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<title xml:lang="en">The revised classification of eukaryotes</title>
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<name sortKey="Adl, Sina M" sort="Adl, Sina M" uniqKey="Adl S" first="Sina M." last="Adl">Sina M. Adl</name>
<affiliation>
<nlm:aff id="A1">Department of Soil Science, University of Saskatchewan, Saskatoon, SK S7N 5A8, Canada</nlm:aff>
</affiliation>
<affiliation>
<nlm:aff id="A2">Department of Biology, Dalhousie University, Halifax, NS B3H 4R2, Canada</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Simpson, Alastair G" sort="Simpson, Alastair G" uniqKey="Simpson A" first="Alastair. G." last="Simpson">Alastair. G. Simpson</name>
<affiliation>
<nlm:aff id="A2">Department of Biology, Dalhousie University, Halifax, NS B3H 4R2, Canada</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Lane, Christopher E" sort="Lane, Christopher E" uniqKey="Lane C" first="Christopher E." last="Lane">Christopher E. Lane</name>
<affiliation>
<nlm:aff id="A3">Department of Biological Sciences, University of Rhode Island, Kingston, RI. 02881, USA</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Lukes, Julius" sort="Lukes, Julius" uniqKey="Lukes J" first="Julius" last="Lukeš">Julius Lukeš</name>
<affiliation>
<nlm:aff id="A4">Institute of Parasitology, Biology Center and Faculty of Sciences, University of South Bohemia, České Budějovice, Czech Republic</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Bass, David" sort="Bass, David" uniqKey="Bass D" first="David" last="Bass">David Bass</name>
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<nlm:aff id="A5">Zoology Department, Natural History Museum, London, SW7 5BD, UK</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Bowser, Samuel S" sort="Bowser, Samuel S" uniqKey="Bowser S" first="Samuel S." last="Bowser">Samuel S. Bowser</name>
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<nlm:aff id="A6">Wadsworth Center, New York State Department of Health, Albany, New York 12201, USA</nlm:aff>
</affiliation>
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<name sortKey="Brown, Matt" sort="Brown, Matt" uniqKey="Brown M" first="Matt" last="Brown">Matt Brown</name>
<affiliation>
<nlm:aff id="A7">Department of Biochemistry, Dalhousie University, Halifax, NS B3H 4R2, Canada</nlm:aff>
</affiliation>
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<author>
<name sortKey="Burki, Fabien" sort="Burki, Fabien" uniqKey="Burki F" first="Fabien" last="Burki">Fabien Burki</name>
<affiliation>
<nlm:aff id="A8">Department of Botany, University of British Columbia, Vancouver, BC V6T 1Z4, Canada</nlm:aff>
</affiliation>
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<name sortKey="Dunthorn, Micah" sort="Dunthorn, Micah" uniqKey="Dunthorn M" first="Micah" last="Dunthorn">Micah Dunthorn</name>
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<nlm:aff id="A9">Department of Ecology, University of Kaiserslautern, 67663 Kaiserslautern, Germany</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Hampl, Vladimir" sort="Hampl, Vladimir" uniqKey="Hampl V" first="Vladimir" last="Hampl">Vladimir Hampl</name>
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<nlm:aff id="A10">Department of Parasitology, Charles University, Prague, 128 43 Praha 2, Czech Republic</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Heiss, Aaron" sort="Heiss, Aaron" uniqKey="Heiss A" first="Aaron" last="Heiss">Aaron Heiss</name>
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<nlm:aff id="A2">Department of Biology, Dalhousie University, Halifax, NS B3H 4R2, Canada</nlm:aff>
</affiliation>
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<name sortKey="Hoppenrath, Mona" sort="Hoppenrath, Mona" uniqKey="Hoppenrath M" first="Mona" last="Hoppenrath">Mona Hoppenrath</name>
<affiliation>
<nlm:aff id="A11">Forschungsinstitut Senckenberg, DZMB - Deutsches Zentrum für Marine Biodiversitätsforschung, D-26382 Wilhelmshaven, Germany</nlm:aff>
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<name sortKey="Lara, Enrique" sort="Lara, Enrique" uniqKey="Lara E" first="Enrique" last="Lara">Enrique Lara</name>
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<nlm:aff id="A12">University of Neuchâtel, Institute of Biology, Neuchâtel CH-2009, Switzerland</nlm:aff>
</affiliation>
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<name sortKey="Legall, Line" sort="Legall, Line" uniqKey="Legall L" first="Line" last="Legall">Line Legall</name>
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<nlm:aff id="A13">Muséum National d'Histoire Naturelle, UMR 7138 Systématique, Adaptation et Evolution, Paris, 75231 cedex Paris 05, France</nlm:aff>
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<name sortKey="Lynn, Denis H" sort="Lynn, Denis H" uniqKey="Lynn D" first="Denis H." last="Lynn">Denis H. Lynn</name>
<affiliation>
<nlm:aff id="A14">Department of Integrative Biology, University of Guelph, Guelph, ON, N1G 2W1, Canada</nlm:aff>
</affiliation>
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<name sortKey="Mcmanus, Hilary" sort="Mcmanus, Hilary" uniqKey="Mcmanus H" first="Hilary" last="Mcmanus">Hilary Mcmanus</name>
<affiliation>
<nlm:aff id="A15">Department of Biological Sciences, LeMoyne College, Syracuse, NY 13214, USA</nlm:aff>
</affiliation>
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<name sortKey="Mitchell, Edward A D" sort="Mitchell, Edward A D" uniqKey="Mitchell E" first="Edward A. D." last="Mitchell">Edward A. D. Mitchell</name>
<affiliation>
<nlm:aff id="A12">University of Neuchâtel, Institute of Biology, Neuchâtel CH-2009, Switzerland</nlm:aff>
</affiliation>
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<name sortKey="Mozley Stanridge, Sharon E" sort="Mozley Stanridge, Sharon E" uniqKey="Mozley Stanridge S" first="Sharon E." last="Mozley-Stanridge">Sharon E. Mozley-Stanridge</name>
<affiliation>
<nlm:aff id="A16">Department of Biology, Middle Georgia College, Cochran, GA 31014, USA</nlm:aff>
</affiliation>
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<name sortKey="Parfrey, Laura Wegener" sort="Parfrey, Laura Wegener" uniqKey="Parfrey L" first="Laura Wegener" last="Parfrey">Laura Wegener Parfrey</name>
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<nlm:aff id="A17">Department of Chemistry and Biochemistry, University of Colorado, Boulder, CO 80309, USA</nlm:aff>
</affiliation>
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<name sortKey="Pawlowski, Jan" sort="Pawlowski, Jan" uniqKey="Pawlowski J" first="Jan" last="Pawlowski">Jan Pawlowski</name>
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<nlm:aff id="A18">Department of Genetics and Evolution, University of Geneva, 1211 Geneva 4, Switzerland</nlm:aff>
</affiliation>
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<name sortKey="Rueckert, Sonja" sort="Rueckert, Sonja" uniqKey="Rueckert S" first="Sonja" last="Rueckert">Sonja Rueckert</name>
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<nlm:aff id="A19">School of Life, Sport and Social Sciences, Edinburgh Napier University, Edinburgh, EH11 4BN, UK</nlm:aff>
</affiliation>
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<name sortKey="Shadwick, Lora" sort="Shadwick, Lora" uniqKey="Shadwick L" first="Lora" last="Shadwick">Lora Shadwick</name>
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<nlm:aff id="A20">Department of Biological Sciences, University of Arkansas, Fayetteville, Arkansas 72701, USA</nlm:aff>
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<name sortKey="Schoch, Conrad" sort="Schoch, Conrad" uniqKey="Schoch C" first="Conrad" last="Schoch">Conrad Schoch</name>
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<nlm:aff id="A21">GenBank taxonomy, NIH/NLM/NCBI, Bethesda, Maryland 20892-6510, USA</nlm:aff>
</affiliation>
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<name sortKey="Smirnov, Alexey" sort="Smirnov, Alexey" uniqKey="Smirnov A" first="Alexey" last="Smirnov">Alexey Smirnov</name>
<affiliation>
<nlm:aff id="A22">Department of Invertebrate Zoology, St.Petersburg State University, St.Petersburg 199034, Russia</nlm:aff>
</affiliation>
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<name sortKey="Spiegel, Frederick W" sort="Spiegel, Frederick W" uniqKey="Spiegel F" first="Frederick W." last="Spiegel">Frederick W. Spiegel</name>
<affiliation>
<nlm:aff id="A20">Department of Biological Sciences, University of Arkansas, Fayetteville, Arkansas 72701, USA</nlm:aff>
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<title xml:lang="en" level="a" type="main">The revised classification of eukaryotes</title>
<author>
<name sortKey="Adl, Sina M" sort="Adl, Sina M" uniqKey="Adl S" first="Sina M." last="Adl">Sina M. Adl</name>
<affiliation>
<nlm:aff id="A1">Department of Soil Science, University of Saskatchewan, Saskatoon, SK S7N 5A8, Canada</nlm:aff>
</affiliation>
<affiliation>
<nlm:aff id="A2">Department of Biology, Dalhousie University, Halifax, NS B3H 4R2, Canada</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Simpson, Alastair G" sort="Simpson, Alastair G" uniqKey="Simpson A" first="Alastair. G." last="Simpson">Alastair. G. Simpson</name>
<affiliation>
<nlm:aff id="A2">Department of Biology, Dalhousie University, Halifax, NS B3H 4R2, Canada</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Lane, Christopher E" sort="Lane, Christopher E" uniqKey="Lane C" first="Christopher E." last="Lane">Christopher E. Lane</name>
<affiliation>
<nlm:aff id="A3">Department of Biological Sciences, University of Rhode Island, Kingston, RI. 02881, USA</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Lukes, Julius" sort="Lukes, Julius" uniqKey="Lukes J" first="Julius" last="Lukeš">Julius Lukeš</name>
<affiliation>
<nlm:aff id="A4">Institute of Parasitology, Biology Center and Faculty of Sciences, University of South Bohemia, České Budějovice, Czech Republic</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Bass, David" sort="Bass, David" uniqKey="Bass D" first="David" last="Bass">David Bass</name>
<affiliation>
<nlm:aff id="A5">Zoology Department, Natural History Museum, London, SW7 5BD, UK</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Bowser, Samuel S" sort="Bowser, Samuel S" uniqKey="Bowser S" first="Samuel S." last="Bowser">Samuel S. Bowser</name>
<affiliation>
<nlm:aff id="A6">Wadsworth Center, New York State Department of Health, Albany, New York 12201, USA</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Brown, Matt" sort="Brown, Matt" uniqKey="Brown M" first="Matt" last="Brown">Matt Brown</name>
<affiliation>
<nlm:aff id="A7">Department of Biochemistry, Dalhousie University, Halifax, NS B3H 4R2, Canada</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Burki, Fabien" sort="Burki, Fabien" uniqKey="Burki F" first="Fabien" last="Burki">Fabien Burki</name>
<affiliation>
<nlm:aff id="A8">Department of Botany, University of British Columbia, Vancouver, BC V6T 1Z4, Canada</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Dunthorn, Micah" sort="Dunthorn, Micah" uniqKey="Dunthorn M" first="Micah" last="Dunthorn">Micah Dunthorn</name>
<affiliation>
<nlm:aff id="A9">Department of Ecology, University of Kaiserslautern, 67663 Kaiserslautern, Germany</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Hampl, Vladimir" sort="Hampl, Vladimir" uniqKey="Hampl V" first="Vladimir" last="Hampl">Vladimir Hampl</name>
<affiliation>
<nlm:aff id="A10">Department of Parasitology, Charles University, Prague, 128 43 Praha 2, Czech Republic</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Heiss, Aaron" sort="Heiss, Aaron" uniqKey="Heiss A" first="Aaron" last="Heiss">Aaron Heiss</name>
<affiliation>
<nlm:aff id="A2">Department of Biology, Dalhousie University, Halifax, NS B3H 4R2, Canada</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Hoppenrath, Mona" sort="Hoppenrath, Mona" uniqKey="Hoppenrath M" first="Mona" last="Hoppenrath">Mona Hoppenrath</name>
<affiliation>
<nlm:aff id="A11">Forschungsinstitut Senckenberg, DZMB - Deutsches Zentrum für Marine Biodiversitätsforschung, D-26382 Wilhelmshaven, Germany</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Lara, Enrique" sort="Lara, Enrique" uniqKey="Lara E" first="Enrique" last="Lara">Enrique Lara</name>
<affiliation>
<nlm:aff id="A12">University of Neuchâtel, Institute of Biology, Neuchâtel CH-2009, Switzerland</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Legall, Line" sort="Legall, Line" uniqKey="Legall L" first="Line" last="Legall">Line Legall</name>
<affiliation>
<nlm:aff id="A13">Muséum National d'Histoire Naturelle, UMR 7138 Systématique, Adaptation et Evolution, Paris, 75231 cedex Paris 05, France</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Lynn, Denis H" sort="Lynn, Denis H" uniqKey="Lynn D" first="Denis H." last="Lynn">Denis H. Lynn</name>
<affiliation>
<nlm:aff id="A14">Department of Integrative Biology, University of Guelph, Guelph, ON, N1G 2W1, Canada</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Mcmanus, Hilary" sort="Mcmanus, Hilary" uniqKey="Mcmanus H" first="Hilary" last="Mcmanus">Hilary Mcmanus</name>
<affiliation>
<nlm:aff id="A15">Department of Biological Sciences, LeMoyne College, Syracuse, NY 13214, USA</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Mitchell, Edward A D" sort="Mitchell, Edward A D" uniqKey="Mitchell E" first="Edward A. D." last="Mitchell">Edward A. D. Mitchell</name>
<affiliation>
<nlm:aff id="A12">University of Neuchâtel, Institute of Biology, Neuchâtel CH-2009, Switzerland</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Mozley Stanridge, Sharon E" sort="Mozley Stanridge, Sharon E" uniqKey="Mozley Stanridge S" first="Sharon E." last="Mozley-Stanridge">Sharon E. Mozley-Stanridge</name>
<affiliation>
<nlm:aff id="A16">Department of Biology, Middle Georgia College, Cochran, GA 31014, USA</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Parfrey, Laura Wegener" sort="Parfrey, Laura Wegener" uniqKey="Parfrey L" first="Laura Wegener" last="Parfrey">Laura Wegener Parfrey</name>
<affiliation>
<nlm:aff id="A17">Department of Chemistry and Biochemistry, University of Colorado, Boulder, CO 80309, USA</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Pawlowski, Jan" sort="Pawlowski, Jan" uniqKey="Pawlowski J" first="Jan" last="Pawlowski">Jan Pawlowski</name>
<affiliation>
<nlm:aff id="A18">Department of Genetics and Evolution, University of Geneva, 1211 Geneva 4, Switzerland</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Rueckert, Sonja" sort="Rueckert, Sonja" uniqKey="Rueckert S" first="Sonja" last="Rueckert">Sonja Rueckert</name>
<affiliation>
<nlm:aff id="A19">School of Life, Sport and Social Sciences, Edinburgh Napier University, Edinburgh, EH11 4BN, UK</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Shadwick, Lora" sort="Shadwick, Lora" uniqKey="Shadwick L" first="Lora" last="Shadwick">Lora Shadwick</name>
<affiliation>
<nlm:aff id="A20">Department of Biological Sciences, University of Arkansas, Fayetteville, Arkansas 72701, USA</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Schoch, Conrad" sort="Schoch, Conrad" uniqKey="Schoch C" first="Conrad" last="Schoch">Conrad Schoch</name>
<affiliation>
<nlm:aff id="A21">GenBank taxonomy, NIH/NLM/NCBI, Bethesda, Maryland 20892-6510, USA</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Smirnov, Alexey" sort="Smirnov, Alexey" uniqKey="Smirnov A" first="Alexey" last="Smirnov">Alexey Smirnov</name>
<affiliation>
<nlm:aff id="A22">Department of Invertebrate Zoology, St.Petersburg State University, St.Petersburg 199034, Russia</nlm:aff>
</affiliation>
</author>
<author>
<name sortKey="Spiegel, Frederick W" sort="Spiegel, Frederick W" uniqKey="Spiegel F" first="Frederick W." last="Spiegel">Frederick W. Spiegel</name>
<affiliation>
<nlm:aff id="A20">Department of Biological Sciences, University of Arkansas, Fayetteville, Arkansas 72701, USA</nlm:aff>
</affiliation>
</author>
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<title level="j">The Journal of eukaryotic microbiology</title>
<idno type="ISSN">1066-5234</idno>
<idno type="eISSN">1550-7408</idno>
<imprint>
<date when="2012">2012</date>
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<div type="abstract" xml:lang="en">
<p id="P2">This revision of the classification of eukaryotes, which updates that of
<xref rid="R1" ref-type="bibr">Adl et al. (2005)</xref>
, retains an emphasis on the protists and incorporates changes since 2005 that have resolved nodes and branches in phylogenetic trees. Whereas the previous revision was successful in re-introducing name stability to the classification, this revision provides a classification for lineages that were then still unresolved. The supergroups have withstood phylogenetic hypothesis testing with some modifications, but despite some progress, problematic nodes at the base of the eukaryotic tree still remain to be statistically resolved. Looking forward, subsequent transformations to our understanding of the diversity of life will be from the discovery of novel lineages in previously under-sampled areas and from environmental genomic information.</p>
</div>
</front>
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<pmc-comment>The publisher of this article does not allow downloading of the full text in XML form.</pmc-comment>
<pmc-dir>properties manuscript</pmc-dir>
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<journal-meta>
<journal-id journal-id-type="nlm-journal-id">9306405</journal-id>
<journal-id journal-id-type="pubmed-jr-id">2391</journal-id>
<journal-id journal-id-type="nlm-ta">J Eukaryot Microbiol</journal-id>
<journal-id journal-id-type="iso-abbrev">J. Eukaryot. Microbiol.</journal-id>
<journal-title-group>
<journal-title>The Journal of eukaryotic microbiology</journal-title>
</journal-title-group>
<issn pub-type="ppub">1066-5234</issn>
<issn pub-type="epub">1550-7408</issn>
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<subject>Article</subject>
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<title-group>
<article-title>The revised classification of eukaryotes</article-title>
</title-group>
<contrib-group>
<contrib contrib-type="author">
<name>
<surname>Adl</surname>
<given-names>Sina M.</given-names>
</name>
<email>sina.adl@usask.ca</email>
<xref ref-type="aff" rid="A1">a</xref>
<xref ref-type="aff" rid="A2">b</xref>
</contrib>
<contrib contrib-type="author">
<name>
<surname>Simpson</surname>
<given-names>Alastair. G.</given-names>
</name>
<xref ref-type="aff" rid="A2">b</xref>
</contrib>
<contrib contrib-type="author">
<name>
<surname>Lane</surname>
<given-names>Christopher E.</given-names>
</name>
<xref ref-type="aff" rid="A3">c</xref>
</contrib>
<contrib contrib-type="author">
<name>
<surname>Lukeš</surname>
<given-names>Julius</given-names>
</name>
<xref ref-type="aff" rid="A4">d</xref>
</contrib>
<contrib contrib-type="author">
<name>
<surname>Bass</surname>
<given-names>David</given-names>
</name>
<xref ref-type="aff" rid="A5">e</xref>
</contrib>
<contrib contrib-type="author">
<name>
<surname>Bowser</surname>
<given-names>Samuel S.</given-names>
</name>
<xref ref-type="aff" rid="A6">f</xref>
</contrib>
<contrib contrib-type="author">
<name>
<surname>Brown</surname>
<given-names>Matt</given-names>
</name>
<xref ref-type="aff" rid="A7">g</xref>
</contrib>
<contrib contrib-type="author">
<name>
<surname>Burki</surname>
<given-names>Fabien</given-names>
</name>
<xref ref-type="aff" rid="A8">h</xref>
</contrib>
<contrib contrib-type="author">
<name>
<surname>Dunthorn</surname>
<given-names>Micah</given-names>
</name>
<xref ref-type="aff" rid="A9">i</xref>
</contrib>
<contrib contrib-type="author">
<name>
<surname>Hampl</surname>
<given-names>Vladimir</given-names>
</name>
<xref ref-type="aff" rid="A10">j</xref>
</contrib>
<contrib contrib-type="author">
<name>
<surname>Heiss</surname>
<given-names>Aaron</given-names>
</name>
<xref ref-type="aff" rid="A2">b</xref>
</contrib>
<contrib contrib-type="author">
<name>
<surname>Hoppenrath</surname>
<given-names>Mona</given-names>
</name>
<xref ref-type="aff" rid="A11">k</xref>
</contrib>
<contrib contrib-type="author">
<name>
<surname>Lara</surname>
<given-names>Enrique</given-names>
</name>
<xref ref-type="aff" rid="A12">l</xref>
</contrib>
<contrib contrib-type="author">
<name>
<surname>leGall</surname>
<given-names>Line</given-names>
</name>
<xref ref-type="aff" rid="A13">m</xref>
</contrib>
<contrib contrib-type="author">
<name>
<surname>Lynn</surname>
<given-names>Denis H.</given-names>
</name>
<xref ref-type="aff" rid="A14">n</xref>
<xref ref-type="author-notes" rid="FN1">1</xref>
</contrib>
<contrib contrib-type="author">
<name>
<surname>McManus</surname>
<given-names>Hilary</given-names>
</name>
<xref ref-type="aff" rid="A15">o</xref>
</contrib>
<contrib contrib-type="author">
<name>
<surname>Mitchell</surname>
<given-names>Edward A. D.</given-names>
</name>
<xref ref-type="aff" rid="A12">l</xref>
</contrib>
<contrib contrib-type="author">
<name>
<surname>Mozley-Stanridge</surname>
<given-names>Sharon E.</given-names>
</name>
<xref ref-type="aff" rid="A16">p</xref>
</contrib>
<contrib contrib-type="author">
<name>
<surname>Parfrey</surname>
<given-names>Laura Wegener</given-names>
</name>
<xref ref-type="aff" rid="A17">q</xref>
</contrib>
<contrib contrib-type="author">
<name>
<surname>Pawlowski</surname>
<given-names>Jan</given-names>
</name>
<xref ref-type="aff" rid="A18">r</xref>
</contrib>
<contrib contrib-type="author">
<name>
<surname>Rueckert</surname>
<given-names>Sonja</given-names>
</name>
<xref ref-type="aff" rid="A19">s</xref>
</contrib>
<contrib contrib-type="author">
<name>
<surname>Shadwick</surname>
<given-names>Lora</given-names>
</name>
<xref ref-type="aff" rid="A20">t</xref>
</contrib>
<contrib contrib-type="author">
<name>
<surname>Schoch</surname>
<given-names>Conrad</given-names>
</name>
<xref ref-type="aff" rid="A21">u</xref>
</contrib>
<contrib contrib-type="author">
<name>
<surname>Smirnov</surname>
<given-names>Alexey</given-names>
</name>
<xref ref-type="aff" rid="A22">v</xref>
</contrib>
<contrib contrib-type="author">
<name>
<surname>Spiegel</surname>
<given-names>Frederick W.</given-names>
</name>
<xref ref-type="aff" rid="A20">t</xref>
</contrib>
</contrib-group>
<aff id="A1">
<label>a</label>
Department of Soil Science, University of Saskatchewan, Saskatoon, SK S7N 5A8, Canada</aff>
<aff id="A2">
<label>b</label>
Department of Biology, Dalhousie University, Halifax, NS B3H 4R2, Canada</aff>
<aff id="A3">
<label>c</label>
Department of Biological Sciences, University of Rhode Island, Kingston, RI. 02881, USA</aff>
<aff id="A4">
<label>d</label>
Institute of Parasitology, Biology Center and Faculty of Sciences, University of South Bohemia, České Budějovice, Czech Republic</aff>
<aff id="A5">
<label>e</label>
Zoology Department, Natural History Museum, London, SW7 5BD, UK</aff>
<aff id="A6">
<label>f</label>
Wadsworth Center, New York State Department of Health, Albany, New York 12201, USA</aff>
<aff id="A7">
<label>g</label>
Department of Biochemistry, Dalhousie University, Halifax, NS B3H 4R2, Canada</aff>
<aff id="A8">
<label>h</label>
Department of Botany, University of British Columbia, Vancouver, BC V6T 1Z4, Canada</aff>
<aff id="A9">
<label>i</label>
Department of Ecology, University of Kaiserslautern, 67663 Kaiserslautern, Germany</aff>
<aff id="A10">
<label>j</label>
Department of Parasitology, Charles University, Prague, 128 43 Praha 2, Czech Republic</aff>
<aff id="A11">
<label>k</label>
Forschungsinstitut Senckenberg, DZMB - Deutsches Zentrum für Marine Biodiversitätsforschung, D-26382 Wilhelmshaven, Germany</aff>
<aff id="A12">
<label>l</label>
University of Neuchâtel, Institute of Biology, Neuchâtel CH-2009, Switzerland</aff>
<aff id="A13">
<label>m</label>
Muséum National d'Histoire Naturelle, UMR 7138 Systématique, Adaptation et Evolution, Paris, 75231 cedex Paris 05, France</aff>
<aff id="A14">
<label>n</label>
Department of Integrative Biology, University of Guelph, Guelph, ON, N1G 2W1, Canada</aff>
<aff id="A15">
<label>o</label>
Department of Biological Sciences, LeMoyne College, Syracuse, NY 13214, USA</aff>
<aff id="A16">
<label>p</label>
Department of Biology, Middle Georgia College, Cochran, GA 31014, USA</aff>
<aff id="A17">
<label>q</label>
Department of Chemistry and Biochemistry, University of Colorado, Boulder, CO 80309, USA</aff>
<aff id="A18">
<label>r</label>
Department of Genetics and Evolution, University of Geneva, 1211 Geneva 4, Switzerland</aff>
<aff id="A19">
<label>s</label>
School of Life, Sport and Social Sciences, Edinburgh Napier University, Edinburgh, EH11 4BN, UK</aff>
<aff id="A20">
<label>t</label>
Department of Biological Sciences, University of Arkansas, Fayetteville, Arkansas 72701, USA</aff>
<aff id="A21">
<label>u</label>
GenBank taxonomy, NIH/NLM/NCBI, Bethesda, Maryland 20892-6510, USA</aff>
<aff id="A22">
<label>v</label>
Department of Invertebrate Zoology, St.Petersburg State University, St.Petersburg 199034, Russia</aff>
<author-notes>
<fn id="FN1" fn-type="present-address">
<label>1</label>
<p id="P1">Present address: Department of Zoology, University of British Columbia, Vancouver, BC, V6T 1Z4, Canada</p>
</fn>
</author-notes>
<pub-date pub-type="nihms-submitted">
<day>23</day>
<month>8</month>
<year>2012</year>
</pub-date>
<pub-date pub-type="ppub">
<month>9</month>
<year>2012</year>
</pub-date>
<pub-date pub-type="pmc-release">
<day>01</day>
<month>9</month>
<year>2013</year>
</pub-date>
<volume>59</volume>
<issue>5</issue>
<fpage>429</fpage>
<lpage>493</lpage>
<pmc-comment>elocation-id from pubmed: 10.1111/j.1550-7408.2012.00644.x</pmc-comment>
<abstract>
<p id="P2">This revision of the classification of eukaryotes, which updates that of
<xref rid="R1" ref-type="bibr">Adl et al. (2005)</xref>
, retains an emphasis on the protists and incorporates changes since 2005 that have resolved nodes and branches in phylogenetic trees. Whereas the previous revision was successful in re-introducing name stability to the classification, this revision provides a classification for lineages that were then still unresolved. The supergroups have withstood phylogenetic hypothesis testing with some modifications, but despite some progress, problematic nodes at the base of the eukaryotic tree still remain to be statistically resolved. Looking forward, subsequent transformations to our understanding of the diversity of life will be from the discovery of novel lineages in previously under-sampled areas and from environmental genomic information.</p>
</abstract>
<kwd-group>
<kwd>Algae</kwd>
<kwd>amoebae</kwd>
<kwd>biodiversity</kwd>
<kwd>ciliates</kwd>
<kwd>flagellates</kwd>
<kwd>fungi</kwd>
<kwd>parasites</kwd>
<kwd>protozoa</kwd>
<kwd>systematics</kwd>
<kwd>taxonomy</kwd>
</kwd-group>
</article-meta>
</front>
</pmc>
</record>

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