Mammal</term><term>Mammalia</term><term>Mandible</term><term>Mandibulae</term><term>Mandibular</term><term>Maxilla</term><term>Mcii</term><term>Mciii</term><term>Medial</term><term>Medial side</term><term>Median</term><term>Medifossette</term><term>Mesaceratherium</term><term>Mesaceratherium welcommi</term><term>Mesocuneiform</term><term>Mesostyle</term><term>Metacarpal</term><term>Metacone</term><term>Metaconid</term><term>Metaloph</term><term>Metapodials</term><term>Mhnt</term><term>Minutum</term><term>Miocene</term><term>Miocene rhinocerotids</term><term>Mirallesi</term><term>Mnhn</term><term>Molar</term><term>Monophyletic</term><term>Mtii</term><term>Mtiii</term><term>Mtiv</term><term>Natural history</term><term>Naturelle</term><term>Navicular</term><term>Node</term><term>Oblique</term><term>Occlusal</term><term>Occlusal view</term><term>Oligocene</term><term>Osborni</term><term>Outgroup</term><term>Paracone</term><term>Paralophid</term><term>Patella</term><term>Paulhiacense</term><term>Perissodactyla</term><term>Phylogenetic</term><term>Plesiaceratherium</term><term>Pleuroceros</term><term>Pleuroceros blanfordi</term><term>Pmnh</term><term>Postcranial</term><term>Postcranials</term><term>Posterior</term><term>Posterior border</term><term>Posterior part</term><term>Posterior tuberosity</term><term>Posterior valley</term><term>Posterolateral</term><term>Posteroproximal</term><term>Praecursor</term><term>Premolar</term><term>Processus</term><term>Processus zygomaticus</term><term>Prosantorhinus</term><term>Protaceratherium</term><term>Protocone</term><term>Protoloph</term><term>Prox</term><term>Proximal</term><term>Proximal border</term><term>Proximal view</term><term>Reversion</term><term>Rhinocerotid</term><term>Rhinocerotidae</term><term>Rhinocerotids</term><term>Rhinocerotina</term><term>Rhinocerotinae</term><term>Right astragalus</term><term>Right cuboid</term><term>Right scaphoid</term><term>Robust</term><term>Ronzotherium</term><term>Sagittal</term><term>Sagittally</term><term>Scaphoid</term><term>Semilunate</term><term>Sensu</term><term>Simorrense</term><term>Squamosal</term><term>Square brackets</term><term>Stratigraphical</term><term>Suprageneric</term><term>Symphysis</term><term>Synapomorphies</term><term>Talus</term><term>Taxon</term><term>Tibia</term><term>Transverse diameter</term><term>Trapezoid</term><term>Trigonias</term><term>Trigonid</term><term>Trochlea</term><term>Tuber</term><term>Tubercle</term><term>Tuberosity</term><term>Ulna</term><term>Unciform</term><term>Upper member</term><term>Upper molars</term><term>Upper premolars</term><term>Welcomme</term><term>Welcommi</term><term>Zinda</term><term>Zoological</term><term>Zoological journal</term><term>Zygomatic</term><term>Zygomatic arch</term><term>Zygomaticus</term></keywords></textClass></profileDesc></teiHeader><front><div type="abstract">Rhinocerotids are particularly abundant and diversified in Neogene deposits of the Indian subcontinent, but their systematics is far from being well defined. Based on the revision of old collections and new findings from the Early Miocene of the Bugti Hills and Zinda Pir, Pakistan, ‘Aceratherium blanfordi Lydekker, 1884’ is a chimera, consisting of two dentally convergent but postcranially distinct rhinocerotid taxa: Pleuroceros blanfordi and Mesaceratherium welcommi sp. nov. Postcranial features appear to be much more diagnostic than craniodental morphology in this case. A phylogenetic analysis based on 282 morphological characters scored for 28 taxa (four outgroups and ingroup including both taxa of interest and a ‘branching group’) strengthens this statement and supports Pleuroceros and Mesaceratherium as monophyletic genera within Rhinocerotinae. Both genera are recognized for the first time outside Europe. In the Bugti Hills, P. blanfordi and M. welcommi are part of an exceptionally diversified rhinocerotid fauna, with up to nine species associated in the same locality (Kumbi 4f). This rhinocerotid assemblage confirms the earliest Miocene age (Agenian/Aquitanian) of the upper member of the Chitarwata Formation as a whole. Coeval homotaxic rhinocerotid faunas from Europe (France, Czech Republic) and East Africa (Uganda, Kenya) support broad and sustainable rhinocerotid interchanges amongst South Asia, Europe, and Africa under compatible environmental conditions throughout earliest Miocene times.</div></front></TEI><istex><corpusName>wiley</corpusName><keywords><teeft><json:string>mhnt</json:string><json:string>miocene</json:string><json:string>linnean society</json:string><json:string>linnean</json:string><json:string>bugti</json:string><json:string>blanfordi</json:string><json:string>pleuroceros</json:string><json:string>lingual</json:string><json:string>mesaceratherium</json:string><json:string>early miocene</json:string><json:string>kumbi</json:string><json:string>welcommi</json:string><json:string>cingulum</json:string><json:string>zoological</json:string><json:string>medial</json:string><json:string>zoological journal</json:string><json:string>lydekker</json:string><json:string>bugti hills</json:string><json:string>node</json:string><json:string>taxon</json:string><json:string>astragalus</json:string><json:string>premolar</json:string><json:string>anterior view</json:string><json:string>diaphysis</json:string><json:string>rhinocerotids</json:string><json:string>protocone</json:string><json:string>balochistan</json:string><json:string>welcomme</json:string><json:string>hypocone</json:string><json:string>tuberosity</json:string><json:string>lateral</json:string><json:string>zinda</json:string><json:string>processus</json:string><json:string>mtiii</json:string><json:string>pleuroceros blanfordi</json:string><json:string>labial</json:string><json:string>tubercle</json:string><json:string>upper molars</json:string><json:string>symphysis</json:string><json:string>earliest miocene</json:string><json:string>heissig</json:string><json:string>mesaceratherium welcommi</json:string><json:string>dera</json:string><json:string>aceratherium</json:string><json:string>postcranial</json:string><json:string>sagittally</json:string><json:string>rhinocerotidae</json:string><json:string>miocene rhinocerotids</json:string><json:string>molar</json:string><json:string>articular</json:string><json:string>mandible</json:string><json:string>anteroposterior</json:string><json:string>occlusal view</json:string><json:string>scaphoid</json:string><json:string>occlusal</json:string><json:string>anteroposterior diameter</json:string><json:string>protaceratherium</json:string><json:string>anterior side</json:string><json:string>mcii</json:string><json:string>proximal</json:string><json:string>tibia</json:string><json:string>ectoloph</json:string><json:string>transverse diameter</json:string><json:string>reversion</json:string><json:string>metaloph</json:string><json:string>labial cingulum</json:string><json:string>rhinocerotid</json:string><json:string>metacone</json:string><json:string>lateral view</json:string><json:string>calcaneus</json:string><json:string>antecrochet</json:string><json:string>dera bugti</json:string><json:string>foramen</json:string><json:string>proximal view</json:string><json:string>mandibular</json:string><json:string>lower cheek teeth</json:string><json:string>lower premolars</json:string><json:string>approximate dimensions</json:string><json:string>semilunate</json:string><json:string>phylogenetic</json:string><json:string>unciform</json:string><json:string>oligocene</json:string><json:string>protoloph</json:string><json:string>trochlea</json:string><json:string>dorsal</json:string><json:string>paulhiacense</json:string><json:string>square brackets</json:string><json:string>mammalia</json:string><json:string>lingual cingulum</json:string><json:string>bracket</json:string><json:string>chitarwata</json:string><json:string>distal view</json:string><json:string>homoplastic</json:string><json:string>robust</json:string><json:string>naturelle</json:string><json:string>upper premolars</json:string><json:string>synapomorphies</json:string><json:string>proximal border</json:string><json:string>maxilla</json:string><json:string>sagittal</json:string><json:string>plesiaceratherium</json:string><json:string>brachypotherium</json:string><json:string>posterior tuberosity</json:string><json:string>articulation</json:string><json:string>insertion</json:string><json:string>gaimersheimense</json:string><json:string>chilotherium</json:string><json:string>cuboid</json:string><json:string>clade</json:string><json:string>entoconid</json:string><json:string>prox</json:string><json:string>mnhn</json:string><json:string>mtiv</json:string><json:string>distal articulation</json:string><json:string>lower molars</json:string><json:string>pmnh</json:string><json:string>calcaneum</json:string><json:string>squamosal</json:string><json:string>navicular</json:string><json:string>mesostyle</json:string><json:string>prosantorhinus</json:string><json:string>dorsal view</json:string><json:string>duvernoy</json:string><json:string>rhinocerotinae</json:string><json:string>mandibulae</json:string><json:string>facet</json:string><json:string>anterior</json:string><json:string>antoine</json:string><json:string>medial side</json:string><json:string>cranial</json:string><json:string>zygomaticus</json:string><json:string>metaconid</json:string><json:string>alicornops</json:string><json:string>outgroup</json:string><json:string>metacarpal</json:string><json:string>dicerorhinus</json:string><json:string>femur</json:string><json:string>sensu</json:string><json:string>fossil</json:string><json:string>groove</json:string><json:string>hooijer</json:string><json:string>posteroproximal</json:string><json:string>crista</json:string><json:string>natural history</json:string><json:string>bonis</json:string><json:string>interossei</json:string><json:string>elasmotheriina</json:string><json:string>apophysis</json:string><json:string>mciii</json:string><json:string>ulna</json:string><json:string>posterior</json:string><json:string>ectometaloph</json:string><json:string>praecursor</json:string><json:string>carpalis</json:string><json:string>extensor carpalis</json:string><json:string>lateral side</json:string><json:string>deciduous</json:string><json:string>tuber</json:string><json:string>lingual groove</json:string><json:string>posterior valley</json:string><json:string>patella</json:string><json:string>zygomatic</json:string><json:string>paracone</json:string><json:string>diceratherium</json:string><json:string>perissodactyla</json:string><json:string>corpus mandibulae</json:string><json:string>ginsburg</json:string><json:string>upper member</json:string><json:string>convex</json:string><json:string>zygomatic arch</json:string><json:string>coeval</json:string><json:string>browni</json:string><json:string>diaceratherium</json:string><json:string>minutum</json:string><json:string>right astragalus</json:string><json:string>albigense</json:string><json:string>rhinocerotina</json:string><json:string>mirallesi</json:string><json:string>posterior part</json:string><json:string>lingual bridge</json:string><json:string>dihedron</json:string><json:string>aceratheriini</json:string><json:string>trigonid</json:string><json:string>ectocuneiform</json:string><json:string>mtii</json:string><json:string>gaindatherium</json:string><json:string>humerus</json:string><json:string>metapodials</json:string><json:string>laterally</json:string><json:string>diceros</json:string><json:string>distal border</json:string><json:string>posterolateral</json:string><json:string>right scaphoid</json:string><json:string>simorrense</json:string><json:string>bicornis</json:string><json:string>incisivum</json:string><json:string>talus</json:string><json:string>postcranials</json:string><json:string>oblique</json:string><json:string>fauna</json:string><json:string>stratigraphical</json:string><json:string>osborni</json:string><json:string>paralophid</json:string><json:string>hypolophid</json:string><json:string>mesocuneiform</json:string><json:string>trigonias</json:string><json:string>douvillei</json:string><json:string>cheek teeth</json:string><json:string>attened</json:string><json:string>brachypus</json:string><json:string>lartet</json:string><json:string>anterior border</json:string><json:string>antoine welcomme</json:string><json:string>posterior border</json:string><json:string>lholi</json:string><json:string>bula</json:string><json:string>gers</json:string><json:string>linnaeus</json:string><json:string>monophyletic</json:string><json:string>suprageneric</json:string><json:string>right cuboid</json:string><json:string>dentition</json:string><json:string>medifossette</json:string><json:string>elasmotheriinae</json:string><json:string>ronzotherium</json:string><json:string>processus zygomaticus</json:string><json:string>external groove</json:string><json:string>gand</json:string><json:string>constricted</json:string><json:string>mammal</json:string><json:string>median</json:string><json:string>trapezoid</json:string><json:string>gutter</json:string><json:string>postfossette</json:string><json:string>eximius</json:string><json:string>carpus</json:string><json:string>occidentalis</json:string><json:string>aceratherium blanfordi</json:string><json:string>terrestris</json:string><json:string>hyrachyus</json:string><json:string>national naturelle</json:string><json:string>osborn</json:string><json:string>armatum</json:string><json:string>proximal facet</json:string><json:string>ectolophid</json:string><json:string>continuous lingual cingulum</json:string><json:string>incisor</json:string><json:string>tapirus</json:string><json:string>bugtirhinus</json:string><json:string>ucbl</json:string><json:string>beonense</json:string><json:string>aprotodon</json:string><json:string>arikarense</json:string><json:string>hispanotherium</json:string><json:string>anderssoni</json:string><json:string>posteriorly</json:string><json:string>nuchal</json:string><json:string>posterior cingulum</json:string><json:string>right mtiii</json:string><json:string>rhinocerotini</json:string><json:string>sumatrensis</json:string><json:string>bugti area</json:string><json:string>teleoceratina</json:string><json:string>fossiles</json:string><json:string>leidy</json:string><json:string>distal</json:string><json:string>pakistan</json:string><json:string>extensor</json:string><json:string>toulouse</json:string><json:string>alveolus</json:string><json:string>medial view</json:string><json:string>lingual side</json:string><json:string>trigonias osborni</json:string><json:string>medial border</json:string><json:string>lateral metapodials</json:string><json:string>phylogenetic analysis</json:string><json:string>lingual fragment</json:string><json:string>right unciform</json:string><json:string>consistency index retention index</json:string><json:string>diaphysis dist</json:string><json:string>intermediate relief</json:string><json:string>chitarwata formation</json:string><json:string>right mtiv</json:string><json:string>right radius</json:string><json:string>right calcaneus</json:string><json:string>natural history museum</json:string><json:string>sister group</json:string><json:string>rhinoceros</json:string><json:string>enamel</json:string><json:string>genus</json:string><json:string>pyramidal</json:string><json:string>mesaceratherium heissig</json:string><json:string>foramen mentale</json:string><json:string>phylogenetic relationships</json:string><json:string>upper cheek teeth</json:string><json:string>protaceratherium albigense</json:string><json:string>postcranial skeleton</json:string><json:string>medial tubercle</json:string><json:string>late miocene</json:string><json:string>collum tali</json:string><json:string>type species</json:string><json:string>processus postorbitalis</json:string><json:string>plesiaceratherium mirallesi</json:string><json:string>pleuroceros pleuroceros</json:string><json:string>upper teeth</json:string><json:string>aquitaine basin</json:string><json:string>articular facets</json:string><json:string>right tibia</json:string><json:string>alicornops simorrense</json:string><json:string>lateral border</json:string><json:string>posterior expansion</json:string><json:string>mandibular symphysis</json:string><json:string>mesaceratherium paulhiacense</json:string><json:string>unique reversion</json:string><json:string>hyrachyus eximius</json:string><json:string>posterior side</json:string><json:string>geological survey</json:string><json:string>lingual wall</json:string><json:string>median valley</json:string><json:string>unique reversions</json:string><json:string>protaceratherium minutum</json:string><json:string>fragment</json:string><json:string>transverse</json:string><json:string>lindsay</json:string></teeft></keywords><author><json:item><name>PIERRE‐OLIVIER ANTOINE</name><affiliations><json:string>Université de Toulouse, UPS (SVT‐OMP), LMTG, 14 Avenue Édouard Belin, F‐31400 Toulouse, France</json:string><json:string>CNRS, LMTG, 14 Avenue Édouard Belin, F‐31400 Toulouse, France</json:string><json:string>IRD, LMTG, 14 Avenue Édouard Belin, F‐31400 Toulouse, France</json:string><json:string>E-mail: poa@lmtg.obs‐mip.fr</json:string></affiliations></json:item><json:item><name>KEVIN F. DOWNING</name><affiliations><json:string>DePaul University, Chicago, IL 60604, USA</json:string></affiliations></json:item><json:item><name>JEAN‐YVES CROCHET</name><affiliations><json:string>Département de Paléontologie, Institut des Sciences de l'Évolution (CNRS‐UMR 5554) Université Montpellier 2, c.c. 064, Place Eugène Bataillon, F‐34095 Montpellier Cedex 5, France</json:string></affiliations></json:item><json:item><name>FRANCIS DURANTHON</name><affiliations><json:string>Laboratoire d'Étude et de Conservation du Patrimoine, Muséum d'Histoire naturelle de Toulouse, 39 allées Jules Guesde, F‐31000 Toulouse, France</json:string></affiliations></json:item><json:item><name>LAWRENCE J. FLYNN</name><affiliations><json:string>Department of Anthropology and Peabody Museum, Harvard University, Cambridge, MA 02138, USA</json:string></affiliations></json:item><json:item><name>LAURENT MARIVAUX</name><affiliations><json:string>Département de Paléontologie, Institut des Sciences de l'Évolution (CNRS‐UMR 5554) Université Montpellier 2, c.c. 064, Place Eugène Bataillon, F‐34095 Montpellier Cedex 5, France</json:string></affiliations></json:item><json:item><name>GRÉGOIRE MÉTAIS</name><affiliations><json:string>Laboratoire de Paléontologie, UMR 5143 CNRS, Muséum National d'Histoire Naturelle, 8 rue Buffon, F‐75005 Paris, France</json:string></affiliations></json:item><json:item><name>ABDUL RAHIM RAJPAR</name><affiliations><json:string>9Earth Sciences Division, Pakistan Museum of Natural History, Garden Avenue, Shakarparian, 44000 Islamabad, Pakistan</json:string></affiliations></json:item><json:item><name>GHAZALA ROOHI</name><affiliations><json:string>9Earth Sciences Division, Pakistan Museum of Natural History, Garden Avenue, Shakarparian, 44000 Islamabad, Pakistan</json:string></affiliations></json:item></author><subject><json:item><lang><json:string>eng</json:string></lang><value>Bugti Hills</value></json:item><json:item><lang><json:string>eng</json:string></lang><value>Chitarwata Formation</value></json:item><json:item><lang><json:string>eng</json:string></lang><value>cladistics</value></json:item><json:item><lang><json:string>eng</json:string></lang><value>Indian subcontinent</value></json:item><json:item><lang><json:string>eng</json:string></lang><value>Mesaceratherium</value></json:item><json:item><lang><json:string>eng</json:string></lang><value>palaeobiogeography</value></json:item><json:item><lang><json:string>eng</json:string></lang><value>Perissodactyla</value></json:item><json:item><lang><json:string>eng</json:string></lang><value>Pleuroceros</value></json:item><json:item><lang><json:string>eng</json:string></lang><value>Zinda Pir</value></json:item></subject><articleId><json:string>ZOJ597</json:string></articleId><arkIstex>ark:/67375/WNG-H9V547SV-F</arkIstex><language><json:string>eng</json:string></language><originalGenre><json:string>article</json:string></originalGenre><abstract>Rhinocerotids are particularly abundant and diversified in Neogene deposits of the Indian subcontinent, but their systematics is far from being well defined. Based on the revision of old collections and new findings from the Early Miocene of the Bugti Hills and Zinda Pir, Pakistan, ‘Aceratherium blanfordi Lydekker, 1884’ is a chimera, consisting of two dentally convergent but postcranially distinct rhinocerotid taxa: Pleuroceros blanfordi and Mesaceratherium welcommi sp. nov. Postcranial features appear to be much more diagnostic than craniodental morphology in this case. A phylogenetic analysis based on 282 morphological characters scored for 28 taxa (four outgroups and ingroup including both taxa of interest and a ‘branching group’) strengthens this statement and supports Pleuroceros and Mesaceratherium as monophyletic genera within Rhinocerotinae. Both genera are recognized for the first time outside Europe. In the Bugti Hills, P. blanfordi and M. welcommi are part of an exceptionally diversified rhinocerotid fauna, with up to nine species associated in the same locality (Kumbi 4f). This rhinocerotid assemblage confirms the earliest Miocene age (Agenian/Aquitanian) of the upper member of the Chitarwata Formation as a whole. Coeval homotaxic rhinocerotid faunas from Europe (France, Czech Republic) and East Africa (Uganda, Kenya) support broad and sustainable rhinocerotid interchanges amongst South Asia, Europe, and Africa under compatible environmental conditions throughout earliest Miocene times.</abstract><qualityIndicators><score>9.556</score><pdfWordCount>26872</pdfWordCount><pdfCharCount>181459</pdfCharCount><pdfVersion>1.3</pdfVersion><pdfPageCount>56</pdfPageCount><pdfPageSize>595.276 x 782.362 pts</pdfPageSize><refBibsNative>true</refBibsNative><abstractWordCount>213</abstractWordCount><abstractCharCount>1524</abstractCharCount><keywordCount>9</keywordCount></qualityIndicators><title>A revision of Aceratherium blanfordi Lydekker, 1884 (Mammalia: Rhinocerotidae) from the Early Miocene of Pakistan: postcranials as a key</title><genre><json:string>article</json:string></genre><host><title>Zoological Journal of the Linnean Society</title><language><json:string>unknown</json:string></language><doi><json:string>10.1111/(ISSN)1096-3642</json:string></doi><issn><json:string>0024-4082</json:string></issn><eissn><json:string>1096-3642</json:string></eissn><publisherId><json:string>ZOJ</json:string></publisherId><volume>160</volume><issue>1</issue><pages><first>139</first><last>194</last><total>56</total></pages><genre><json:string>journal</json:string></genre></host><namedEntities><unitex><date><json:string>1910</json:string><json:string>1995</json:string><json:string>2010</json:string><json:string>2004</json:string></date><geogName></geogName><orgName><json:string>France, Czech Republic</json:string><json:string>Pakistan Received</json:string><json:string>International Commission On Zoological Nomenclature</json:string><json:string>Uganda, Kenya</json:string><json:string>FROM PAKISTAN Table</json:string><json:string>Claude-Bernard University</json:string><json:string>Pakistan Museum of Natural History</json:string><json:string>Department of Anthropology</json:string><json:string>Harvard University</json:string><json:string>Pakistan Museum of Natural History, Islamabad</json:string><json:string>American Museum of Natural History</json:string><json:string>Kenya and Uganda</json:string><json:string>Pakistan/ East Africa</json:string><json:string>Pakistan Museum of Natural History, Garden Avenue, Shakarparian</json:string></orgName><orgName_funder></orgName_funder><orgName_provider></orgName_provider><persName><json:string>M. A. K. Bugti</json:string><json:string>Sophie Hervet</json:string><json:string>Christine Argot</json:string><json:string>Léonard Ginsburg</json:string><json:string>Claire Sagne</json:string><json:string>Jules Guesde</json:string><json:string>Miocene</json:string><json:string>John C. Barry</json:string><json:string>H. Lindsay</json:string><json:string>Jean-Loup Welcomme</json:string><json:string>Sulaiman Lobe</json:string><json:string>L. Welcomme</json:string><json:string>Mouloud Benammi</json:string><json:string>Édouard Belin</json:string><json:string>Sulaiman Range</json:string><json:string>Peter J. Whybrow</json:string><json:string>Annie Antoine</json:string><json:string>Jean-Jacques Jaeger</json:string><json:string>Peter Hayward</json:string><json:string>Jérôme Proriol</json:string><json:string>David Pilbeam</json:string><json:string>Jerry J. Hooker</json:string><json:string>Marc Delcorso</json:string><json:string>Kamal Majidullah</json:string><json:string>Dario De Franceschi</json:string><json:string>U. Cheema</json:string><json:string>Time Scale</json:string><json:string>Malcolm C. McKenna</json:string><json:string>Will Downs</json:string><json:string>Pascal Tassy</json:string><json:string>Andy Currant</json:string><json:string>Michèle E. Morgan</json:string><json:string>H. E. Iqbal</json:string><json:string>Hans Larsson</json:string><json:string>Susanne Cote</json:string><json:string>Q. Comparisons</json:string><json:string>Abel Prieur</json:string><json:string>Pir</json:string></persName><placeName><json:string>Pakistan</json:string><json:string>Gaj</json:string><json:string>IL</json:string><json:string>Paris</json:string><json:string>Montpellier</json:string><json:string>Munich</json:string><json:string>Islamabad</json:string><json:string>Europe</json:string><json:string>America</json:string><json:string>Czech Republic</json:string><json:string>Toulouse</json:string><json:string>Chicago</json:string><json:string>York</json:string><json:string>MA</json:string><json:string>France</json:string><json:string>Karachi</json:string><json:string>Cambridge</json:string><json:string>Lyon</json:string><json:string>PAKISTAN</json:string></placeName><ref_url></ref_url><ref_bibl><json:string>MHNT Pak 1190</json:string><json:string>MHNT Pak 1026, 1047</json:string><json:string>MHNT Pak 1590</json:string><json:string>MHNT Pak 1990</json:string><json:string>MHNT Pak 1156</json:string><json:string>Répelin, 1917</json:string><json:string>MHNT Pak 1435</json:string><json:string>MHNT Pak 1139</json:string><json:string>Antoine et al., 2003a</json:string><json:string>Crusafont et al., 1955</json:string><json:string>Guérin & Pickford, 2003</json:string><json:string>PMNH Z2269; Downing, 2005: 3</json:string><json:string>[4]</json:string><json:string>MHNT Pak 1480</json:string><json:string>MHNT Pak 1063</json:string><json:string>Lindsay et al. (2005)</json:string><json:string>MHNT Pak 1100</json:string><json:string>MHNT Pak 1167</json:string><json:string>Farris, 1988</json:string><json:string>Mihlbachler, 2005</json:string><json:string>MHNT Pak 1967</json:string><json:string>MHNT Pak 1091</json:string><json:string>Antoine et al., 2008</json:string><json:string>MHNT Pak 1195</json:string><json:string>MHNT Pak 1111</json:string><json:string>[12]</json:string><json:string>Collinson & Hooker, 2003</json:string><json:string>Cuvier, 1822</json:string><json:string>Roman, 1912</json:string><json:string>MHNT Pak 1260</json:string><json:string>MHNT Pak 1085</json:string><json:string>MHNT Pak 1068</json:string><json:string>MHNT Pak 1868</json:string><json:string>MHNT Pak 1150</json:string><json:string>MHNT Pak 1096</json:string><json:string>Pal 1198</json:string><json:string>Barry et al., 2002</json:string><json:string>[6]</json:string><json:string>Antoine et al., 2003a, b, 2004</json:string><json:string>MHNT Pak 1040</json:string><json:string>MHNT Pak 1144</json:string><json:string>Wiens, 2001: 690</json:string><json:string>MHNT Pak 2458</json:string><json:string>MHNT Pak 1648</json:string><json:string>MHNT Pak 1051</json:string><json:string>MHNT Pak 1676</json:string><json:string>MHNT Pak 1138</json:string><json:string>Antoine et al. 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(2006)</json:string><json:string>MHNT Pak 1143</json:string><json:string>MHNT Pak 1089</json:string><json:string>MHNT Pak 2215</json:string><json:string>MHNT Pak 1109</json:string><json:string>MHNT Pak 1194, 1195</json:string><json:string>Heissig & Fejfar, 2007: 19</json:string><json:string>MHNT Pak 1050</json:string><json:string>Welcomme et al., 1997, 1999, 2001</json:string><json:string>MHNT Pak 1033</json:string><json:string>in 29 specimens of the recent rhinocerotine Diceros bicornis; Guérin, 1980: 131</json:string><json:string>MHNT Pak 1061</json:string><json:string>MHNT Pak 1044</json:string><json:string>MHNT Pak 1444</json:string><json:string>MHNT Pak 1027</json:string><json:string>Gentry, 1987</json:string><json:string>Freeland, 1991</json:string><json:string>Antoine et al., 2006</json:string><json:string>MHNT Pak 1193</json:string><json:string>Welcomme et al., 1997, 2001</json:string><json:string>Laudet & Antoine, 2004</json:string><json:string>MHNT Pak 1159</json:string><json:string>MHNT Pak 1438</json:string><json:string>Antoine, 1997</json:string><json:string>Blanford, 1876, 1879</json:string><json:string>MHNT Pak 1085, 1198</json:string><json:string>[3]</json:string><json:string>MHNT Pak 1120</json:string><json:string>MHNT Pak 1066</json:string><json:string>MHNT Pak 1103</json:string><json:string>MHNT Pak 1049</json:string><json:string>MHNT Pak 760, 1012a, 1045, 1258</json:string><json:string>Lydekker, 1881, 1884</json:string><json:string>Barbour, 1906</json:string><json:string>MHNT Pak 1131</json:string><json:string>MHNT Pak 2203</json:string><json:string>De Franceschi et al., 2008</json:string><json:string>Downing, 2005</json:string><json:string>Swofford, 1998</json:string><json:string>MHNT Pak 1021</json:string><json:string>MHNT Pak 1088</json:string><json:string>Lucas, 1900</json:string><json:string>MHNT Pak 1032B</json:string><json:string>Borissiak, 1944</json:string><json:string>Antoine et al., 2000</json:string><json:string>Leidy, 1851</json:string><json:string>sensu Heissig, 1972</json:string><json:string>MHNT Pak 1153</json:string><json:string>MHNT Pak 1099</json:string><json:string>MHNT Pak 1136</json:string><json:string>MHNT Pak 1032b</json:string><json:string>MHNT Pak 1119</json:string><json:string>Antoine et al. (2003b)</json:string><json:string>[5]</json:string><json:string>MHNT Pak 1060</json:string><json:string>Antoine, Duranthon & Tassy, 1997</json:string><json:string>MHNT Pak 1164</json:string><json:string>MHNT Pak 1147</json:string><json:string>MHNT Pak 1964</json:string><json:string>Fischer Von Waldheim, 1814</json:string><json:string>Guérin, 1980</json:string><json:string>MHNT Pak 1192</json:string><json:string>MHNT Pak 1054</json:string><json:string>Welcomme et al., 1997: 532</json:string><json:string>MHNT Pak 2126</json:string><json:string>MHNT Pak 1733</json:string><json:string>MHNT Pak 1437</json:string><json:string>Richard, 1937</json:string><json:string>Métais et al. 2009: 163</json:string><json:string>MHNT Pak 1065</json:string><json:string>MHNT Pak 1206</json:string><json:string>de Bonis, 1973</json:string><json:string>MHNT Pak 1093</json:string><json:string>MHNT Pak 1126, 1128</json:string><json:string>Antoine et al.</json:string><json:string>Welcomme et al., 1999, 2001</json:string><json:string>MHNT Pak 1113</json:string><json:string>MHNT Pak 1059</json:string><json:string>Lartet, 1837</json:string><json:string>[7]</json:string><json:string>Lindsay et al., 2005</json:string><json:string>Lihoreau et al., 2009</json:string><json:string>MHNT Pak 1087</json:string><json:string>Heissig, 1976</json:string><json:string>MHNT Pak 1124</json:string><json:string>MHNT Pak 1032A</json:string><json:string>MHNT Pak 1107</json:string><json:string>Forster-Cooper, 1934</json:string><json:string>Pilgrim, 1910, 1912</json:string><json:string>Antoine, 2002, 2003</json:string><json:string>MHNT Pak 1152</json:string><json:string>MHNT Pak 1098</json:string><json:string>MHNT Pak 1552</json:string><json:string>MHNT Pak 1135</json:string><json:string>MHNT Pak 1032a</json:string><json:string>Gradstein et al., 2005</json:string><json:string>Métais et al., 2009</json:string><json:string>MHNT Pak 1163</json:string><json:string>Lindsay et al.</json:string><json:string>MHNT Pak 1025</json:string><json:string>MHNT Pak 1129</json:string><json:string>FROM PAKISTAN ORDER PERISSODACTYLA OWEN, 1848</json:string><json:string>Antoine et al., 2004</json:string><json:string>MHNT Pak 1191</json:string><json:string>MHNT Pak 1991</json:string><json:string>Levels 3bis and 4; Table 30; Antoine & Welcomme, 2000</json:string><json:string>MHNT Pak 1019</json:string><json:string>Antoine et al., 2003b</json:string><json:string>MHNT Pak 1064</json:string><json:string>MHNT Pak 1101</json:string><json:string>MHNT Pak 1047</json:string><json:string>MHNT Pak 1864</json:string><json:string>sensu Antoine, 2003</json:string><json:string>MHNT Pak 1709</json:string><json:string>MHNT Pak 1092</json:string><json:string>Pilgrim, 1912</json:string><json:string>MHNT Pak 1196</json:string><json:string>Tassy, 1986</json:string><json:string>MHNT Pak 1058</json:string><json:string>Welcomme et al., 2001</json:string><json:string>MHNT Pak 1049, 1063, 1065</json:string><json:string>Pickford, 1986</json:string><json:string>MHNT Pak 1140</json:string><json:string>[2]</json:string><json:string>Guthrie, 1984</json:string><json:string>Cote et al., 2007</json:string><json:string>Osborn, 1900</json:string><json:string>Métais et al., 2009: 163</json:string><json:string>MHNT Pak 1113, 1114</json:string><json:string>Hooijer, 1966, 1971, 1973</json:string><json:string>MHNT Pak 1151</json:string><json:string>MHNT Pak 1097</json:string><json:string>Métais et al. (2009)</json:string><json:string>MHNT Pak 1134</json:string><json:string>Heissig, 1969</json:string><json:string>MHNT Pak 1117</json:string><json:string>Antoine, 2002</json:string><json:string>Antoine, Duranthon & Welcomme, 2003b</json:string><json:string>von Linnaeus, 1758</json:string><json:string>MHNT Pak 1019, 1022</json:string><json:string>MHNT Pak 1162</json:string><json:string>Beliajeva, 1954</json:string><json:string>MHNT Pak 2234</json:string><json:string>1966, 1973</json:string><json:string>MHNT Pak 1128</json:string></ref_bibl><bibl></bibl></unitex></namedEntities><ark><json:string>ark:/67375/WNG-H9V547SV-F</json:string></ark><categories><wos><json:string>1 - science</json:string><json:string>2 - zoology</json:string></wos><scienceMetrix><json:string>1 - natural sciences</json:string><json:string>2 - biology</json:string><json:string>3 - evolutionary biology</json:string></scienceMetrix><scopus><json:string>1 - Life Sciences</json:string><json:string>2 - Agricultural and Biological Sciences</json:string><json:string>3 - Animal Science and Zoology</json:string><json:string>1 - Life Sciences</json:string><json:string>2 - Agricultural and Biological Sciences</json:string><json:string>3 - Ecology, Evolution, Behavior and Systematics</json:string></scopus></categories><publicationDate>2010</publicationDate><copyrightDate>2010</copyrightDate><doi><json:string>10.1111/j.1096-3642.2009.00597.x</json:string></doi><id>3C10EB81E8129E9F8E7C93E4DEC42360A31E54B3</id><score>1</score><fulltext><json:item><extension>pdf</extension><original>true</original><mimetype>application/pdf</mimetype><uri>https://api.istex.fr/document/3C10EB81E8129E9F8E7C93E4DEC42360A31E54B3/fulltext/pdf</uri></json:item><json:item><extension>zip</extension><original>false</original><mimetype>application/zip</mimetype><uri>https://api.istex.fr/document/3C10EB81E8129E9F8E7C93E4DEC42360A31E54B3/fulltext/zip</uri></json:item><istex:fulltextTEI uri="https://api.istex.fr/document/3C10EB81E8129E9F8E7C93E4DEC42360A31E54B3/fulltext/tei"><teiHeader><fileDesc><titleStmt><title level="a" type="main">A revision of <hi rend="italic">Aceratherium blanfordi</hi> Lydekker, 1884 (Mammalia: Rhinocerotidae) from the Early Miocene of Pakistan: postcranials as a key</title></titleStmt><publicationStmt><authority>ISTEX</authority><publisher>Blackwell Publishing Ltd</publisher><pubPlace>Oxford, UK</pubPlace><availability><licence>© 2010 The Linnean Society of London</licence></availability><date type="published" when="2010-09"></date></publicationStmt><notesStmt><note type="content-type" subtype="article" source="article" scheme="https://content-type.data.istex.fr/ark:/67375/XTP-6N5SZHKN-D">article</note><note type="publication-type" subtype="journal" scheme="https://publication-type.data.istex.fr/ark:/67375/JMC-0GLKJH51-B">journal</note></notesStmt><sourceDesc><biblStruct type="article"><analytic><title level="a" type="main">A revision of <hi rend="italic">Aceratherium blanfordi</hi> Lydekker, 1884 (Mammalia: Rhinocerotidae) from the Early Miocene of Pakistan: postcranials as a key</title><title level="a" type="short">MIOCENE RHINOCEROTIDS FROM PAKISTAN</title><author xml:id="author-0000" role="corresp"><persName><forename type="first">PIERRE‐OLIVIER</forename><surname>ANTOINE</surname></persName><affiliation>Université de Toulouse, UPS (SVT‐OMP), LMTG, 14 Avenue Édouard Belin, F‐31400 Toulouse, France<address><country key="FR"></country></address></affiliation><affiliation>CNRS, LMTG, 14 Avenue Édouard Belin, F‐31400 Toulouse, France<address><country key="FR"></country></address></affiliation><affiliation>IRD, LMTG, 14 Avenue Édouard Belin, F‐31400 Toulouse, France<address><country key="FR"></country></address></affiliation><affiliation>E‐mail: poa@lmtg.obs‐mip.fr</affiliation></author><author xml:id="author-0001"><persName><forename type="first">KEVIN F.</forename><surname>DOWNING</surname></persName><affiliation>DePaul University, Chicago, IL 60604, USA<address><country key="US"></country></address></affiliation></author><author xml:id="author-0002"><persName><forename type="first">JEAN‐YVES</forename><surname>CROCHET</surname></persName><affiliation>Département de Paléontologie, Institut des Sciences de l'Évolution (CNRS‐UMR 5554) Université Montpellier 2, c.c. 064, Place Eugène Bataillon, F‐34095 Montpellier Cedex 5, France<address><country key="FR"></country></address></affiliation></author><author xml:id="author-0003"><persName><forename type="first">FRANCIS</forename><surname>DURANTHON</surname></persName><affiliation>Laboratoire d'Étude et de Conservation du Patrimoine, Muséum d'Histoire naturelle de Toulouse, 39 allées Jules Guesde, F‐31000 Toulouse, France<address><country key="FR"></country></address></affiliation></author><author xml:id="author-0004"><persName><forename type="first">LAWRENCE J.</forename><surname>FLYNN</surname></persName><affiliation>Department of Anthropology and Peabody Museum, Harvard University, Cambridge, MA 02138, USA<address><country key="US"></country></address></affiliation></author><author xml:id="author-0005"><persName><forename type="first">LAURENT</forename><surname>MARIVAUX</surname></persName><affiliation>Département de Paléontologie, Institut des Sciences de l'Évolution (CNRS‐UMR 5554) Université Montpellier 2, c.c. 064, Place Eugène Bataillon, F‐34095 Montpellier Cedex 5, France<address><country key="FR"></country></address></affiliation></author><author xml:id="author-0006"><persName><forename type="first">GRÉGOIRE</forename><surname>MÉTAIS</surname></persName><affiliation>Laboratoire de Paléontologie, UMR 5143 CNRS, Muséum National d'Histoire Naturelle, 8 rue Buffon, F‐75005 Paris, France<address><country key="FR"></country></address></affiliation></author><author xml:id="author-0007"><persName><forename type="first">ABDUL RAHIM</forename><surname>RAJPAR</surname></persName><affiliation>9Earth Sciences Division, Pakistan Museum of Natural History, Garden Avenue, Shakarparian, 44000 Islamabad, Pakistan<address><country key="PK"></country></address></affiliation></author><author xml:id="author-0008"><persName><forename type="first">GHAZALA</forename><surname>ROOHI</surname></persName><affiliation>9Earth Sciences Division, Pakistan Museum of Natural History, Garden Avenue, Shakarparian, 44000 Islamabad, Pakistan<address><country key="PK"></country></address></affiliation></author><idno type="istex">3C10EB81E8129E9F8E7C93E4DEC42360A31E54B3</idno><idno type="ark">ark:/67375/WNG-H9V547SV-F</idno><idno type="DOI">10.1111/j.1096-3642.2009.00597.x</idno><idno type="unit">ZOJ597</idno><idno type="toTypesetVersion">file:ZOJ.ZOJ597.pdf</idno></analytic><monogr><title level="j" type="main">Zoological Journal of the Linnean Society</title><title level="j" type="alt">ZOOLOGICAL JOURNAL OF THE LINNEAN SOCIETY</title><idno type="pISSN">0024-4082</idno><idno type="eISSN">1096-3642</idno><idno type="book-DOI">10.1111/(ISSN)1096-3642</idno><idno type="book-part-DOI">10.1111/zoj.2010.160.issue-1</idno><idno type="product">ZOJ</idno><idno type="publisherDivision">ST</idno><imprint><biblScope unit="vol">160</biblScope><biblScope unit="issue">1</biblScope><biblScope unit="page" from="139">139</biblScope><biblScope unit="page" to="194">194</biblScope><biblScope unit="page-count">56</biblScope><publisher>Blackwell Publishing Ltd</publisher><pubPlace>Oxford, UK</pubPlace><date type="published" when="2010-09"></date></imprint></monogr></biblStruct></sourceDesc></fileDesc><profileDesc><abstract xml:lang="en" style="main"><p>Rhinocerotids are particularly abundant and diversified in Neogene deposits of the Indian subcontinent, but their systematics is far from being well defined. Based on the revision of old collections and new findings from the Early Miocene of the Bugti Hills and Zinda Pir, Pakistan, ‘<hi rend="italic">Aceratherium blanfordi</hi> Lydekker, 1884’ is a chimera, consisting of two dentally convergent but postcranially distinct rhinocerotid taxa: <hi rend="italic">Pleuroceros blanfordi</hi> and <hi rend="bold"><hi rend="italic">Mesaceratherium welcommi</hi> sp. nov.</hi> Postcranial features appear to be much more diagnostic than craniodental morphology in this case. A phylogenetic analysis based on 282 morphological characters scored for 28 taxa (four outgroups and ingroup including both taxa of interest and a ‘branching group’) strengthens this statement and supports <hi rend="italic">Pleuroceros</hi> and <hi rend="italic">Mesaceratherium</hi> as monophyletic genera within Rhinocerotinae. Both genera are recognized for the first time outside Europe. In the Bugti Hills, <hi rend="italic">P. blanfordi</hi> and <hi rend="italic">M. welcommi</hi> are part of an exceptionally diversified rhinocerotid fauna, with up to nine species associated in the same locality (Kumbi 4f). This rhinocerotid assemblage confirms the earliest Miocene age (Agenian/Aquitanian) of the upper member of the Chitarwata Formation as a whole. Coeval homotaxic rhinocerotid faunas from Europe (France, Czech Republic) and East Africa (Uganda, Kenya) support broad and sustainable rhinocerotid interchanges amongst South Asia, Europe, and Africa under compatible environmental conditions throughout earliest Miocene times.</p><p>© 2010 The Linnean Society of London, <hi rend="italic">Zoological Journal of the Linnean Society</hi>, 2010, <hi rend="bold">160</hi>, 139–194.</p></abstract><textClass><keywords xml:lang="en"><term xml:id="k1">Bugti Hills</term><term xml:id="k2">Chitarwata Formation</term><term xml:id="k3">cladistics</term><term xml:id="k4">Indian subcontinent</term><term xml:id="k5"><hi rend="italic">Mesaceratherium</hi></term><term xml:id="k6">palaeobiogeography</term><term xml:id="k7">Perissodactyla</term><term xml:id="k8"><hi rend="italic">Pleuroceros</hi></term><term xml:id="k9">Zinda Pir</term></keywords><keywords rend="tocHeading1"><term>Original Articles</term></keywords></textClass><langUsage><language ident="en"></language></langUsage></profileDesc></teiHeader></istex:fulltextTEI><json:item><extension>txt</extension><original>false</original><mimetype>text/plain</mimetype><uri>https://api.istex.fr/document/3C10EB81E8129E9F8E7C93E4DEC42360A31E54B3/fulltext/txt</uri></json:item></fulltext><metadata><istex:metadataXml wicri:clean="Wiley, elements deleted: body"><istex:xmlDeclaration>version="1.0" encoding="UTF-8" standalone="yes"</istex:xmlDeclaration><istex:document><component version="2.0" type="serialArticle" xml:lang="en"><header><publicationMeta level="product"><publisherInfo><publisherName>Blackwell Publishing Ltd</publisherName><publisherLoc>Oxford, UK</publisherLoc></publisherInfo><doi origin="wiley" registered="yes">10.1111/(ISSN)1096-3642</doi><issn type="print">0024-4082</issn><issn type="electronic">1096-3642</issn><idGroup><id type="product" value="ZOJ"></id><id type="publisherDivision" value="ST"></id></idGroup><titleGroup><title type="main" sort="ZOOLOGICAL JOURNAL OF THE LINNEAN SOCIETY">Zoological Journal of the Linnean Society</title></titleGroup></publicationMeta><publicationMeta level="part" position="09001"><doi origin="wiley">10.1111/zoj.2010.160.issue-1</doi><numberingGroup><numbering type="journalVolume" number="160">160</numbering><numbering type="journalIssue" number="1">1</numbering></numberingGroup><coverDate startDate="2010-09">September 2010</coverDate></publicationMeta><publicationMeta level="unit" type="article" position="6" status="forIssue"><doi origin="wiley">10.1111/j.1096-3642.2009.00597.x</doi><idGroup><id type="unit" value="ZOJ597"></id></idGroup><countGroup><count type="pageTotal" number="56"></count></countGroup><titleGroup><title type="tocHeading1">Original Articles</title></titleGroup><copyright>© 2010 The Linnean Society of London</copyright><eventGroup><event type="xmlConverted" agent="Converter:BPG_TO_WML3G version:2.3.16 mode:FullText" date="2010-08-18"></event><event type="firstOnline" date="2010-08-18"></event><event type="publishedOnlineFinalForm" date="2010-08-18"></event><event type="xmlConverted" agent="Converter:WILEY_ML3G_TO_WILEY_ML3GV2 version:3.8.8" date="2014-02-11"></event><event type="xmlConverted" agent="Converter:WML3G_To_WML3G version:4.1.7 mode:FullText,remove_FC" date="2014-11-04"></event></eventGroup><numberingGroup><numbering type="pageFirst" number="139">139</numbering><numbering type="pageLast" number="194">194</numbering></numberingGroup><correspondenceTo>
E‐mail: <email normalForm="poa@lmtg.obs-mip.fr">poa@lmtg.obs‐mip.fr</email></correspondenceTo><objectNameGroup><objectName elementName="appendix">Appendices</objectName></objectNameGroup><linkGroup><link type="toTypesetVersion" href="file:ZOJ.ZOJ597.pdf"></link></linkGroup></publicationMeta><contentMeta><unparsedEditorialHistory>Received 22 January 2009; accepted for publication 28 April 2009</unparsedEditorialHistory><countGroup><count type="figureTotal" number="13"></count><count type="tableTotal" number="30"></count><count type="formulaTotal" number="0"></count><count type="referenceTotal" number="98"></count><count type="wordTotal" number="30759"></count><count type="linksPubMed" number="0"></count><count type="linksCrossRef" number="0"></count></countGroup><titleGroup><title type="main">A revision of <i>Aceratherium blanfordi</i> Lydekker, 1884 (Mammalia: Rhinocerotidae) from the Early Miocene of Pakistan: postcranials as a key</title><title type="shortAuthors">P.‐O. ANTOINE <i>ET AL.</i></title><title type="short">MIOCENE RHINOCEROTIDS FROM PAKISTAN</title></titleGroup><creators><creator creatorRole="author" xml:id="cr1" affiliationRef="#a1 #a2 #a3" corresponding="yes"><personName><givenNames>PIERRE‐OLIVIER</givenNames><familyName>ANTOINE</familyName></personName></creator><creator creatorRole="author" xml:id="cr2" affiliationRef="#a4"><personName><givenNames>KEVIN F.</givenNames><familyName>DOWNING</familyName></personName></creator><creator creatorRole="author" xml:id="cr3" affiliationRef="#a5"><personName><givenNames>JEAN‐YVES</givenNames><familyName>CROCHET</familyName></personName></creator><creator creatorRole="author" xml:id="cr4" affiliationRef="#a6"><personName><givenNames>FRANCIS</givenNames><familyName>DURANTHON</familyName></personName></creator><creator creatorRole="author" xml:id="cr5" affiliationRef="#a7"><personName><givenNames>LAWRENCE J.</givenNames><familyName>FLYNN</familyName></personName></creator><creator creatorRole="author" xml:id="cr6" affiliationRef="#a5"><personName><givenNames>LAURENT</givenNames><familyName>MARIVAUX</familyName></personName></creator><creator creatorRole="author" xml:id="cr7" affiliationRef="#a8"><personName><givenNames>GRÉGOIRE</givenNames><familyName>MÉTAIS</familyName></personName></creator><creator creatorRole="author" xml:id="cr8" affiliationRef="#a9"><personName><givenNames>ABDUL RAHIM</givenNames><familyName>RAJPAR</familyName></personName></creator><creator creatorRole="author" xml:id="cr9" affiliationRef="#a9"><personName><givenNames>GHAZALA</givenNames><familyName>ROOHI</familyName></personName></creator></creators><affiliationGroup><affiliation xml:id="a1" countryCode="FR"><unparsedAffiliation>Université de Toulouse, UPS (SVT‐OMP), LMTG, 14 Avenue Édouard Belin, F‐31400 Toulouse, France</unparsedAffiliation></affiliation><affiliation xml:id="a2" countryCode="FR"><unparsedAffiliation>CNRS, LMTG, 14 Avenue Édouard Belin, F‐31400 Toulouse, France</unparsedAffiliation></affiliation><affiliation xml:id="a3" countryCode="FR"><unparsedAffiliation>IRD, LMTG, 14 Avenue Édouard Belin, F‐31400 Toulouse, France</unparsedAffiliation></affiliation><affiliation xml:id="a4" countryCode="US"><unparsedAffiliation>DePaul University, Chicago, IL 60604, USA</unparsedAffiliation></affiliation><affiliation xml:id="a5" countryCode="FR"><unparsedAffiliation>Département de Paléontologie, Institut des Sciences de l'Évolution (CNRS‐UMR 5554) Université Montpellier 2, c.c. 064, Place Eugène Bataillon, F‐34095 Montpellier Cedex 5, France</unparsedAffiliation></affiliation><affiliation xml:id="a6" countryCode="FR"><unparsedAffiliation>Laboratoire d'Étude et de Conservation du Patrimoine, Muséum d'Histoire naturelle de Toulouse, 39 allées Jules Guesde, F‐31000 Toulouse, France</unparsedAffiliation></affiliation><affiliation xml:id="a7" countryCode="US"><unparsedAffiliation>Department of Anthropology and Peabody Museum, Harvard University, Cambridge, MA 02138, USA</unparsedAffiliation></affiliation><affiliation xml:id="a8" countryCode="FR"><unparsedAffiliation>Laboratoire de Paléontologie, UMR 5143 CNRS, Muséum National d'Histoire Naturelle, 8 rue Buffon, F‐75005 Paris, France</unparsedAffiliation></affiliation><affiliation xml:id="a9" countryCode="PK"><unparsedAffiliation><sup>9</sup>Earth Sciences Division, Pakistan Museum of Natural History, Garden Avenue, Shakarparian, 44000 Islamabad, Pakistan</unparsedAffiliation></affiliation></affiliationGroup><keywordGroup xml:lang="en"><keyword xml:id="k1">Bugti Hills</keyword><keyword xml:id="k2">Chitarwata Formation</keyword><keyword xml:id="k3">cladistics</keyword><keyword xml:id="k4">Indian subcontinent</keyword><keyword xml:id="k5"><i>Mesaceratherium</i></keyword><keyword xml:id="k6">palaeobiogeography</keyword><keyword xml:id="k7">Perissodactyla</keyword><keyword xml:id="k8"><i>Pleuroceros</i></keyword><keyword xml:id="k9">Zinda Pir</keyword></keywordGroup><abstractGroup><abstract type="main" xml:lang="en"><p>Rhinocerotids are particularly abundant and diversified in Neogene deposits of the Indian subcontinent, but their systematics is far from being well defined. Based on the revision of old collections and new findings from the Early Miocene of the Bugti Hills and Zinda Pir, Pakistan, ‘<i>Aceratherium blanfordi</i> Lydekker, 1884’ is a chimera, consisting of two dentally convergent but postcranially distinct rhinocerotid taxa: <i>Pleuroceros blanfordi</i> and <b><i>Mesaceratherium welcommi</i> sp. nov.</b> Postcranial features appear to be much more diagnostic than craniodental morphology in this case. A phylogenetic analysis based on 282 morphological characters scored for 28 taxa (four outgroups and ingroup including both taxa of interest and a ‘branching group’) strengthens this statement and supports <i>Pleuroceros</i> and <i>Mesaceratherium</i> as monophyletic genera within Rhinocerotinae. Both genera are recognized for the first time outside Europe. In the Bugti Hills, <i>P. blanfordi</i> and <i>M. welcommi</i> are part of an exceptionally diversified rhinocerotid fauna, with up to nine species associated in the same locality (Kumbi 4f). This rhinocerotid assemblage confirms the earliest Miocene age (Agenian/Aquitanian) of the upper member of the Chitarwata Formation as a whole. Coeval homotaxic rhinocerotid faunas from Europe (France, Czech Republic) and East Africa (Uganda, Kenya) support broad and sustainable rhinocerotid interchanges amongst South Asia, Europe, and Africa under compatible environmental conditions throughout earliest Miocene times.</p><p>© 2010 The Linnean Society of London, <i>Zoological Journal of the Linnean Society</i>, 2010, <b>160</b>, 139–194.</p></abstract></abstractGroup></contentMeta></header></component></istex:document></istex:metadataXml><mods version="3.6"><titleInfo lang="en"><title>A revision of Aceratherium blanfordi Lydekker, 1884 (Mammalia: Rhinocerotidae) from the Early Miocene of Pakistan: postcranials as a key</title></titleInfo><titleInfo type="abbreviated" lang="en"><title>MIOCENE RHINOCEROTIDS FROM PAKISTAN</title></titleInfo><titleInfo type="alternative" contentType="CDATA" lang="en"><title>A revision of Aceratherium blanfordi Lydekker, 1884 (Mammalia: Rhinocerotidae) from the Early Miocene of Pakistan: postcranials as a key</title></titleInfo><name type="personal"><namePart type="given">PIERRE‐OLIVIER</namePart><namePart type="family">ANTOINE</namePart><affiliation>Université de Toulouse, UPS (SVT‐OMP), LMTG, 14 Avenue Édouard Belin, F‐31400 Toulouse, France</affiliation><affiliation>CNRS, LMTG, 14 Avenue Édouard Belin, F‐31400 Toulouse, France</affiliation><affiliation>IRD, LMTG, 14 Avenue Édouard Belin, F‐31400 Toulouse, France</affiliation><affiliation>E-mail: poa@lmtg.obs‐mip.fr</affiliation><role><roleTerm type="text">author</roleTerm></role></name><name type="personal"><namePart type="given">KEVIN F.</namePart><namePart type="family">DOWNING</namePart><affiliation>DePaul University, Chicago, IL 60604, USA</affiliation><role><roleTerm type="text">author</roleTerm></role></name><name type="personal"><namePart type="given">JEAN‐YVES</namePart><namePart type="family">CROCHET</namePart><affiliation>Département de Paléontologie, Institut des Sciences de l'Évolution (CNRS‐UMR 5554) Université Montpellier 2, c.c. 064, Place Eugène Bataillon, F‐34095 Montpellier Cedex 5, France</affiliation><role><roleTerm type="text">author</roleTerm></role></name><name type="personal"><namePart type="given">FRANCIS</namePart><namePart type="family">DURANTHON</namePart><affiliation>Laboratoire d'Étude et de Conservation du Patrimoine, Muséum d'Histoire naturelle de Toulouse, 39 allées Jules Guesde, F‐31000 Toulouse, France</affiliation><role><roleTerm type="text">author</roleTerm></role></name><name type="personal"><namePart type="given">LAWRENCE J.</namePart><namePart type="family">FLYNN</namePart><affiliation>Department of Anthropology and Peabody Museum, Harvard University, Cambridge, MA 02138, USA</affiliation><role><roleTerm type="text">author</roleTerm></role></name><name type="personal"><namePart type="given">LAURENT</namePart><namePart type="family">MARIVAUX</namePart><affiliation>Département de Paléontologie, Institut des Sciences de l'Évolution (CNRS‐UMR 5554) Université Montpellier 2, c.c. 064, Place Eugène Bataillon, F‐34095 Montpellier Cedex 5, France</affiliation><role><roleTerm type="text">author</roleTerm></role></name><name type="personal"><namePart type="given">GRÉGOIRE</namePart><namePart type="family">MÉTAIS</namePart><affiliation>Laboratoire de Paléontologie, UMR 5143 CNRS, Muséum National d'Histoire Naturelle, 8 rue Buffon, F‐75005 Paris, France</affiliation><role><roleTerm type="text">author</roleTerm></role></name><name type="personal"><namePart type="given">ABDUL RAHIM</namePart><namePart type="family">RAJPAR</namePart><affiliation>9Earth Sciences Division, Pakistan Museum of Natural History, Garden Avenue, Shakarparian, 44000 Islamabad, Pakistan</affiliation><role><roleTerm type="text">author</roleTerm></role></name><name type="personal"><namePart type="given">GHAZALA</namePart><namePart type="family">ROOHI</namePart><affiliation>9Earth Sciences Division, Pakistan Museum of Natural History, Garden Avenue, Shakarparian, 44000 Islamabad, Pakistan</affiliation><role><roleTerm type="text">author</roleTerm></role></name><typeOfResource>text</typeOfResource><genre type="article" displayLabel="article" authority="ISTEX" authorityURI="https://content-type.data.istex.fr" valueURI="https://content-type.data.istex.fr/ark:/67375/XTP-6N5SZHKN-D">article</genre><originInfo><publisher>Blackwell Publishing Ltd</publisher><place><placeTerm type="text">Oxford, UK</placeTerm></place><dateIssued encoding="w3cdtf">2010-09</dateIssued><edition>Received 22 January 2009; 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Based on the revision of old collections and new findings from the Early Miocene of the Bugti Hills and Zinda Pir, Pakistan, ‘Aceratherium blanfordi Lydekker, 1884’ is a chimera, consisting of two dentally convergent but postcranially distinct rhinocerotid taxa: Pleuroceros blanfordi and Mesaceratherium welcommi sp. nov. Postcranial features appear to be much more diagnostic than craniodental morphology in this case. A phylogenetic analysis based on 282 morphological characters scored for 28 taxa (four outgroups and ingroup including both taxa of interest and a ‘branching group’) strengthens this statement and supports Pleuroceros and Mesaceratherium as monophyletic genera within Rhinocerotinae. Both genera are recognized for the first time outside Europe. In the Bugti Hills, P. blanfordi and M. welcommi are part of an exceptionally diversified rhinocerotid fauna, with up to nine species associated in the same locality (Kumbi 4f). This rhinocerotid assemblage confirms the earliest Miocene age (Agenian/Aquitanian) of the upper member of the Chitarwata Formation as a whole. Coeval homotaxic rhinocerotid faunas from Europe (France, Czech Republic) and East Africa (Uganda, Kenya) support broad and sustainable rhinocerotid interchanges amongst South Asia, Europe, and Africa under compatible environmental conditions throughout earliest Miocene times.</abstract><abstract>© 2010 The Linnean Society of London, Zoological Journal of the Linnean Society, 2010, 160, 139–194.</abstract><subject lang="en"><genre>keywords</genre><topic>Bugti Hills</topic><topic>Chitarwata Formation</topic><topic>cladistics</topic><topic>Indian subcontinent</topic><topic>Mesaceratherium</topic><topic>palaeobiogeography</topic><topic>Perissodactyla</topic><topic>Pleuroceros</topic><topic>Zinda Pir</topic></subject><relatedItem type="host"><titleInfo><title>Zoological Journal of the Linnean Society</title></titleInfo><genre type="journal" authority="ISTEX" authorityURI="https://publication-type.data.istex.fr" valueURI="https://publication-type.data.istex.fr/ark:/67375/JMC-0GLKJH51-B">journal</genre><identifier type="ISSN">0024-4082</identifier><identifier type="eISSN">1096-3642</identifier><identifier type="DOI">10.1111/(ISSN)1096-3642</identifier><identifier type="PublisherID">ZOJ</identifier><part><date>2010</date><detail type="volume"><caption>vol.</caption><number>160</number></detail><detail type="issue"><caption>no.</caption><number>1</number></detail><extent unit="pages"><start>139</start><end>194</end><total>56</total></extent></part></relatedItem><identifier type="istex">3C10EB81E8129E9F8E7C93E4DEC42360A31E54B3</identifier><identifier type="ark">ark:/67375/WNG-H9V547SV-F</identifier><identifier type="DOI">10.1111/j.1096-3642.2009.00597.x</identifier><identifier type="ArticleID">ZOJ597</identifier><accessCondition type="use and reproduction" contentType="copyright">© 2010 The Linnean Society of London</accessCondition><recordInfo><recordContentSource authority="ISTEX" authorityURI="https://loaded-corpus.data.istex.fr" valueURI="https://loaded-corpus.data.istex.fr/ark:/67375/XBH-L0C46X92-X">wiley</recordContentSource><recordOrigin>Blackwell Publishing Ltd</recordOrigin></recordInfo></mods><json:item><extension>json</extension><original>false</original><mimetype>application/json</mimetype><uri>https://api.istex.fr/document/3C10EB81E8129E9F8E7C93E4DEC42360A31E54B3/metadata/json</uri></json:item></metadata><serie></serie></istex></record>Pour manipuler ce document sous Unix (Dilib)
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