Addendum to ‘Revision of the Late Jurassic teleosaurid genus Machimosaurus (Crocodylomorpha, Thalattosuchia)’
Identifieur interne : 000004 ( Pmc/Corpus ); précédent : 000003; suivant : 000005Addendum to ‘Revision of the Late Jurassic teleosaurid genus Machimosaurus (Crocodylomorpha, Thalattosuchia)’
Auteurs : Mark T. Young ; Stéphane Hua ; Lorna Steel ; Davide Foffa ; Stephen L. Brusatte ; Silvan Thüring ; Octávio Mateus ; José Ignacio Ruiz-Ome Aca ; Philipe Havlik ; Yves Lepage ; Marco Brandalise De AndradeSource :
- Royal Society Open Science [ 2054-5703 ] ; 2015.
Url:
DOI: 10.1098/rsos.150024
PubMed: NONE
PubMed Central: 4448809
Links to Exploration step
PMC:4448809Le document en format XML
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(Crocodylomorpha, Thalattosuchia)’</title>
<author><name sortKey="Young, Mark T" sort="Young, Mark T" uniqKey="Young M" first="Mark T." last="Young">Mark T. Young</name>
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<author><name sortKey="Hua, Stephane" sort="Hua, Stephane" uniqKey="Hua S" first="Stéphane" last="Hua">Stéphane Hua</name>
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<author><name sortKey="Steel, Lorna" sort="Steel, Lorna" uniqKey="Steel L" first="Lorna" last="Steel">Lorna Steel</name>
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<author><name sortKey="Foffa, Davide" sort="Foffa, Davide" uniqKey="Foffa D" first="Davide" last="Foffa">Davide Foffa</name>
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<author><name sortKey="Thuring, Silvan" sort="Thuring, Silvan" uniqKey="Thuring S" first="Silvan" last="Thüring">Silvan Thüring</name>
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<author><name sortKey="Mateus, Octavio" sort="Mateus, Octavio" uniqKey="Mateus O" first="Octávio" last="Mateus">Octávio Mateus</name>
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<affiliation><nlm:aff id="af9"><addr-line>Museu da Lourinhã</addr-line>
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<author><name sortKey="Ruiz Ome Aca, Jose Ignacio" sort="Ruiz Ome Aca, Jose Ignacio" uniqKey="Ruiz Ome Aca J" first="José Ignacio" last="Ruiz-Ome Aca">José Ignacio Ruiz-Ome Aca</name>
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<author><name sortKey="Havlik, Philipe" sort="Havlik, Philipe" uniqKey="Havlik P" first="Philipe" last="Havlik">Philipe Havlik</name>
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,<addr-line>Sigwartstrasse 10, Tübingen 72072, Germany</addr-line>
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<author><name sortKey="Lepage, Yves" sort="Lepage, Yves" uniqKey="Lepage Y" first="Yves" last="Lepage">Yves Lepage</name>
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<author><name sortKey="De Andrade, Marco Brandalise" sort="De Andrade, Marco Brandalise" uniqKey="De Andrade M" first="Marco Brandalise" last="De Andrade">Marco Brandalise De Andrade</name>
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,<institution>Pontifícia Universidade Católica do Rio Grande do Sul</institution>
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<sourceDesc><biblStruct><analytic><title xml:lang="en" level="a" type="main">Addendum to ‘Revision of the Late Jurassic teleosaurid genus <italic>Machimosaurus</italic>
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<author><name sortKey="Young, Mark T" sort="Young, Mark T" uniqKey="Young M" first="Mark T." last="Young">Mark T. Young</name>
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</nlm:aff>
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<author><name sortKey="Steel, Lorna" sort="Steel, Lorna" uniqKey="Steel L" first="Lorna" last="Steel">Lorna Steel</name>
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</nlm:aff>
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<author><name sortKey="Foffa, Davide" sort="Foffa, Davide" uniqKey="Foffa D" first="Davide" last="Foffa">Davide Foffa</name>
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</nlm:aff>
</affiliation>
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<author><name sortKey="Thuring, Silvan" sort="Thuring, Silvan" uniqKey="Thuring S" first="Silvan" last="Thüring">Silvan Thüring</name>
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<author><name sortKey="Mateus, Octavio" sort="Mateus, Octavio" uniqKey="Mateus O" first="Octávio" last="Mateus">Octávio Mateus</name>
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,<institution>Universidade Nova de Lisboa</institution>
,<addr-line>Quinta da Torre, Caparica 2829-516, Portugal</addr-line>
</nlm:aff>
</affiliation>
<affiliation><nlm:aff id="af9"><addr-line>Museu da Lourinhã</addr-line>
,<institution>Rua João Luis de Moura 9</institution>
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</nlm:aff>
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<author><name sortKey="Havlik, Philipe" sort="Havlik, Philipe" uniqKey="Havlik P" first="Philipe" last="Havlik">Philipe Havlik</name>
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</nlm:aff>
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</nlm:aff>
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<author><name sortKey="De Andrade, Marco Brandalise" sort="De Andrade, Marco Brandalise" uniqKey="De Andrade M" first="Marco Brandalise" last="De Andrade">Marco Brandalise De Andrade</name>
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</author>
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<journal-id journal-id-type="iso-abbrev">R Soc Open Sci</journal-id>
<journal-id journal-id-type="publisher-id">RSOS</journal-id>
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<article-id pub-id-type="doi">10.1098/rsos.150024</article-id>
<article-id pub-id-type="publisher-id">rsos150024</article-id>
<article-categories><subj-group subj-group-type="heading"><subject>Biology (Whole Organism)</subject>
</subj-group>
</article-categories>
<title-group><article-title>Addendum to ‘Revision of the Late Jurassic teleosaurid genus <italic>Machimosaurus</italic>
(Crocodylomorpha, Thalattosuchia)’</article-title>
<alt-title alt-title-type="short">Addendum to Machimosaurus revision</alt-title>
</title-group>
<contrib-group><contrib contrib-type="author"><name><surname>Young</surname>
<given-names>Mark T.</given-names>
</name>
<xref ref-type="aff" rid="af1">1</xref>
<xref ref-type="aff" rid="af2">2</xref>
<xref ref-type="corresp" rid="cor1"></xref>
</contrib>
<contrib contrib-type="author"><name><surname>Hua</surname>
<given-names>Stéphane</given-names>
</name>
<xref ref-type="aff" rid="af3">3</xref>
</contrib>
<contrib contrib-type="author"><name><surname>Steel</surname>
<given-names>Lorna</given-names>
</name>
<xref ref-type="aff" rid="af4">4</xref>
</contrib>
<contrib contrib-type="author"><name><surname>Foffa</surname>
<given-names>Davide</given-names>
</name>
<xref ref-type="aff" rid="af1">1</xref>
<xref ref-type="aff" rid="af5">5</xref>
</contrib>
<contrib contrib-type="author"><name><surname>Brusatte</surname>
<given-names>Stephen L.</given-names>
</name>
<xref ref-type="aff" rid="af1">1</xref>
<xref ref-type="aff" rid="af6">6</xref>
</contrib>
<contrib contrib-type="author"><name><surname>Thüring</surname>
<given-names>Silvan</given-names>
</name>
<xref ref-type="aff" rid="af7">7</xref>
</contrib>
<contrib contrib-type="author"><name><surname>Mateus</surname>
<given-names>Octávio</given-names>
</name>
<xref ref-type="aff" rid="af8">8</xref>
<xref ref-type="aff" rid="af9">9</xref>
<contrib-id contrib-id-type="orcid" authenticated="false">http://orcid.org/0000-0003-1253-3616</contrib-id>
</contrib>
<contrib contrib-type="author"><name><surname>Ruiz-Omeñaca</surname>
<given-names>José Ignacio</given-names>
</name>
<xref ref-type="aff" rid="af10">10</xref>
</contrib>
<contrib contrib-type="author"><name><surname>Havlik</surname>
<given-names>Philipe</given-names>
</name>
<xref ref-type="aff" rid="af11">11</xref>
</contrib>
<contrib contrib-type="author"><name><surname>Lepage</surname>
<given-names>Yves</given-names>
</name>
<xref ref-type="aff" rid="af12">12</xref>
</contrib>
<contrib contrib-type="author"><name><surname>de Andrade</surname>
<given-names>Marco Brandalise</given-names>
</name>
<xref ref-type="aff" rid="af13">13</xref>
</contrib>
</contrib-group>
<aff id="af1"><label>1</label>
<addr-line>School of GeoSciences</addr-line>
,<institution>University of Edinburgh, The King's Buildings</institution>
,<addr-line>Edinburgh EH9 3JFE, UK</addr-line>
</aff>
<aff id="af2"><label>2</label>
<addr-line>School of Ocean and Earth Science, National Oceanography Centre</addr-line>
,<institution>University of Southampton</institution>
,<addr-line>Southampton SO14 3ZH, UK</addr-line>
</aff>
<aff id="af3"><label>3</label>
<addr-line>Le Musée des Dinosaures d'Espéraza</addr-line>
,<addr-line>Espéraza 11260, France</addr-line>
</aff>
<aff id="af4"><label>4</label>
<addr-line>Department of Earth Sciences</addr-line>
,<institution>Natural History Museum</institution>
,<addr-line>London SW7 5BD, UK</addr-line>
</aff>
<aff id="af5"><label>5</label>
<addr-line>School of Earth Sciences</addr-line>
,<institution>University of Bristol, Wills Memorial Building</institution>
,<addr-line>Bristol BS8 1RJ, UK</addr-line>
</aff>
<aff id="af6"><label>6</label>
<addr-line>National Museums Scotland</addr-line>
,<addr-line>Chambers St., Edinburgh EH1 1JF, UK</addr-line>
</aff>
<aff id="af7"><label>7</label>
<addr-line>Naturmuseum Solothurn</addr-line>
,<institution>Klosterplatz 2</institution>
,<addr-line>Solothurn 4500, Switzerland</addr-line>
</aff>
<aff id="af8"><label>8</label>
<addr-line>GeoBioTec, Departamento de Ciências da Terra, Faculdade de Ciências e Tecnologia</addr-line>
,<institution>Universidade Nova de Lisboa</institution>
,<addr-line>Quinta da Torre, Caparica 2829-516, Portugal</addr-line>
</aff>
<aff id="af9"><label>9</label>
<addr-line>Museu da Lourinhã</addr-line>
,<institution>Rua João Luis de Moura 9</institution>
,<addr-line>Lourinhã 2530-158, Portugal</addr-line>
</aff>
<aff id="af10"><label>10</label>
<addr-line>Museo del Jurásico de Asturias (MUJA)</addr-line>
,<addr-line>Colunga 33328, Spain</addr-line>
</aff>
<aff id="af11"><label>11</label>
<addr-line>Paläontologische Sammlung, Senckenberg Center for Human Evolution and Palaeoenvironment</addr-line>
,<institution>Eberhard Karls Universität</institution>
,<addr-line>Sigwartstrasse 10, Tübingen 72072, Germany</addr-line>
</aff>
<aff id="af12"><label>12</label>
<addr-line>Sciences et Géologie Normandes</addr-line>
,<addr-line>Le Havre 76620, France</addr-line>
</aff>
<aff id="af13"><label>13</label>
<addr-line>Faculdade de Biociências</addr-line>
,<institution>Pontifícia Universidade Católica do Rio Grande do Sul</institution>
,<addr-line>Av. Ipiranga 6681, 90619-900 Porto Alegre, Brazil</addr-line>
</aff>
<author-notes><corresp id="cor1">Author for correspondence: Mark T. Young e-mail: <email>myoung5@staffmail.ed.ac.uk</email>
</corresp>
</author-notes>
<pub-date pub-type="collection"><month>2</month>
<year>2015</year>
</pub-date>
<pub-date pub-type="epub"><day>25</day>
<month>2</month>
<year>2015</year>
</pub-date>
<pub-date pub-type="pmc-release"><day>25</day>
<month>2</month>
<year>2015</year>
</pub-date>
<pmc-comment> PMC Release delay is 0 months and
0 days and was based on the . </pmc-comment>
<volume>2</volume>
<issue>2</issue>
<elocation-id>150024</elocation-id>
<history><date date-type="received"><day>14</day>
<month>1</month>
<year>2015</year>
</date>
<date date-type="accepted"><day>29</day>
<month>1</month>
<year>2015</year>
</date>
</history>
<permissions><copyright-statement>© 2015 The Authors.</copyright-statement>
<copyright-year>2015</copyright-year>
<license license-type="open-access" xlink:href="http://creativecommons.org/licenses/by/4.0/"><license-p>Published by the Royal Society under the terms of the Creative Commons Attribution License <ext-link ext-link-type="uri" xlink:href="http://creativecommons.org/licenses/by/4.0/">http://creativecommons.org/licenses/by/4.0/</ext-link>
, which permits unrestricted use, provided the original author and source are credited.</license-p>
</license>
</permissions>
<self-uri content-type="pdf" xlink:href="rsos150024.pdf"></self-uri>
<custom-meta-group><custom-meta><meta-name>cover-date</meta-name>
<meta-value>February, 2015</meta-value>
</custom-meta>
</custom-meta-group>
</article-meta>
</front>
<body><sec sec-type="intro" id="s1"><label>1.</label>
<title>Introduction</title>
<p>This addendum is to ensure that the ICZN (International Commission on Zoological Nomenclature) Code criteria for the availability of new nomenclatural acts are satisfied. In our original publication [<xref rid="RSOS150024C1" ref-type="bibr">1</xref>
], the newly established taxon <italic>Machimosaurus</italic>
<italic>buffetauti</italic>
was not named in accordance with the ICZN Code, specifically it failed to fulfil Article 8.5.3. This addendum, being a ‘corrigendum’ in the sense of the Glossary of the Code, should be consulted alongside the original publication [<xref rid="RSOS150024C1" ref-type="bibr">1</xref>
]. However, it is this addendum that fulfils the ICZN Code criteria for establishing the name <italic>M. buffetauti</italic>
. As such, the date of publication and nominal authority of that nomenclatural act is the date that this addendum has been published.</p>
<p>This addendum fulfils Article 8.5.3 of the ICZN Code, namely to ‘be registered in the <italic>Official Register of Zoological Nomenclature</italic>
(ZooBank) (see Article 78.2.4)’. Thus, we have officially registered the new nomenculatural act <italic>M. buffetauti</italic>
(in addition to registering the genus, type species and the publication establishing them). Note that at present (14/Jan/2015), ZooBank only accepts new names, as other nomenclatural acts (such as amendment of spellings) are not currently supported. As such, <italic>Machimosaurus</italic>
is currently registered under <italic>Madrimosaurus</italic>
(see [<xref rid="RSOS150024C1" ref-type="bibr">1</xref>
] for details). Below are the resultant LSIDs (Life Science Identifiers), and we provide an updated Systematic Palaeontology section from Young <italic>et al</italic>
. [<xref rid="RSOS150024C1" ref-type="bibr">1</xref>
] thus ensuring <italic>M. buffetauti</italic>
sp. nov. is now a valid name.</p>
<p>In addition to being registered with ZooBank, and having the LSIDs recorded within, the electronic edition of this work was published in a journal with an ISSN, and has been archived and is available from the following digital repositories: CLOCKSS and PORTICO.</p>
<p><bold>Institutional abbreviations.</bold>
DFMMh, Dinosaurier-Freilichtmuseum Münchehagen, Lower Saxony, Germany; GPIT, Paläontologische Sammlung der Eberhard Karls Universität Tübingen, Germany; MPV, Musée paléontologique (Paléospace) de Villers-sur-Mer, Normandy, France; NHMUK, Natural History Museum, London, United Kingdom; SMNS, Staatliches Museum für Naturkunde Stuttgart, Germany.</p>
</sec>
<sec id="s2"><label>2.</label>
<title>Zoobank LSIDs</title>
<list list-type="simple"><list-item><p><bold>Original publication (Young <italic>et al</italic>
. [<xref rid="RSOS150024C1" ref-type="bibr">1</xref>
])</bold>
</p>
</list-item>
<list-item><p>urn:lsid:zoobank.org:pub:9125E890-E1E3–4FF8-BB24-D693DFE4F286</p>
</list-item>
<list-item><p><bold>This addendum</bold>
</p>
</list-item>
<list-item><p>urn:lsid:zoobank.org:pub:E4733A09-2781–43BE-8F2C-621D5C29B7C6</p>
</list-item>
<list-item><p><bold>The publication von Meyer, 1837 [<xref rid="RSOS150024C2" ref-type="bibr">2</xref>
]</bold>
</p>
</list-item>
<list-item><p>urn:lsid:zoobank.org:pub:7B719821-F101–4380-A590-12E2D86704EC</p>
</list-item>
<list-item><p><bold>Genus</bold>
<bold><italic>Machimosaurus</italic>
</bold>
<bold>(note that it is registered under the original mis-spelling of</bold>
<bold><italic>Madrimosaurus</italic>
</bold>
<bold>von Meyer, 1837 [<xref rid="RSOS150024C2" ref-type="bibr">2</xref>
])</bold>
</p>
</list-item>
<list-item><p>urn:lsid:zoobank.org:act:DEB4DD43–173A-4B21-B4F0–1C0075B30C3D</p>
</list-item>
<list-item><p><bold>Type species,</bold>
<bold><italic>Machimosaurus hugii</italic>
</bold>
[<xref rid="RSOS150024C2" ref-type="bibr">2</xref>
]</p>
</list-item>
<list-item><p>urn:lsid:zoobank.org:act:D44630DF-5C84–4A73–8D13-A2CF4F4948C5</p>
</list-item>
<list-item><p><bold>New Species</bold>
, <bold><italic>Machimosaurus buffetauti</italic>
</bold>
</p>
</list-item>
<list-item><p>urn:lsid:zoobank.org:act:EAA53B27–2A4C-4395-B776-CDCFD715C413</p>
</list-item>
</list>
</sec>
<sec id="s3"><label>3.</label>
<title>Systematic palaeontology</title>
<p><bold><italic>Machimosaurus</italic>
</bold>
<bold>von Meyer, 1837 [<xref rid="RSOS150024C2" ref-type="bibr">2</xref>
]</bold>
<bold>emend. von Meyer, 1838</bold>
[<xref rid="RSOS150024C3" ref-type="bibr">3</xref>
]</p>
<p><italic>Machimosaurus buffetauti</italic>
sp. nov.
<list list-type="simple"><list-item><p>— v 1873 <italic>Steneosaurus burgensis</italic>
nomen nudum—Jarrin [<xref rid="RSOS150024C4" ref-type="bibr">4</xref>
]</p>
</list-item>
<list-item><p>— v 1876 <italic>Steneosaurus burgensis</italic>
nomen nudum—Jarrin [<xref rid="RSOS150024C5" ref-type="bibr">5</xref>
]</p>
</list-item>
<list-item><p>— v 1905 <italic>Steneosaurus burgensis chanuti</italic>
nomen nudum—Chanel [<xref rid="RSOS150024C6" ref-type="bibr">6</xref>
, figs. 1–3]</p>
</list-item>
<list-item><p>— v 1982 <italic>Machimosaurus hugii</italic>
von Meyer—Buffetaut [<xref rid="RSOS150024C7" ref-type="bibr">7</xref>
, Plate 1, figs. A–D]</p>
</list-item>
<list-item><p>— v 1982 <italic>Machimosaurus hugii</italic>
von Meyer—Buffetaut [<xref rid="RSOS150024C8" ref-type="bibr">8</xref>
, Plate 1]</p>
</list-item>
<list-item><p>— v 2004 <italic>Machimosaurus hugii</italic>
von Meyer—Karl & Tichy [<xref rid="RSOS150024C9" ref-type="bibr">9</xref>
, figs. 1–2]</p>
</list-item>
<list-item><p>— v 2006 <italic>Machimosaurus hugii</italic>
von Meyer—Karl <italic>et al</italic>
. [10, fig. 8]</p>
</list-item>
<list-item><p>— v 2008 <italic>Machimosaurus hugii</italic>
von Meyer—Lepage <italic>et al</italic>
. [<xref rid="RSOS150024C11" ref-type="bibr">11</xref>
, figs. 1–7]</p>
</list-item>
<list-item><p>— v 2008 <italic>Machimosaurus hugii</italic>
von Meyer—Pierce <italic>et al</italic>
. (<italic>partim</italic>
) [<xref rid="RSOS150024C12" ref-type="bibr">12</xref>
]</p>
</list-item>
<list-item><p>— v 2013 <italic>Machimosaurus hugii</italic>
von Meyer—Martin & Vincent [<xref rid="RSOS150024C13" ref-type="bibr">13</xref>
, figs. 1–9]</p>
</list-item>
<list-item><p>— v 2014 <italic>Machimosaurus buffetauti</italic>
sp. nov. et invalid—Young <italic>et al</italic>
. [<xref rid="RSOS150024C1" ref-type="bibr">1</xref>
, figs. 1–9]</p>
</list-item>
<list-item><p>— v 2014 <italic>Machimosaurus buffetauti</italic>
invalid name—Young <italic>et al</italic>
. [<xref rid="RSOS150024C14" ref-type="bibr">14</xref>
, figs. 2–6]</p>
</list-item>
</list>
</p>
<p><bold>Holotype.</bold>
SMNS 91415: complete skull and mandible, with partial post-cranial skeleton ([<xref rid="RSOS150024C1" ref-type="bibr">1</xref>
, figs. 22–27], [<xref rid="RSOS150024C13" ref-type="bibr">13</xref>
, figs. 1–9] and [<xref rid="RSOS150024C14" ref-type="bibr">14</xref>
, fig. 2]). The holotype repository is the Staaliches Museum für Naturkunde Stuttgart.</p>
<p><bold>Holotype locality.</bold>
Am Hörnle Quarry, Neuffen, Baden-Württemberg, Germany [<xref rid="RSOS150024C13" ref-type="bibr">13</xref>
].</p>
<p><bold>Holotype horizon.</bold>
Lacunosamergel Formation, <italic>Ataxioceras hypselocyclum</italic>
Sub-Mediterranean ammonite Zone (Weißer Jura gamma 2), lower Kimmeridgian, Upper Jurassic [<xref rid="RSOS150024C13" ref-type="bibr">13</xref>
].</p>
<p><bold>Etymology.</bold>
’Buffetaut's pugnacious lizard’. Named in honour of Eric Buffetaut (b. 1950), whose research has greatly elucidated thalattosuchian and crocodylomorph evolution.</p>
<p><bold>Referred specimens.</bold>
MPV V.1600.Bo and V.1601.Bo—anterior half of rostrum (premaxilla, maxilla and dentary) in occlusion and a maxilla-nasal fragment (Calcaires Coquilliers Formation; <italic>Pictonia baylei</italic>
Sub-Boreal ammonite Zone, lowermost Kimmeridgian of Cricqueboeuf, Normandy, Northern France [<xref rid="RSOS150024C1" ref-type="bibr">1</xref>
, figs. 12–14]) [<xref rid="RSOS150024C7" ref-type="bibr">7</xref>
,<xref rid="RSOS150024C11" ref-type="bibr">11</xref>
].</p>
<p>MPV KIM027—Cricqueboeuf, Normandy, France (Calcaires à <italic>Harpagodes</italic>
Member of the Marnes de Bléville Formation, <italic>Rasenia cymodoce</italic>
Sub-Boreal ammonite Zone, lower Kimmeridgian [<xref rid="RSOS150024C14" ref-type="bibr">14</xref>
, fig. 5]).</p>
<p>DFMMh FV 330, DFMMh FV 541: isolated tooth crowns (Langenberg Formation; Langenberg near Oker, Lower Saxony, Germany; Kimmeridgian [<xref rid="RSOS150024C1" ref-type="bibr">1</xref>
, fig. 21C-H] and [<xref rid="RSOS150024C14" ref-type="bibr">14</xref>
, figs. 3–4]) [<xref rid="RSOS150024C9" ref-type="bibr">9</xref>
,<xref rid="RSOS150024C10" ref-type="bibr">10</xref>
].</p>
<p>Musée de Brou (Bourg-en-Bresse, France), specimen number unknown—a complete skull and mandible in articulation (Calcaires à ptérocères Formation, lower Kimmeridgian; Montmerle, Bourg-en-Bresse, département de l'Ain, France [<xref rid="RSOS150024C8" ref-type="bibr">8</xref>
]).</p>
<p><bold>Geological range.</bold>
Lower Kimmeridgian.</p>
<p><bold>Geographical range.</bold>
Europe (France and Germany). An isolated tooth from Smallmouth Sands, England (NHMUK PV R1774 [<xref rid="RSOS150024C1" ref-type="bibr">1</xref>
, fig. 21A-B] and [<xref rid="RSOS150024C14" ref-type="bibr">14</xref>
, fig. 6]) is possibly referable to this taxon, as are isolated teeth from Czarnogłowy, Poland (GPIT/RE/328, GPIT/RE/9280 and GPIT/RE/9281 [<xref rid="RSOS150024C1" ref-type="bibr">1</xref>
, figs. 31–32]).</p>
<p><bold>Species diagnosis.</bold>
Teleosaurid crocodylomorph within the genus <italic>Machimosaurus</italic>
with the following unique combination of characters (autapomorphic characters are indicated by an asterisk *): 21–22 alveoli per maxilla (approx. 16–17 of which are anterior to the palatines); 24/25 alveoli per dentary (19–20 of which are adjacent to the mandibular symphysis); low post-symphyseal dentary tooth count (2 pairs)*; inter-alveolar spaces between the maxillary and dentary alveoli are variable in size (thalattosuchian symplesiomorphy); orbits are sub-circular in shape (transverse and anteroposterior axes are sub-equal; the <italic>Steneosaurus brevior</italic>
holotype also has circular orbits)*; the quadrates have a single large circular depression on the dorsal surface near the hemicondyles*; basioccipital apophysis has a ‘U-shaped’ cross-section (teleosaurid symplesiomorphy); the inter-basioccipital tubera notch is a wide and gentle inverse semi-circle when seen in occipital view (teleosaurid symplesiomorphy); basioccipital tuberosities (basal tubera) are reduced in size when seen in occipital view (apomorphy shared with <italic>Machimosaurus mosae</italic>
); axis neural arch dorsal margin is strongly concave when seen in lateral view*; axis postzygapophyses terminate significantly posterior to posterior-surface of the centrum view (somewhat similar to that seen in <italic>Steneosaurus durobrivensis</italic>
); coracoid glenoid process (process near the glenoid fossa that projects posterodorsally) is elongate, extending considerably from the head of the coracoid, and is a sub-isosceles triangle in shape when seen in lateral view*; coracoid postglenoid process anterior margin is very slightly concave, and terminates approximately in the same frontal plane as the glenoid*; coracoid postglenoid process posterior margin is strongly concave, and terminates approximately in the same frontal plane as the posterior-end of the glenoid process*; dorsal osteoderm ornamentation is composed of small-to-large irregularly shaped pits arranged in a random manner, that are well separated from one another (somewhat similar to <italic>Steneosaurus leedsi</italic>
).</p>
<p><bold><italic>Steneosaurus burgensis</italic>
</bold>
. The name <italic>S. burgensis</italic>
, and <italic>S. burgensis chanuti</italic>
, have been applied to the <italic>Machimosaurus</italic>
skull from Ain, France [<xref rid="RSOS150024C4" ref-type="bibr">4</xref>
–<xref rid="RSOS150024C6" ref-type="bibr">6</xref>
,<xref rid="RSOS150024C8" ref-type="bibr">8</xref>
]. These specific and sub-specific names are however <italic>nomina nuda</italic>
. Both Jarrin [<xref rid="RSOS150024C5" ref-type="bibr">5</xref>
] and Chanel [<xref rid="RSOS150024C6" ref-type="bibr">6</xref>
] stated that the Ain skull was sent to Caen for preparation and study by Eugène Eudes-Deslongchamps, who proposed the name <italic>Steneosaurus burgensis</italic>
for the specimen, in consultation with the Société d'émulation de l'Ain. Neither Jarrin [<xref rid="RSOS150024C4" ref-type="bibr">4</xref>
,<xref rid="RSOS150024C5" ref-type="bibr">5</xref>
] nor Chanel [<xref rid="RSOS150024C6" ref-type="bibr">6</xref>
] established the name under Article 12 of the ICZN Code as: they did not describe the specimen, nor did they provide a definition of the species; they simply reported that the name was proposed by Eudes-Deslongchamps. Unfortunately, Eudes-Deslongchamps never published his description of the Ain skull [<xref rid="RSOS150024C6" ref-type="bibr">6</xref>
,<xref rid="RSOS150024C8" ref-type="bibr">8</xref>
]. The specimen was not formally described until 1982, and then it was referred to <italic>Machimosaurus hugii</italic>
[<xref rid="RSOS150024C8" ref-type="bibr">8</xref>
]. The sub-specific epithet <italic>chanuti</italic>
was apparently established by those who did not fully understand zoological nomenclature [<xref rid="RSOS150024C8" ref-type="bibr">8</xref>
], as it was added to the Ain skull's specimen plaque solely to honour the discoverer [<xref rid="RSOS150024C6" ref-type="bibr">6</xref>
,<xref rid="RSOS150024C8" ref-type="bibr">8</xref>
].</p>
<p>Our decision to establish a new taxon based on SMNS 91415, and not formally establish <italic>S. burgensis</italic>
for the Ain skull was for several reasons: (i) the Ain skull is partially reconstructed, and it is unclear how much is plaster and how much is real bone [<xref rid="RSOS150024C8" ref-type="bibr">8</xref>
]; (ii) the cranium and lower jaw of the Ain skull are in articulation, meaning that the palatal and dorsal mandibular morphologies cannot be seen [<xref rid="RSOS150024C6" ref-type="bibr">6</xref>
,<xref rid="RSOS150024C8" ref-type="bibr">8</xref>
]; (iii) the German skull SMNS 91415 has the cranium and lower jaw disarticulated, allowing these morphologies to be observed [<xref rid="RSOS150024C13" ref-type="bibr">13</xref>
] and (iv) SMNS 91415 has associated post-cranial material, greatly aiding in comparisons with other <italic>Machimosaurus</italic>
taxa, in particular, <italic>M. mosae</italic>
.</p>
</sec>
</body>
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