Review of the Psychodinae from Mallorca, Spain, with description of Pericomaunipennata, sp. n. (Diptera, Psychodidae)
Identifieur interne : 000105 ( Pmc/Corpus ); précédent : 000104; suivant : 000106Review of the Psychodinae from Mallorca, Spain, with description of Pericomaunipennata, sp. n. (Diptera, Psychodidae)
Auteurs : Gunnar Mikalsen Kvifte ; Morten Stokkan ; Rüdiger WagnerSource :
- ZooKeys [ 1313-2989 ] ; 2016.
Abstract
We review the
Url:
DOI: 10.3897/zookeys.577.7679
PubMed: 27110192
PubMed Central: 4829887
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<record><TEI><teiHeader><fileDesc><titleStmt><title xml:lang="en">Review of the <named-content content-type="taxon-name"><named-content content-type="subfamily">Psychodinae</named-content></named-content> from Mallorca, Spain, with description of <italic><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">unipennata</named-content></named-content></italic>, sp. n. (<named-content content-type="taxon-name"><named-content content-type="order">Diptera</named-content></named-content>, <named-content content-type="taxon-name"><named-content content-type="family">Psychodidae</named-content></named-content>)</title><author><name sortKey="Kvifte, Gunnar Mikalsen" sort="Kvifte, Gunnar Mikalsen" uniqKey="Kvifte G" first="Gunnar Mikalsen" last="Kvifte">Gunnar Mikalsen Kvifte</name><affiliation><nlm:aff id="A1"><addr-line>Dept. of Zoology-Limnology, University of Kassel, Heinrich-Plett-Strasse 40, 34132 Kassel-Oberzwehren, Germany</addr-line></nlm:aff></affiliation><affiliation><nlm:aff id="A2"><addr-line>Dept. of Natural History, University Museum of Bergen, P.O. Box 7800, University of Bergen, 5040 Bergen, Norway</addr-line></nlm:aff></affiliation></author><author><name sortKey="Stokkan, Morten" sort="Stokkan, Morten" uniqKey="Stokkan M" first="Morten" last="Stokkan">Morten Stokkan</name><affiliation><nlm:aff id="A2"><addr-line>Dept. of Natural History, University Museum of Bergen, P.O. Box 7800, University of Bergen, 5040 Bergen, Norway</addr-line></nlm:aff></affiliation><affiliation><nlm:aff id="A3"><addr-line>Department of Biodiversity and Conservation, Instituto Mediterraneo de Estudios Avanzados (IMEDEA, CSIC-UIB), Esporles, Spain</addr-line></nlm:aff></affiliation></author><author><name sortKey="Wagner, Rudiger" sort="Wagner, Rudiger" uniqKey="Wagner R" first="Rüdiger" last="Wagner">Rüdiger Wagner</name><affiliation><nlm:aff id="A1"><addr-line>Dept. of Zoology-Limnology, University of Kassel, Heinrich-Plett-Strasse 40, 34132 Kassel-Oberzwehren, Germany</addr-line></nlm:aff></affiliation></author></titleStmt><publicationStmt><idno type="wicri:source">PMC</idno><idno type="pmid">27110192</idno><idno type="pmc">4829887</idno><idno type="url">http://www.ncbi.nlm.nih.gov/pmc/articles/PMC4829887</idno><idno type="RBID">PMC:4829887</idno><idno type="doi">10.3897/zookeys.577.7679</idno><date when="2016">2016</date><idno type="wicri:Area/Pmc/Corpus">000105</idno></publicationStmt><sourceDesc><biblStruct><analytic><title xml:lang="en" level="a" type="main">Review of the <named-content content-type="taxon-name"><named-content content-type="subfamily">Psychodinae</named-content></named-content> from Mallorca, Spain, with description of <italic><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">unipennata</named-content></named-content></italic>, sp. n. (<named-content content-type="taxon-name"><named-content content-type="order">Diptera</named-content></named-content>, <named-content content-type="taxon-name"><named-content content-type="family">Psychodidae</named-content></named-content>)</title><author><name sortKey="Kvifte, Gunnar Mikalsen" sort="Kvifte, Gunnar Mikalsen" uniqKey="Kvifte G" first="Gunnar Mikalsen" last="Kvifte">Gunnar Mikalsen Kvifte</name><affiliation><nlm:aff id="A1"><addr-line>Dept. of Zoology-Limnology, University of Kassel, Heinrich-Plett-Strasse 40, 34132 Kassel-Oberzwehren, Germany</addr-line></nlm:aff></affiliation><affiliation><nlm:aff id="A2"><addr-line>Dept. of Natural History, University Museum of Bergen, P.O. Box 7800, University of Bergen, 5040 Bergen, Norway</addr-line></nlm:aff></affiliation></author><author><name sortKey="Stokkan, Morten" sort="Stokkan, Morten" uniqKey="Stokkan M" first="Morten" last="Stokkan">Morten Stokkan</name><affiliation><nlm:aff id="A2"><addr-line>Dept. of Natural History, University Museum of Bergen, P.O. Box 7800, University of Bergen, 5040 Bergen, Norway</addr-line></nlm:aff></affiliation><affiliation><nlm:aff id="A3"><addr-line>Department of Biodiversity and Conservation, Instituto Mediterraneo de Estudios Avanzados (IMEDEA, CSIC-UIB), Esporles, Spain</addr-line></nlm:aff></affiliation></author><author><name sortKey="Wagner, Rudiger" sort="Wagner, Rudiger" uniqKey="Wagner R" first="Rüdiger" last="Wagner">Rüdiger Wagner</name><affiliation><nlm:aff id="A1"><addr-line>Dept. of Zoology-Limnology, University of Kassel, Heinrich-Plett-Strasse 40, 34132 Kassel-Oberzwehren, Germany</addr-line></nlm:aff></affiliation></author></analytic><series><title level="j">ZooKeys</title><idno type="ISSN">1313-2989</idno><idno type="eISSN">1313-2970</idno><imprint><date when="2016">2016</date></imprint></series></biblStruct></sourceDesc></fileDesc><profileDesc><textClass></textClass></profileDesc></teiHeader><front><div type="abstract" xml:lang="en"><label>Abstract</label><p>We review the <named-content content-type="taxon-name"><named-content content-type="subfamily">Psychodinae</named-content></named-content> of Mallorca, recognising fifteen species based on recent collections and available literature. Previously unpublished data is presented for eleven species, of which <italic><named-content content-type="taxon-name"><named-content content-type="genus">Neoarisemus</named-content><named-content content-type="species">ibericus</named-content></named-content></italic> Wagner, 1978, <italic><named-content content-type="taxon-name"><named-content content-type="genus">Mormia</named-content><named-content content-type="species">tenebricosa</named-content></named-content></italic> (Vaillant, 1954), <italic><named-content content-type="taxon-name"><named-content content-type="genus">Clogmia</named-content><named-content content-type="species">albipunctata</named-content></named-content></italic> (Williston, 1893), <italic><named-content content-type="taxon-name"><named-content content-type="genus">Lepiseodina</named-content><named-content content-type="species">rothschildi</named-content></named-content></italic> (Eaton, 1913), <italic><named-content content-type="taxon-name"><named-content content-type="genus">Paramormia</named-content><named-content content-type="species">ustulata</named-content></named-content></italic> (Walker, 1856), <italic><named-content content-type="taxon-name"><named-content content-type="genus">Philosepedon</named-content><named-content content-type="species">pyrenaicus</named-content></named-content></italic> Vaillant, 1974 and <named-content content-type="taxon-name"><named-content content-type="genus">Psychoda</named-content> (<named-content content-type="subgenus">Psycha</named-content>) <named-content content-type="species">grisescens</named-content></named-content> Tonnoir, 1922 are first records for Mallorca. An old record of <italic><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">trifasciata</named-content></named-content></italic> (Meigen, 1804) is considered doubtful. <italic><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">unipennata</named-content></named-content></italic><bold>sp. n</bold> is described and illustrated based on a male collected at Deía. Distributional data are reviewed for all newly recorded species. Based on the <named-content content-type="taxon-name"><named-content content-type="subfamily">Psychodinae</named-content></named-content> fauna, the zoogeographical affinities of Mallorca are briefly discussed.</p></div></front><back><div1 type="bibliography"><listBibl><biblStruct><analytic><author><name sortKey="Alcover, Mm" uniqKey="Alcover M">MM Alcover</name></author><author><name sortKey="Ballart, C" uniqKey="Ballart C">C Ballart</name></author><author><name sortKey="Martin Sanchez, J" uniqKey="Martin Sanchez J">J Martín-Sánchez</name></author><author><name sortKey="Serra, T" uniqKey="Serra T">T Serra</name></author><author><name sortKey="Castillejo, S" uniqKey="Castillejo S">S Castillejo</name></author><author><name sortKey="Portus, M" uniqKey="Portus M">M Portús</name></author><author><name sortKey="Gallego, M" uniqKey="Gallego M">M Gállego</name></author></analytic></biblStruct><biblStruct><analytic><author><name sortKey="Andersen, T" uniqKey="Andersen T">T 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<front><journal-meta><journal-id journal-id-type="nlm-ta">Zookeys</journal-id><journal-id journal-id-type="iso-abbrev">Zookeys</journal-id><journal-id journal-id-type="publisher-id">ZooKeys</journal-id><journal-title-group><journal-title>ZooKeys</journal-title></journal-title-group><issn pub-type="ppub">1313-2989</issn><issn pub-type="epub">1313-2970</issn><publisher><publisher-name>Pensoft Publishers</publisher-name></publisher></journal-meta><article-meta><article-id pub-id-type="pmid">27110192</article-id><article-id pub-id-type="pmc">4829887</article-id><article-id pub-id-type="doi">10.3897/zookeys.577.7679</article-id><article-categories><subj-group subj-group-type="heading"><subject>Research Article</subject></subj-group></article-categories><title-group><article-title>Review of the <named-content content-type="taxon-name"><named-content content-type="subfamily">Psychodinae</named-content></named-content> from Mallorca, Spain, with description of <italic><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">unipennata</named-content></named-content></italic>, sp. n. (<named-content content-type="taxon-name"><named-content content-type="order">Diptera</named-content></named-content>, <named-content content-type="taxon-name"><named-content content-type="family">Psychodidae</named-content></named-content>)</article-title></title-group><contrib-group><contrib contrib-type="author"><name><surname>Kvifte</surname><given-names>Gunnar Mikalsen</given-names></name><xref ref-type="aff" rid="A1">1</xref><xref ref-type="aff" rid="A2">2</xref></contrib><contrib contrib-type="author"><name><surname>Stokkan</surname><given-names>Morten</given-names></name><xref ref-type="aff" rid="A2">2</xref><xref ref-type="aff" rid="A3">3</xref></contrib><contrib contrib-type="author"><name><surname>Wagner</surname><given-names>Rüdiger</given-names></name><xref ref-type="aff" rid="A1">1</xref></contrib></contrib-group><aff id="A1"><label>1</label><addr-line>Dept. of Zoology-Limnology, University of Kassel, Heinrich-Plett-Strasse 40, 34132 Kassel-Oberzwehren, Germany</addr-line></aff><aff id="A2"><label>2</label><addr-line>Dept. of Natural History, University Museum of Bergen, P.O. Box 7800, University of Bergen, 5040 Bergen, Norway</addr-line></aff><aff id="A3"><label>3</label><addr-line>Department of Biodiversity and Conservation, Instituto Mediterraneo de Estudios Avanzados (IMEDEA, CSIC-UIB), Esporles, Spain</addr-line></aff><author-notes><corresp>Corresponding author: Gunnar Mikalsen Kvifte (<email xlink:type="simple">Gunnar.Kvifte@uib.no</email>)</corresp><fn fn-type="edited-by"><p>Academic editor: J. Salmela</p></fn></author-notes><pub-date pub-type="collection"><year>2016</year></pub-date><pub-date pub-type="epub"><day>5</day><month>4</month><year>2016</year></pub-date><issue>577</issue><fpage>149</fpage><lpage>160</lpage><history><date date-type="received"><day>2</day><month>1</month><year>2016</year></date><date date-type="accepted"><day>6</day><month>3</month><year>2016</year></date></history><permissions><copyright-statement>Gunnar Mikalsen Kvifte, Morten Stokkan, Rüdiger Wagner</copyright-statement><license license-type="creative-commons-attribution" xlink:href="http://creativecommons.org/licenses/by/4.0"><license-p>This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.</license-p></license></permissions><self-uri content-type="zoobank" xlink:type="simple" xlink:href="http://zoobank.org/99814AF8-365D-43BA-AEE7-521ECCF64238">http://zoobank.org/99814AF8-365D-43BA-AEE7-521ECCF64238</self-uri><abstract><label>Abstract</label><p>We review the <named-content content-type="taxon-name"><named-content content-type="subfamily">Psychodinae</named-content></named-content> of Mallorca, recognising fifteen species based on recent collections and available literature. Previously unpublished data is presented for eleven species, of which <italic><named-content content-type="taxon-name"><named-content content-type="genus">Neoarisemus</named-content><named-content content-type="species">ibericus</named-content></named-content></italic> Wagner, 1978, <italic><named-content content-type="taxon-name"><named-content content-type="genus">Mormia</named-content><named-content content-type="species">tenebricosa</named-content></named-content></italic> (Vaillant, 1954), <italic><named-content content-type="taxon-name"><named-content content-type="genus">Clogmia</named-content><named-content content-type="species">albipunctata</named-content></named-content></italic> (Williston, 1893), <italic><named-content content-type="taxon-name"><named-content content-type="genus">Lepiseodina</named-content><named-content content-type="species">rothschildi</named-content></named-content></italic> (Eaton, 1913), <italic><named-content content-type="taxon-name"><named-content content-type="genus">Paramormia</named-content><named-content content-type="species">ustulata</named-content></named-content></italic> (Walker, 1856), <italic><named-content content-type="taxon-name"><named-content content-type="genus">Philosepedon</named-content><named-content content-type="species">pyrenaicus</named-content></named-content></italic> Vaillant, 1974 and <named-content content-type="taxon-name"><named-content content-type="genus">Psychoda</named-content> (<named-content content-type="subgenus">Psycha</named-content>) <named-content content-type="species">grisescens</named-content></named-content> Tonnoir, 1922 are first records for Mallorca. An old record of <italic><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">trifasciata</named-content></named-content></italic> (Meigen, 1804) is considered doubtful. <italic><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">unipennata</named-content></named-content></italic><bold>sp. n</bold> is described and illustrated based on a male collected at Deía. Distributional data are reviewed for all newly recorded species. Based on the <named-content content-type="taxon-name"><named-content content-type="subfamily">Psychodinae</named-content></named-content> fauna, the zoogeographical affinities of Mallorca are briefly discussed.</p></abstract><kwd-group><label>Keywords</label><kwd>Moth flies</kwd><kwd>Balearic Isles</kwd><kwd>new species</kwd><kwd>faunistics</kwd><kwd>distribution</kwd><kwd>check list</kwd><pmc-comment>PageBreak</pmc-comment>
</kwd-group></article-meta><notes><sec sec-type="Citation"><title>Citation</title><p>Kvifte GM, Stokkan M, Wagner R (2016) Review of the Psychodinae from Mallorca, Spain, with description of <italic>Pericoma unipennata</italic>, sp. n. (Diptera, Psychodidae). ZooKeys 577: 149–160. doi: <ext-link ext-link-type="doi" xlink:href="10.3897/zookeys.577.7679">10.3897/zookeys.577.7679</ext-link></p></sec></notes></front><body><sec sec-type="Introduction"><title>Introduction</title><p>The <named-content content-type="taxon-name"><named-content content-type="family">Psychodidae</named-content></named-content> (moth flies and sand flies) are a species-rich and widespread group of small insects mainly associated with humid habitats. The most thoroughly studied faunas of the group are found in Europe, from where more than 500 species have been described and new discoveries continue to be made (<xref rid="B29" ref-type="bibr">Wagner 2004</xref>, <xref rid="B30" ref-type="bibr">Wagner and Kvifte 2015</xref>). The most poorly studied moth fly faunas of Europe are those of the Iberian peninsula (<xref rid="B28" ref-type="bibr">Wagner 2001</xref>) and the Balearic islands of Spain.</p><p>The most thoroughly studied moth flies from the Balearic islands belong to the subfamily <named-content content-type="taxon-name"><named-content content-type="subfamily">Phlebotominae</named-content></named-content>, the sand flies. On Mallorca, the largest of the Balearic islands, phlebotomine sand flies have received much attention due to their significance in veterinary medicine as vectors of canine leishmaniasis. This disease is widespread and long established, to the point that some local dog breeds have evolved resistance to the parasite (<xref rid="B20" ref-type="bibr">Solano-Gallego et al. 2000</xref>). Four species of <italic><named-content content-type="taxon-name"><named-content content-type="genus">Phlebotomus</named-content></named-content></italic> and one of <italic><named-content content-type="taxon-name"><named-content content-type="genus">Sergentomyia</named-content></named-content></italic> have been recorded from the island, of which <italic><named-content content-type="taxon-name"><named-content content-type="genus">Phlebotomus</named-content><named-content content-type="species">ariasi</named-content></named-content></italic> Tonnoir, 1921 has been suggested as a doubtful record (<xref rid="B1" ref-type="bibr">Alcover et al. 2014</xref>).</p><p>Other subfamilies of <named-content content-type="taxon-name"><named-content content-type="family">Psychodidae</named-content></named-content> have been studied far less. <xref rid="B27" ref-type="bibr">Wagner (1990)</xref> listed a single species, namely <italic><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">barbarica</named-content></named-content></italic> Vaillant, 1955 based on <xref rid="B23" ref-type="bibr">Vaillant (1978)</xref>. <xref rid="B31" ref-type="bibr">Wagner et al. (2002)</xref> further included a record of <italic><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">trifasciata</named-content></named-content></italic> (Meigen, 1818), most likely based on a typographical error in <xref rid="B23" ref-type="bibr">Vaillant (1978)</xref>. The <italic><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">trifasciata</named-content></named-content></italic> record was not mentioned by <xref rid="B29" ref-type="bibr">Wagner (2004)</xref>, who instead listed <italic><named-content content-type="taxon-name"><named-content content-type="genus">Psychoda</named-content><named-content content-type="species">minuta</named-content></named-content></italic> Banks, 1894, <italic><named-content content-type="taxon-name"><named-content content-type="genus">Psychoda</named-content><named-content content-type="species">phalaenoides</named-content></named-content></italic> (Linnaeus, 1758), <named-content content-type="taxon-name"><named-content content-type="genus">Psychoda</named-content> (<named-content content-type="subgenus">Tineria</named-content>) <named-content content-type="species">alternata</named-content></named-content> Say, 1824 and <named-content content-type="taxon-name"><named-content content-type="genus">Psychoda</named-content> (<named-content content-type="subgenus">Tinearia</named-content>) <named-content content-type="species">lativentris</named-content></named-content> Berdén, 1952 based on unpublished material (the latter two placed in <italic><named-content content-type="taxon-name"><named-content content-type="genus">Tinearia</named-content></named-content></italic> Schellenberg, 1803, treated by <xref rid="B29" ref-type="bibr">Wagner 2004</xref> as a genus).</p><p>In the present study, we review existing records of Mallorcan <named-content content-type="taxon-name"><named-content content-type="subfamily">Psychodinae</named-content></named-content> and present new material for eleven species; seven of which are previously unknown from Mallorca. In addition, we describe <italic><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">unipennata</named-content></named-content></italic> sp. n as new to science.</p></sec><sec sec-type="materials|methods"><title>Material and methods</title><p>Specimens were collected mainly by sweep netting and with aspirators from vegetation near the presumed larval habitats and preserved in 70–100% alcohol. Male specimens were sorted, dissected and mounted on slides in euparal (material in coll. <abbrev content-type="institution" xlink:title="Alexander-König Zoologischer Forschungsmuseum, Bonn, Germany">ZFMK</abbrev> and <abbrev content-type="institution" xlink:title="Entomology Collections, Dept. of Natural History, University Museum of Bergen, Bergen, Norway">ZMUB</abbrev>) or Canada balsam (material in coll. <abbrev content-type="institution" xlink:title="Private collection of Rüdiger Wagner, Kassel">RW</abbrev>). Morphological terminology is according to <xref rid="B19" ref-type="bibr">Quate and Brown (2004)</xref> and <xref rid="B14" ref-type="bibr">Kvifte (2014</xref>, <xref rid="B15" ref-type="bibr">2015</xref>). The „median moveable appendage“ in <xref rid="B17" ref-type="bibr">Kvifte et al. (2013)</xref> is here recognised as a <italic>parameral sclerite</italic>. Measurements are given in µm with an accuracy of 3 µm; except wing length which is given in mm to an accuracy of 100 µm.</p><p>Both literature records and new material that we have examined are included in our present checklist. Tribe-level classification is given according to <xref rid="B4" ref-type="bibr">Duckhouse (1987)</xref>, while genus-level taxonomy is according to <xref rid="B29" ref-type="bibr">Wagner (2004)</xref> except where noted otherwise in the text. Species recorded as new to Mallorca are marked with an asterisk (*).</p><pmc-comment>PageBreak</pmc-comment>
<p>The material is deposited in the following collections:</p><p><abbrev content-type="institution">RW
<def><p> Private collection of Rüdiger Wagner, Kassel </p></def></abbrev></p><p><abbrev content-type="institution">ZFMK
<def><p> Alexander-König Zoologischer Forschungsmuseum, Bonn, Germany </p></def></abbrev></p><p><abbrev content-type="institution">ZMUB
<def><p> Entomology Collections, Dept. of Natural History, University Museum of Bergen, Bergen, Norway </p></def></abbrev></p></sec><sec><title>Species list</title><sec sec-type="Maruinini"><title><named-content content-type="taxon-name"><named-content content-type="tribe">Maruinini</named-content></named-content></title><sec sec-type="taxon-treatment"><sec-meta><kwd-group><label>Taxon classification</label><kwd><named-content content-type="kingdom">Animalia</named-content></kwd><kwd><named-content content-type="order">Diptera</named-content></kwd><kwd><named-content content-type="family">Psychodidae</named-content></kwd></kwd-group></sec-meta><label>*</label><title><named-content content-type="taxon-name"><named-content content-type="genus">Neoarisemus</named-content><named-content content-type="species">ibericus</named-content></named-content></title><p><named-content content-type="taxon-authority">Wagner, 1978</named-content></p><sec sec-type="treatment-First record from Mallorca"><title>First record from Mallorca.</title><p>Puigpunyant, <named-content content-type="dwc:verbatimCoordinates">39.6167°N, 2.5167°E</named-content>, <named-content content-type="date">6.x.1981</named-content>, H. Malicky leg. <named-content content-type="specimen-count">1♂</named-content> (<abbrev content-type="institution" xlink:title="Private collection of Rüdiger Wagner, Kassel">RW</abbrev>).</p></sec><sec sec-type="treatment-Remarks"><title>Remarks.</title><p>Previously only recorded from the type locality in northern Spain.</p></sec></sec></sec><sec sec-type="Mormiini"><title><named-content content-type="taxon-name"><named-content content-type="tribe">Mormiini</named-content></named-content></title><sec sec-type="taxon-treatment"><sec-meta><kwd-group><label>Taxon classification</label><kwd><named-content content-type="kingdom">Animalia</named-content></kwd><kwd><named-content content-type="order">Diptera</named-content></kwd><kwd><named-content content-type="family">Psychodidae</named-content></kwd></kwd-group></sec-meta><label>*</label><title><named-content content-type="taxon-name"><named-content content-type="genus">Mormia</named-content><named-content content-type="species">tenebricosa</named-content></named-content></title><p><named-content content-type="taxon-authority">(Vaillant, 1954)</named-content></p><sec sec-type="treatment-First records from Mallorca"><title>First records from Mallorca.</title><p>Calobra, <named-content content-type="dwc:verbatimCoordinates">39.85°N, 2.8°E</named-content>, <named-content content-type="date">9.v.1978</named-content>, 90 m a.s.l., H. Malicky leg. <named-content content-type="specimen-count">1♂</named-content> (<abbrev content-type="institution" xlink:title="Private collection of Rüdiger Wagner, Kassel">RW</abbrev>);</p><p>Deiá, town fountain, <named-content content-type="dwc:verbatimCoordinates">39.748072°N, 2.643385°E</named-content>, <named-content content-type="date">8.ii.2015</named-content>, G.Kvifte, M. Stokkan & C. Garcia leg. <named-content content-type="specimen-count">1♂</named-content> (<abbrev content-type="institution" xlink:title="Alexander-König Zoologischer Forschungsmuseum, Bonn, Germany">ZFMK</abbrev>);</p><p>Puigpunyent, <named-content content-type="dwc:verbatimCoordinates">39.62°N, 2.85°E</named-content>, <named-content content-type="date">12.v.1978</named-content>, 200m a.s.l., H. Malicky leg., <named-content content-type="specimen-count">1♂</named-content>; same but <named-content content-type="date">1.X.1979</named-content>, <named-content content-type="specimen-count">6 ♂♂</named-content>; same but <named-content content-type="date">6.x.1979</named-content>, <named-content content-type="specimen-count">6♂♂</named-content> (all <abbrev content-type="institution" xlink:title="Private collection of Rüdiger Wagner, Kassel">RW</abbrev>);</p><p>South slope of Piug Major, north of Soller, <named-content content-type="dwc:verbatimCoordinates">39.783°N, 2.767°E</named-content>, <named-content content-type="date">7.–9.v.1978</named-content>, 700 m a.s.l., H. Malicky leg. <named-content content-type="specimen-count">2♂♂</named-content> (<abbrev content-type="institution" xlink:title="Private collection of Rüdiger Wagner, Kassel">RW</abbrev>).</p></sec><sec sec-type="treatment-Remarks"><title>Remarks.</title><p>Previously known from France, mainland Spain, Italy, Morocco and Algeria (<xref rid="B22" ref-type="bibr">Vaillant 1974</xref>). The generic and subgeneric classification of <named-content content-type="taxon-name"><named-content content-type="subtribe">Mormiina</named-content></named-content> is unstable and requires revision; therefore the placement in <italic><named-content content-type="taxon-name"><named-content content-type="genus">Mormia</named-content></named-content></italic> may change.</p></sec></sec></sec><sec sec-type="Paramormiini"><title><named-content content-type="taxon-name"><named-content content-type="tribe">Paramormiini</named-content></named-content></title><sec sec-type="taxon-treatment"><sec-meta><kwd-group><label>Taxon classification</label><kwd><named-content content-type="kingdom">Animalia</named-content></kwd><kwd><named-content content-type="order">Diptera</named-content></kwd><kwd><named-content content-type="family">Psychodidae</named-content></kwd></kwd-group></sec-meta><label>*</label><title><named-content content-type="taxon-name"><named-content content-type="genus">Clogmia</named-content><named-content content-type="species">albipunctata</named-content></named-content></title><p><named-content content-type="taxon-authority">(Williston, 1893)</named-content></p><sec sec-type="treatment-First records from Mallorca"><title>First records from Mallorca.</title><p>Esporles, IMEDEA research center, <named-content content-type="dwc:verbatimCoordinates">39.666438°N, 2.580863°E</named-content>, <named-content content-type="date">13.iv.2015</named-content>, M. Stokkan leg. <named-content content-type="specimen-count">1♀</named-content> (ZMFK);</p><p>E of Puigpunyent, <named-content content-type="dwc:verbatimCoordinates">39.616667°N, 2.85°E</named-content>, <named-content content-type="date">1.x.1979</named-content>, 200m a.s.l., H. Malicky leg. <named-content content-type="specimen-count">1♀</named-content>; same but <named-content content-type="date">6.x.1979</named-content>, <named-content content-type="specimen-count">4♀♀</named-content> (all <abbrev content-type="institution" xlink:title="Private collection of Rüdiger Wagner, Kassel">RW</abbrev>).</p></sec><pmc-comment>PageBreak</pmc-comment>
<sec sec-type="treatment-Remarks"><title>Remarks.</title><p>A widespread near-cosmopolitan species, first recorded from Spain by <xref rid="B21" ref-type="bibr">Tonnoir (1920)</xref> under the synonym <italic><named-content content-type="taxon-name"><named-content content-type="genus">Telmatoscopus</named-content><named-content content-type="species">meridionalis</named-content></named-content></italic> (Eaton, 1894). Its biology is summarised in <xref rid="B3" ref-type="bibr">Boumans et al. (2009)</xref> and <xref rid="B12" ref-type="bibr">Ježek et al. (2012)</xref>; for a review of its taxonomy see <xref rid="B6" ref-type="bibr">Ibañez-Bernal (2008)</xref>.</p></sec></sec><sec sec-type="taxon-treatment"><sec-meta><kwd-group><label>Taxon classification</label><kwd><named-content content-type="kingdom">Animalia</named-content></kwd><kwd><named-content content-type="order">Diptera</named-content></kwd><kwd><named-content content-type="family">Psychodidae</named-content></kwd></kwd-group></sec-meta><label>*</label><title><named-content content-type="taxon-name"><named-content content-type="genus">Lepiseodina</named-content><named-content content-type="species">rothschildi</named-content></named-content></title><p><named-content content-type="taxon-authority">(Eaton, 1912)</named-content></p><sec sec-type="treatment-First records from Mallorca"><title>First record from Mallorca.</title><p>SW Pollensa, <named-content content-type="dwc:verbatimCoordinates">39.8833°N, 2.9833°E</named-content>, <named-content content-type="date">3–5.x.1981</named-content>, H. Malicky leg. <named-content content-type="specimen-count">1♂</named-content> (<abbrev content-type="institution" xlink:title="Private collection of Rüdiger Wagner, Kassel">RW</abbrev>).</p></sec><sec sec-type="treatment-Remarks"><title>Remarks.</title><p>Placed in <italic><named-content content-type="taxon-name"><named-content content-type="genus">Lepiseodina</named-content></named-content></italic> Enderlein, 1937 (type species <italic><named-content content-type="taxon-name"><named-content content-type="genus">Psychoda</named-content><named-content content-type="species">tristis</named-content></named-content></italic> Meigen, 1830) by <xref rid="B9" ref-type="bibr">Ježek (1990)</xref> because of its asymmetric genitalia. The placement in <italic><named-content content-type="taxon-name"><named-content content-type="genus">Clogmia</named-content></named-content></italic> by <xref rid="B8" ref-type="bibr">Ježek (1984)</xref>, <xref rid="B24" ref-type="bibr">Vaillant (1989)</xref> and <xref rid="B29" ref-type="bibr">Wagner (2004)</xref> may be a valid option to consider for the future when further character systems are explored.</p><p>Previous records are from Austria, Belgium, Czech Republic, France, Germany, Great Britain, Ireland and Slovakia (<xref rid="B18" ref-type="bibr">Oboňa and Ježek 2012</xref>).</p></sec></sec><sec sec-type="taxon-treatment"><sec-meta><kwd-group><label>Taxon classification</label><kwd><named-content content-type="kingdom">Animalia</named-content></kwd><kwd><named-content content-type="order">Diptera</named-content></kwd><kwd><named-content content-type="family">Psychodidae</named-content></kwd></kwd-group></sec-meta><label>*</label><title><named-content content-type="taxon-name"><named-content content-type="genus">Paramormia</named-content><named-content content-type="species">ustulata</named-content></named-content></title><p><named-content content-type="taxon-authority">(Walker, 1856)</named-content></p><sec sec-type="treatment-First records from Mallorca"><title>First records from Mallorca.</title><p>Banyalbufar, <named-content content-type="dwc:verbatimCoordinates">39.691635°N, 2.514367°E</named-content>, <named-content content-type="date">25.x.2012</named-content>, G.Kvifte leg. <named-content content-type="specimen-count">1♂</named-content><named-content content-type="specimen-count">1♀</named-content> (<abbrev content-type="institution" xlink:title="Entomology Collections, Dept. of Natural History, University Museum of Bergen, Bergen, Norway">ZMUB</abbrev>);</p><p>Pond west of Cala Figuera, <named-content content-type="dwc:verbatimCoordinates">39.335635°N, 3.152597°E</named-content>, <named-content content-type="date">11.ii.2015</named-content>, G.Kvifte leg, <named-content content-type="specimen-count">1♂</named-content> (<abbrev content-type="institution" xlink:title="Alexander-König Zoologischer Forschungsmuseum, Bonn, Germany">ZFMK</abbrev>).</p></sec><sec sec-type="treatment-Remarks"><title>Remarks.</title><p>A widespread species or complex of species occurring in the Holarctic region. <xref rid="B11" ref-type="bibr">Ježek and Yağci (2005)</xref> list occurences from the following countries: Afghanistan, Algeria, the Azores, Belgium, Bosnia-Herzegovina, Bulgaria, the Canary Islands, China, Corsica, Czech Republic, Denmark, France, Germany, Great Britain, Greece, Hungary, Iran, Ireland, Israel, Italy, Macedonia, Madeira, Mongolia, Morocco, the Netherlands, Olanda island, Poland, Romania, Sardinia, Serbia, Slovakia, Slovenia, Spain, Sweden, Switzerland, Tunisia, Turkey and the USA.</p></sec></sec></sec><sec sec-type="Pericomaini"><title><named-content content-type="taxon-name"><named-content content-type="tribe">Pericomaini</named-content></named-content></title><sec sec-type="taxon-treatment"><sec-meta><kwd-group><label>Taxon classification</label><kwd><named-content content-type="kingdom">Animalia</named-content></kwd><kwd><named-content content-type="order">Diptera</named-content></kwd><kwd><named-content content-type="family">Psychodidae</named-content></kwd></kwd-group></sec-meta><title><named-content content-type="taxon-name" xlink:href="http://zoobank.org/14619D1C-5A7E-4197-B99C-2C004DE54E83"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">unipennata</named-content></named-content></title><p><named-content content-type="taxon-authority">Kvifte, Stokkan & Wagner</named-content><named-content content-type="taxon-status">sp. n.</named-content></p><p>http://zoobank.org/14619D1C-5A7E-4197-B99C-2C004DE54E83</p><sec sec-type="treatment-type material"><title>Type material.</title><p><named-content content-type="dwc:typeStatus">Holotype</named-content> male. Deiá, town fountain, <named-content content-type="dwc:verbatimCoordinates">39.748072°N, 2.643385°E</named-content>, <named-content content-type="date">8.ii.2015</named-content>, G.Kvifte, M. Stokkan & C. Garcia leg (<abbrev content-type="institution" xlink:title="Entomology Collections, Dept. of Natural History, University Museum of Bergen, Bergen, Norway">ZMUB</abbrev>).</p><p><bold>Diagnosis.</bold><italic><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">unipennata</named-content></named-content></italic> can be separated from all other <italic><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content></named-content></italic> species on the presence of one feather-tipped and four spatulate tenacula on each surstylus, as well as the following combination of characters: parameral sheath with shallow U-sha<pmc-comment>PageBreak</pmc-comment>ped apical indentation 1/6th as deep as width of parameral appendage base, gonostyle with narrow distal part 1/4th as long as broad basal part, distiphallic spatula with sclerotized side margins not converging apically, parameral appendage concave at lateral sides with distal 1/8th protruding over distiphallic spatula.</p></sec><sec sec-type="treatment-Description"><title>Description.</title><p>Male (n=1). Head (Fig. <xref ref-type="fig" rid="F1">1A</xref>) longer than wide; vertex rounded with posterior lobe pointed, about half length of head; eyebridge comprising four rows of facets, separated by five facet diameters; interocular suture broadly V-shaped with median swelling; 7 supraocular setae present, 3 on ventral side and 4 on dorsal side; frontal scar patch inversely T-shaped, reaching level of uppermost facet row of eyebridge; clypeus subrectangular, weakly concave at anterior margin, densely setose, not <pmc-comment>PageBreak</pmc-comment>protruding in front of level of eyes; palp with 4th segment fleshy, corrugated, length of palpus segments 93 : 106 : 129 : 231; labellum fleshy, micropilose with 11 larger setae; antennae with 14 fusiform flagellomeres; scape cylindrical, pedicel elongate globular to barrell-shaped; ascoids present on flagellomeres 4-8; flagellomere 14 with apiculus about as long as base of segment; length of antennal segments 69 : 63 : 48 : 45 : 45 : 42 : 42 : 42 : 42 : 42 : 36 : 36 : 33 : 27 : 24 : 36.</p><fig id="F1" orientation="portrait" position="float"><label>Figure 1.</label><caption><p><italic><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">unipennata</named-content></named-content></italic> sp. n., holotype male. <bold>A</bold> Head <bold>B</bold> Wing <bold>C</bold> Aedeagus <bold>D</bold> Male genitalia <bold>E</bold> Tenacula. The following abbreviations are used: <abbrev>bp<def><p> – basiphallus</p></def></abbrev>, <abbrev>c r<def><p> – cercal region</p></def></abbrev>, <abbrev>dp lm<def><p> – distiphallic lateral margin</p></def></abbrev>, <abbrev>dp s<def><p> – distiphallic spatula</p></def></abbrev>, <abbrev>f t<def><p> – feathery tenacula</p></def></abbrev>, <abbrev>gcx<def><p> – gonocoxite</p></def></abbrev>, <abbrev>gst<def><p> – gonostyle</p></def></abbrev>, <abbrev>hyp<def><p> – hypandrium</p></def></abbrev>, <abbrev>pm<def><p> – parameral sclerite</p></def></abbrev>, <abbrev>pm j<def><p> – parameral joint</p></def></abbrev>, <abbrev>pms<def><p> – parameral sheath</p></def></abbrev>, <abbrev>s t<def><p> – spatuliform tenacula</p></def></abbrev>.</p></caption><graphic id="oo_84031.jpg" xlink:href="zookeys-577-149-g001"></graphic></fig><p>Thorax without accessory organs; mesonotum and scutellum covered in setae alveoli except on lateral margins; anepisternum and laterotergite covered with setae alveoli; coxae and trochanters with dorsal and ventral stripes of setae alveoli, mid coxa with apicoanterior setose projection; legs without special features;</p><p>Wing (Fig. <xref ref-type="fig" rid="F1">1B</xref>) 2.3 mm long, subovate, membrane without setation or infuscation; radial fork very slightly distad of medial fork, both slightly distad of CuA<sub>2</sub>; C with two breaks; Sc weakly curved towards C; apical section of R<sub>2</sub> curved anteriorly; wing apex between R<sub>4</sub> and R<sub>5</sub>; medial fork incomplete; apical section of CuA<sub>2</sub> curved posteriorly; jugum obtuse;</p><p>Genitalia (Fig. <xref ref-type="fig" rid="F1">1C, D</xref>) with hypandrium of even width; aedeagus (Fig. <xref ref-type="fig" rid="F1">1D</xref>) with basiphallus compressed laterally; distiphallus consisting of two phallomeres forming a spatula with sclerotized lateral margins, lateral margins not converging distally; parameral sheath fused with gonocoxites basally, distally narrowing to nearly half its width at base; distally with narrow median incision; parameral sclerite meeting distal section of basiphallus, jointed to parameral sheath with slightly curved transverse band-like sclerite; parameral sclerite half length of distiphallus, consisting of two fused sclerites which are concave laterally, median suture complete; gonocoxites (Fig. <xref ref-type="fig" rid="F1">1C</xref>) stout, reniform, parabasal process absent but setose field present near connection with parameral sheath; gonocoxal condyles apparently fused with parameral sheath, gonostyle (Fig. <xref ref-type="fig" rid="F1">1C</xref>) with basal four 5ths liver-shaped, stout, distal 5th pointed, weakly sinous; epandrium slightly wider than long, expanding distally, number of apertures not discernable in specimen, carrying sparse hairs on ventral surface; surstyli about as long as epandrium, with lateral inner third more weakly sclerotized and densely micropilose; apex of surstylus with five aseriate tenacula of which one apical is feathery at the apex, four subapical tenacula spatulate; hypoproct obtusely isosceles trapezoid, micropilose; epiproct oval, densely pilose.</p></sec><sec sec-type="treatment-Etymology"><title>Etymology.</title><p><italic>unipennata</italic> = with one feather; refers to the presence of a single feather-tipped and four spatulate tenacula on its surstyli.</p></sec><sec sec-type="treatment-Biology"><title>Biology.</title><p>The specimen was collected at a spring with many bryophytes growing in a seepage stream. Bryophyte material was collected and extracted, however, no psychodid larvae or pupae were found.</p></sec></sec><sec sec-type="taxon-treatment"><sec-meta><kwd-group><label>Taxon classification</label><kwd><named-content content-type="kingdom">Animalia</named-content></kwd><kwd><named-content content-type="order">Diptera</named-content></kwd><kwd><named-content content-type="family">Psychodidae</named-content></kwd></kwd-group></sec-meta><title><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">barbarica</named-content></named-content></title><p><named-content content-type="taxon-authority">Vaillant, 1955</named-content></p><sec sec-type="treatment-New records"><title>New records.</title><p>Calobra, <named-content content-type="dwc:verbatimCoordinates">39.85°N, 2.8°E</named-content>, <named-content content-type="date">9.v.1978</named-content>, 90m a.s.l., H. Malicky leg. <named-content content-type="specimen-count">1 ♂</named-content> (<abbrev content-type="institution" xlink:title="Private collection of Rüdiger Wagner, Kassel">RW</abbrev>);</p><p>Deiá, town fountain, <named-content content-type="dwc:verbatimCoordinates">39.748072°N, 2.643385°E</named-content>, <named-content content-type="date">8.ii.2015</named-content>, G.Kvifte, M. Stokkan & C. Garcia leg. <named-content content-type="specimen-count">3♂♂</named-content> (<abbrev content-type="institution" xlink:title="Entomology Collections, Dept. of Natural History, University Museum of Bergen, Bergen, Norway">ZMUB</abbrev>);</p><pmc-comment>PageBreak</pmc-comment>
<p>Esporles, Torrent de San Vic, <named-content content-type="dwc:verbatimCoordinates">39.670459°N, 2.569193°E</named-content>, <named-content content-type="date">11.ii.2015</named-content>, G.Kvifte leg. <named-content content-type="specimen-count">1♂</named-content> (ZMFK);</p><p>Puigpunyent, <named-content content-type="dwc:verbatimCoordinates">39.616667°N, 2.85°E</named-content>, <named-content content-type="date">15.v.1978</named-content>, 200m a.s.l., H. Malicky leg. <named-content content-type="specimen-count">12 ♂♂</named-content>; same but <named-content content-type="date">1.x.1979</named-content>, <named-content content-type="specimen-count">5 ♂♂</named-content>; same but <named-content content-type="date">6.x.1979</named-content>, <named-content content-type="specimen-count">2 ♂♂</named-content> (all <abbrev content-type="institution" xlink:title="Private collection of Rüdiger Wagner, Kassel">RW</abbrev>);</p><p>South slope of Piug Major, north of Soller, <named-content content-type="dwc:verbatimCoordinates">39.783333°N, 2.766667°E</named-content>, <named-content content-type="date">7.–9.v.1978</named-content>, 700 m a.s.l., H. Malicky leg. <named-content content-type="specimen-count">11♂♂</named-content> (<abbrev content-type="institution" xlink:title="Private collection of Rüdiger Wagner, Kassel">RW</abbrev>).</p></sec><sec sec-type="treatment-Literature record"><title>Literature record.</title><p><xref rid="B23" ref-type="bibr">Vaillant (1978)</xref>.</p></sec><sec sec-type="treatment-Remarks"><title>Remarks.</title><p>This species was described from Algeria and subsequently recorded from Mallorca by <xref rid="B23" ref-type="bibr">Vaillant (1978)</xref>. It also occurs in mainland Spain and in North Africa (<xref rid="B23" ref-type="bibr">Vaillant 1978</xref>).</p></sec></sec><sec sec-type="taxon-treatment"><sec-meta><kwd-group><label>Taxon classification</label><kwd><named-content content-type="kingdom">Animalia</named-content></kwd><kwd><named-content content-type="order">Diptera</named-content></kwd><kwd><named-content content-type="family">Psychodidae</named-content></kwd></kwd-group></sec-meta><title><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">trifasciata</named-content></named-content></title><p><named-content content-type="taxon-authority">(Meigen, 1818)</named-content></p><sec sec-type="treatment-Literature record"><title>Literature record.</title><p><xref rid="B31" ref-type="bibr">Wagner et al. (2002)</xref>.</p></sec><sec sec-type="treatment-Remarks"><title>Remarks.</title><p><xref rid="B23" ref-type="bibr">Vaillant (1978)</xref> wrote “[<italic><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content></named-content></italic>] <italic><named-content content-type="taxon-name"><named-content content-type="species">trifasciata</named-content></named-content></italic> ist ebenfalls gemein auf der Mallorca-Insel [...]” (“<italic><named-content content-type="taxon-name"><named-content content-type="species">trifasciata</named-content></named-content></italic> is also common on the island of Mallorca”) in the paragraph summarizing the geographical distribution of <italic><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">barbarica</named-content></named-content></italic> Vaillant, 1955; this is probably a lapsus as Mallorca is not mentioned in the species account for <italic><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">trifasciata</named-content></named-content></italic>. No other records are available in the literature prior to the listing by <xref rid="B31" ref-type="bibr">Wagner et al. (2002)</xref>, which was based on <xref rid="B23" ref-type="bibr">Vaillant (1978)</xref>. In <xref rid="B29" ref-type="bibr">Wagner (2004)</xref> and the present paper we deem the records from the Baleares to be doubtful and in need of verification through examination of specimens.</p></sec></sec></sec><sec sec-type="Psychodini"><title><named-content content-type="taxon-name"><named-content content-type="tribe">Psychodini</named-content></named-content></title><sec sec-type="taxon-treatment"><sec-meta><kwd-group><label>Taxon classification</label><kwd><named-content content-type="kingdom">Animalia</named-content></kwd><kwd><named-content content-type="order">Diptera</named-content></kwd><kwd><named-content content-type="family">Psychodidae</named-content></kwd></kwd-group></sec-meta><label>*</label><title><named-content content-type="taxon-name"><named-content content-type="genus">Philosepedon</named-content><named-content content-type="species">pyrenaicus</named-content></named-content></title><p><named-content content-type="taxon-authority">Vaillant, 1974</named-content></p><sec sec-type="treatment-First records from Mallorca"><title>First records from Mallorca.</title><p>Deiá, town fountain, <named-content content-type="dwc:verbatimCoordinates">39.748072°N, 2.643385°E</named-content>, <named-content content-type="date">8.ii.2015</named-content>, G.Kvifte leg. <named-content content-type="specimen-count">1♂</named-content> (ZMFK);</p><p>Esporles, town fountain, <named-content content-type="dwc:verbatimCoordinates">39.669519°N, 2.576953°E</named-content>, <named-content content-type="date">10.ii.2015</named-content>, G.Kvifte leg. <named-content content-type="specimen-count">1♂</named-content> (ZMFK);</p><p>Palma de Mallorca, <named-content content-type="dwc:verbatimCoordinates">39.570725°N, 2.641432°E</named-content>, <named-content content-type="date">7.ii.2015</named-content>, G.Kvifte leg. <named-content content-type="specimen-count">3♂</named-content><named-content content-type="specimen-count">1♀</named-content> (<named-content content-type="specimen-count">2♂</named-content> ZMFK, <named-content content-type="specimen-count">1♂</named-content><named-content content-type="specimen-count">1♀</named-content><abbrev content-type="institution" xlink:title="Entomology Collections, Dept. of Natural History, University Museum of Bergen, Bergen, Norway">ZMUB</abbrev>).</p></sec><sec sec-type="treatment-Remarks"><title>Remarks.</title><p>The males recorded here were identified as <italic><named-content content-type="taxon-name"><named-content content-type="genus">Philosepedon</named-content><named-content content-type="species">pyrenaicus</named-content></named-content></italic> according to the key in <xref rid="B22" ref-type="bibr">Vaillant (1974)</xref> because of the eyes separated by three facet diameters, the wing “mittwinkel” 105° and the left phallomere of the aedeagus being longer than the right. However, the specimens differ in that the 1st flagellomere is longer than the combined length of the scape and pedicel, and the eyebridge is two facet rows wide at several points. We deem these differences not to be taxonomically meaningful, as the differences in proportions and eyebridge composition could be interpreted as interspecific variation.</p><pmc-comment>PageBreak</pmc-comment>
</sec></sec><sec sec-type="taxon-treatment"><sec-meta><kwd-group><label>Taxon classification</label><kwd><named-content content-type="kingdom">Animalia</named-content></kwd><kwd><named-content content-type="order">Diptera</named-content></kwd><kwd><named-content content-type="family">Psychodidae</named-content></kwd></kwd-group></sec-meta><title><named-content content-type="taxon-name"><named-content content-type="genus">Psychoda</named-content>(<named-content content-type="subgenus">Psychodocha</named-content>)
<named-content content-type="species">cinerea</named-content></named-content></title><p><named-content content-type="taxon-authority">Banks, 1894</named-content></p><sec sec-type="treatment-Literature record"><title>Literature record.</title><p><xref rid="B29" ref-type="bibr">Wagner (2004)</xref></p></sec><sec sec-type="treatment-New records"><title>New records.</title><p>Cala Figuera, <named-content content-type="dwc:verbatimCoordinates">39.332245°N, 3.166440°E</named-content>, <named-content content-type="date">11.ii.2015</named-content>, G. Kvifte leg. <named-content content-type="specimen-count">2♂♂</named-content> (ZMFK);</p><p>Deiá, town fountain, <named-content content-type="dwc:verbatimCoordinates">39.748072°N, 2.643385°E</named-content>, <named-content content-type="date">8.ii.2015</named-content>, G.Kvifte, M. Stokkan & C. Garcia leg. <named-content content-type="specimen-count">1♂</named-content> (<abbrev content-type="institution" xlink:title="Entomology Collections, Dept. of Natural History, University Museum of Bergen, Bergen, Norway">ZMUB</abbrev>);</p><p>Esporles, Torrent de San Vic, <named-content content-type="dwc:verbatimCoordinates">39.670459°N, 2.569193°E</named-content>, <named-content content-type="date">10.ii.2015</named-content>, G.Kvifte leg. <named-content content-type="specimen-count">1♂</named-content> (ZMFK).</p></sec></sec><sec sec-type="taxon-treatment"><sec-meta><kwd-group><label>Taxon classification</label><kwd><named-content content-type="kingdom">Animalia</named-content></kwd><kwd><named-content content-type="order">Diptera</named-content></kwd><kwd><named-content content-type="family">Psychodidae</named-content></kwd></kwd-group></sec-meta><title><named-content content-type="taxon-name"><named-content content-type="genus">Psychoda</named-content>(<named-content content-type="subgenus">Psychodocha</named-content>)
<named-content content-type="species">gemina</named-content></named-content></title><p><named-content content-type="taxon-authority">(Eaton, 1904)</named-content></p><sec sec-type="treatment-Literature record"><title>Literature record.</title><p><xref rid="B29" ref-type="bibr">Wagner (2004)</xref></p></sec></sec><sec sec-type="taxon-treatment"><sec-meta><kwd-group><label>Taxon classification</label><kwd><named-content content-type="kingdom">Animalia</named-content></kwd><kwd><named-content content-type="order">Diptera</named-content></kwd><kwd><named-content content-type="family">Psychodidae</named-content></kwd></kwd-group></sec-meta><title><named-content content-type="taxon-name"><named-content content-type="genus">Psychoda</named-content>(<named-content content-type="subgenus">Psychoda</named-content>)
<named-content content-type="species">phalaenoides</named-content></named-content></title><p><named-content content-type="taxon-authority">(Linnaeus, 1758)</named-content></p><sec sec-type="treatment-Literature record"><title>Literature record.</title><p><xref rid="B29" ref-type="bibr">Wagner (2004)</xref>.</p></sec></sec><sec sec-type="taxon-treatment"><sec-meta><kwd-group><label>Taxon classification</label><kwd><named-content content-type="kingdom">Animalia</named-content></kwd><kwd><named-content content-type="order">Diptera</named-content></kwd><kwd><named-content content-type="family">Psychodidae</named-content></kwd></kwd-group></sec-meta><label>*</label><title><named-content content-type="taxon-name"><named-content content-type="genus">Psychoda</named-content>(<named-content content-type="subgenus">Psycha</named-content>)
<named-content content-type="species">grisescens</named-content></named-content></title><p><named-content content-type="taxon-authority">Tonnoir, 1922</named-content></p><sec sec-type="treatment-First records from Mallorca"><title>First records from Mallorca.</title><p>Esporles, Torrent de San Vic, <named-content content-type="dwc:verbatimCoordinates">39.670459°N, 2.569193°E</named-content>, <named-content content-type="date">11.ii.2015</named-content>, G.Kvifte leg. <named-content content-type="specimen-count">1♂</named-content>.</p></sec><sec sec-type="treatment-Remarks"><title>Remarks.</title><p>Recorded from Algeria, Austria, Belgium, Bosnia-Herzegovina, Czech Republic, Denmark, the Faroe Islands, France, Germany, Great Britain, Greece, Hungary, Ireland, Italy, Morocco, the Netherlands, Norway, Slovakia, Slovenia, Sweden, Tunisia and Turkey (<xref rid="B2" ref-type="bibr">Andersen 1999</xref>, <xref rid="B10" ref-type="bibr">Ježek 2004</xref>, <xref rid="B11" ref-type="bibr">Ježek and Yağci 2005</xref>).</p></sec></sec><sec sec-type="taxon-treatment"><sec-meta><kwd-group><label>Taxon classification</label><kwd><named-content content-type="kingdom">Animalia</named-content></kwd><kwd><named-content content-type="order">Diptera</named-content></kwd><kwd><named-content content-type="family">Psychodidae</named-content></kwd></kwd-group></sec-meta><title><named-content content-type="taxon-name"><named-content content-type="genus">Psychoda</named-content>(<named-content content-type="subgenus">Psychodula</named-content>)
<named-content content-type="species">minuta</named-content></named-content></title><p><named-content content-type="taxon-authority">Banks, 1894</named-content></p><sec sec-type="treatment-Literature record"><title>Literature record.</title><p><xref rid="B29" ref-type="bibr">Wagner (2004)</xref>.</p></sec><sec sec-type="treatment-New record"><title>New record.</title><p>Esporles, Torrent de San Vic, <named-content content-type="dwc:verbatimCoordinates">39.670459°N, 2.569193°E</named-content>, <named-content content-type="date">10.ii.2015</named-content>, G.Kvifte leg. <named-content content-type="specimen-count">1♂</named-content>.</p></sec></sec><sec sec-type="taxon-treatment"><sec-meta><kwd-group><label>Taxon classification</label><kwd><named-content content-type="kingdom">Animalia</named-content></kwd><kwd><named-content content-type="order">Diptera</named-content></kwd><kwd><named-content content-type="family">Psychodidae</named-content></kwd></kwd-group></sec-meta><title><named-content content-type="taxon-name"><named-content content-type="genus">Psychoda</named-content>(<named-content content-type="subgenus">Tinearia</named-content>)
<named-content content-type="species">alternata</named-content></named-content></title><p><named-content content-type="taxon-authority">+Say, 1824</named-content></p><sec sec-type="treatment-Literature record"><title>Literature record.</title><p><xref rid="B29" ref-type="bibr">Wagner (2004)</xref>.</p></sec><sec sec-type="treatment-New record"><title>New record.</title><p>Esporles, town fountain, <named-content content-type="dwc:verbatimCoordinates">39.669519°N, 2.576953°E</named-content>, <named-content content-type="date">10.ii.2015</named-content>, G.Kvifte leg. <named-content content-type="specimen-count">1♀</named-content> (ZMFK).</p><pmc-comment>PageBreak</pmc-comment>
</sec><sec sec-type="treatment-Remarks"><title>Remarks.</title><p><italic><named-content content-type="taxon-name"><named-content content-type="genus">Tinearia</named-content></named-content></italic> Schellenberg, 1803 was treated as a genus by <xref rid="B29" ref-type="bibr">Wagner (2004)</xref> following <xref rid="B7" ref-type="bibr">Ježek (1977)</xref>. This species group is undoubtedly monophyletic, however based on morphology and some analyses of molecular data it appears to be deeply nested within <italic><named-content content-type="taxon-name"><named-content content-type="genus">Psychoda</named-content></named-content></italic> s.l. (<xref rid="B25" ref-type="bibr">Vaillant 1990</xref>, <xref rid="B5" ref-type="bibr">Espindola et al. 2012</xref>, <xref rid="B16" ref-type="bibr">Kvifte and Andersen 2012</xref>, Kvifte unpubl.). Inclusion of <italic><named-content content-type="taxon-name"><named-content content-type="genus">Tinearia</named-content></named-content></italic> as a subgenus within <italic><named-content content-type="taxon-name"><named-content content-type="genus">Psychoda</named-content></named-content></italic> would thus allow ease of identification of monophyletic units, whereas recognising <italic><named-content content-type="taxon-name"><named-content content-type="genus">Tinearia</named-content></named-content></italic> as a separate taxon would render <italic><named-content content-type="taxon-name"><named-content content-type="genus">Psychoda</named-content></named-content></italic> paraphyletic. For a discussion of the monophyly criterion in supraspecific taxonomy, see <xref rid="B13" ref-type="bibr">Komarek and Beutel (2006)</xref>.</p></sec></sec><sec sec-type="taxon-treatment"><sec-meta><kwd-group><label>Taxon classification</label><kwd><named-content content-type="kingdom">Animalia</named-content></kwd><kwd><named-content content-type="order">Diptera</named-content></kwd><kwd><named-content content-type="family">Psychodidae</named-content></kwd></kwd-group></sec-meta><title><named-content content-type="taxon-name"><named-content content-type="genus">Psychoda</named-content>(<named-content content-type="subgenus">Tinearia</named-content>)
<named-content content-type="species">lativentris</named-content></named-content></title><p><named-content content-type="taxon-authority">Berdén, 1952</named-content></p><sec sec-type="treatment-Literature record"><title>Literature record.</title><p><xref rid="B29" ref-type="bibr">Wagner (2004)</xref></p></sec><sec sec-type="treatment-Remarks"><title>Remarks.</title><p>For genus taxonomy, see remarks under <italic><named-content content-type="taxon-name"><named-content content-type="genus">Psychoda</named-content><named-content content-type="species">alternata</named-content></named-content></italic> above.</p></sec></sec></sec></sec><sec sec-type="Discussion"><title>Discussion</title><p>The material examined in the present study was collected opportunistically and does not reflect the diversity of suitable habitats on Mallorca. Nevertheless, a few preliminary conclusions about the diversity and zoogeographic affinities of the fauna can be made.</p><p>Most of the species encountered are widespread European, Holarctic or even cosmopolitan species (<italic><named-content content-type="taxon-name"><named-content content-type="genus">Clogmia</named-content><named-content content-type="species">albipunctata</named-content></named-content></italic>, <italic><named-content content-type="taxon-name"><named-content content-type="genus">Paramormia</named-content><named-content content-type="species">ustulata</named-content></named-content></italic>, <italic><named-content content-type="taxon-name"><named-content content-type="genus">Psychoda</named-content></named-content></italic> spp. and arguably <italic><named-content content-type="taxon-name"><named-content content-type="genus">Lepiseodina</named-content><named-content content-type="species">rothschildi</named-content></named-content></italic>). Four species appear to have more limited distributions as local West Mediterranean elements, namely <italic><named-content content-type="taxon-name"><named-content content-type="genus">Neoarisemus</named-content><named-content content-type="species">ibericus</named-content></named-content></italic>, <italic><named-content content-type="taxon-name"><named-content content-type="genus">Mormia</named-content><named-content content-type="species">tenebricosa</named-content></named-content></italic>, <italic><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">barbarica</named-content></named-content></italic> and <italic><named-content content-type="taxon-name"><named-content content-type="genus">Philosepedon</named-content><named-content content-type="species">pyrenaicus</named-content></named-content></italic>. A single species, <italic><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">unipennata</named-content></named-content></italic> sp. n., has yet to be recorded outside of Mallorca but may have been overlooked elsewhere.</p><p>Our records of <italic><named-content content-type="taxon-name"><named-content content-type="genus">Neoarisemus</named-content><named-content content-type="species">ibericus</named-content></named-content></italic> and <italic><named-content content-type="taxon-name"><named-content content-type="genus">Philosepedon</named-content><named-content content-type="species">pyrenaicus</named-content></named-content></italic> are the first since the original descriptions, which in both cases were based on very few specimens collected in Northern Spain: Montes Universales and the Pyrenees respectively. The records from Mallorca represent a major range extension for both species, suggesting them to be more widespread than previously expected and that they may have been overlooked elsewhere.</p><p>Both <italic><named-content content-type="taxon-name"><named-content content-type="genus">Mormia</named-content><named-content content-type="species">tenebricosa</named-content></named-content></italic> and <italic><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">barbarica</named-content></named-content></italic> have similar distribution patterns; occurring on the north and south coasts of the West Mediterranean. Both species appear widespread in North Africa, having been recorded from Morocco, Algeria and Tunisia (<xref rid="B22" ref-type="bibr">Vaillant 1974</xref>, <xref rid="B23" ref-type="bibr">1978</xref>, <xref rid="B26" ref-type="bibr">Wagner 1987</xref>). They differ in their European distributions, with the range of <italic><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">barbarica</named-content></named-content></italic> extending northeast to the Pyrenees and <italic><named-content content-type="taxon-name"><named-content content-type="genus">Mormia</named-content><named-content content-type="species">tenebricosa</named-content></named-content></italic> reaching the Western Alps in southern France and Italy.</p><p><xref rid="B22" ref-type="bibr">Vaillant (1974)</xref> mentions consistent minor morphological differences between adult males of <italic><named-content content-type="taxon-name"><named-content content-type="genus">Mormia</named-content><named-content content-type="species">tenebricosa</named-content></named-content></italic> in the North African and European populations, but refrained from using them to delimit species. We consider that these differences warrant further study as they may be evidence of cryptic species or ongoing speciation; DNA sequences will be useful in illuminating this question.</p><pmc-comment>PageBreak</pmc-comment>
<p><italic><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">unipennata</named-content></named-content></italic> sp. n. appears to be a member of the Mediterranean <italic><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">modesta</named-content></named-content></italic> Tonnoir, 1922 species group as defined by <xref rid="B23" ref-type="bibr">Vaillant (1978</xref>, p. 226). In this group, <italic><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">modesta</named-content></named-content></italic> has a wide Mediterranean distribution whereas <italic><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">alhambrana</named-content></named-content></italic> Vaillant, 1978, <italic><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">graecica</named-content></named-content></italic> Vaillant, 1978 and <italic><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">motasi</named-content></named-content></italic> Vaillant, 1978 are localized endemics in Southern Spain, Southern Balkan and the Romanian Carpathian mountains, respectively. The <named-content content-type="taxon-name"><named-content content-type="family">Psychodidae</named-content></named-content> of the Iberian peninsula are too incompletely known to tell whether <italic><named-content content-type="taxon-name"><named-content content-type="genus">Pericoma</named-content><named-content content-type="species">unipennata</named-content></named-content></italic> sp. n. is an island endemic of the Balearic islands or whether it is more widespread in the Mediterranean region.</p></sec><sec sec-type="supplementary-material"><title>Supplementary Material</title><supplementary-material id="zookeys.577.7679-treatment1" content-type="local-data"><caption><title>XML Treatment for
<named-content content-type="genus">Neoarisemus</named-content><named-content content-type="species">ibericus</named-content></title></caption><media xlink:href="zookeys.577.7679-treatment1.xml" mimetype="text" mime-subtype="xml"></media></supplementary-material><supplementary-material id="zookeys.577.7679-treatment2" content-type="local-data"><caption><title>XML Treatment for
<named-content content-type="genus">Mormia</named-content><named-content content-type="species">tenebricosa</named-content></title></caption><media xlink:href="zookeys.577.7679-treatment2.xml" mimetype="text" mime-subtype="xml"></media></supplementary-material><supplementary-material id="zookeys.577.7679-treatment3" content-type="local-data"><caption><title>XML Treatment for
<named-content content-type="genus">Clogmia</named-content><named-content content-type="species">albipunctata</named-content></title></caption><media xlink:href="zookeys.577.7679-treatment3.xml" mimetype="text" mime-subtype="xml"></media></supplementary-material><supplementary-material id="zookeys.577.7679-treatment4" content-type="local-data"><caption><title>XML Treatment for
<named-content content-type="genus">Lepiseodina</named-content><named-content content-type="species">rothschildi</named-content></title></caption><media xlink:href="zookeys.577.7679-treatment4.xml" mimetype="text" mime-subtype="xml"></media></supplementary-material><supplementary-material id="zookeys.577.7679-treatment5" content-type="local-data"><caption><title>XML Treatment for
<named-content content-type="genus">Paramormia</named-content><named-content content-type="species">ustulata</named-content></title></caption><media xlink:href="zookeys.577.7679-treatment5.xml" mimetype="text" mime-subtype="xml"></media></supplementary-material><supplementary-material id="zookeys.577.7679-treatment6" content-type="local-data"><caption><title>XML Treatment for
<named-content content-type="genus">Pericoma</named-content><named-content content-type="species">unipennata</named-content></title></caption><media xlink:href="zookeys.577.7679-treatment6.xml" mimetype="text" mime-subtype="xml"></media></supplementary-material><supplementary-material id="zookeys.577.7679-treatment7" content-type="local-data"><caption><title>XML Treatment for
<named-content content-type="genus">Pericoma</named-content><named-content content-type="species">barbarica</named-content></title></caption><media xlink:href="zookeys.577.7679-treatment7.xml" mimetype="text" mime-subtype="xml"></media></supplementary-material><supplementary-material id="zookeys.577.7679-treatment8" content-type="local-data"><caption><title>XML Treatment for
<named-content content-type="genus">Pericoma</named-content><named-content content-type="species">trifasciata</named-content></title></caption><media xlink:href="zookeys.577.7679-treatment8.xml" mimetype="text" mime-subtype="xml"></media></supplementary-material><supplementary-material id="zookeys.577.7679-treatment9" content-type="local-data"><caption><title>XML Treatment for
<named-content content-type="genus">Philosepedon</named-content><named-content content-type="species">pyrenaicus</named-content></title></caption><media xlink:href="zookeys.577.7679-treatment9.xml" mimetype="text" mime-subtype="xml"></media></supplementary-material><supplementary-material id="zookeys.577.7679-treatment10" content-type="local-data"><caption><title>XML Treatment for
<named-content content-type="genus">Psychoda</named-content>(<named-content content-type="subgenus">Psychodocha</named-content>)
<named-content content-type="species">cinerea</named-content></title></caption><media xlink:href="zookeys.577.7679-treatment10.xml" mimetype="text" mime-subtype="xml"></media></supplementary-material><supplementary-material id="zookeys.577.7679-treatment11" content-type="local-data"><caption><title>XML Treatment for
<named-content content-type="genus">Psychoda</named-content>(<named-content content-type="subgenus">Psychodocha</named-content>)
<named-content content-type="species">gemina</named-content></title></caption><media xlink:href="zookeys.577.7679-treatment11.xml" mimetype="text" mime-subtype="xml"></media></supplementary-material><supplementary-material id="zookeys.577.7679-treatment12" content-type="local-data"><caption><title>XML Treatment for
<named-content content-type="genus">Psychoda</named-content>(<named-content content-type="subgenus">Psychoda</named-content>)
<named-content content-type="species">phalaenoides</named-content></title></caption><media xlink:href="zookeys.577.7679-treatment12.xml" mimetype="text" mime-subtype="xml"></media></supplementary-material><supplementary-material id="zookeys.577.7679-treatment13" content-type="local-data"><caption><title>XML Treatment for
<named-content content-type="genus">Psychoda</named-content>(<named-content content-type="subgenus">Psycha</named-content>)
<named-content content-type="species">grisescens</named-content></title></caption><media xlink:href="zookeys.577.7679-treatment13.xml" mimetype="text" mime-subtype="xml"></media></supplementary-material><supplementary-material id="zookeys.577.7679-treatment14" content-type="local-data"><caption><title>XML Treatment for
<named-content content-type="genus">Psychoda</named-content>(<named-content content-type="subgenus">Psychodula</named-content>)
<named-content content-type="species">minuta</named-content></title></caption><media xlink:href="zookeys.577.7679-treatment14.xml" mimetype="text" mime-subtype="xml"></media></supplementary-material><supplementary-material id="zookeys.577.7679-treatment15" content-type="local-data"><caption><title>XML Treatment for
<named-content content-type="genus">Psychoda</named-content>(<named-content content-type="subgenus">Tinearia</named-content>)
<named-content content-type="species">alternata</named-content></title></caption><media xlink:href="zookeys.577.7679-treatment15.xml" mimetype="text" mime-subtype="xml"></media></supplementary-material><supplementary-material id="zookeys.577.7679-treatment16" content-type="local-data"><caption><title>XML Treatment for
<named-content content-type="genus">Psychoda</named-content>(<named-content content-type="subgenus">Tinearia</named-content>)
<named-content content-type="species">lativentris</named-content></title></caption><media xlink:href="zookeys.577.7679-treatment16.xml" mimetype="text" mime-subtype="xml"></media></supplementary-material></sec></body><back><ack><title>Acknowledgements</title><p>We are grateful to Hans Malicky, Lunz am See, for useful material, to Pelayo Secades, Bergen, for valuable help in arranging GMK’s field stays in Mallorca, and to Cecilia Enriques García, Bergen, for assistance in the field. Michelle Vine, Gold Coast, kindly checked the English. We thank our colleagues Greg Curler, Sergio Ibañez-Bernal, Jan Ježek and Jukka Salmela for their helpful reviews of the manuscript. 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