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A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)

Identifieur interne : 000268 ( Pmc/Curation ); précédent : 000267; suivant : 000269

A systematic revision of Operclipygus Marseul (Coleoptera, Histeridae, Exosternini)

Auteurs : Michael S. Caterino [États-Unis] ; Alexey K. Tishechkin

Source :

RBID : PMC:3652427

Abstract

We revise the large Neotropical genus Operclipygus Marseul, in the histerid tribe Exosternini (Histeridae: Histerinae). We synonymize 3 species, move 14 species from other genera, sink the genus Tribalister Horn into Operclipygus, and describe 138 species as new, bringing the total to 177 species of Operclipygus. Keys are provided for the identification of all species, and the majority of the species are illustrated by habitus and male genitalia illustrations. The species are diverse throughout tropical South and Central America, with only a few species extending into the temperate parts of North America. The majority of species can be recognized by the presence of a distinct stria or sulcus along the apical margin of the pygidium, though it is not exclusive to the genus. Natural history details for species of Operclipygus are scant, as most specimens have been collected through the use of passive flight interception traps. Many are probably generally associated with decaying vegetation and leaf litter, where they prey on small arthropods. But a small proportion are known inquilines, with social insects such as ants and termites, and also with some burrowing mammals, such as Ctenomys Blainville. The genus now includes the following species groups and species: Operclipygus sulcistrius group [Operclipygus lucanoidessp. n., Operclipygus schmidtisp. n., Operclipygus simplistriussp. n., Operclipygus sulcistrius Marseul, 1870], Operclipygus mirabilis group [Operclipygus mirabilis (Wenzel & Dybas, 1941) comb. n., Operclipygus pustulifersp. n., Operclipygus plaumannisp. n., Operclipygus sinuatussp. n., Operclipygus mutucasp. n., Operclipygus carinistrius (Lewis, 1908) comb. n., Operclipygus parensissp. n., Operclipygus schlingerisp. n.], Operclipygus kerga group [Operclipygus kerga (Marseul, 1870), Operclipygus planifronssp. n., Operclipygus punctistriussp. n.], Operclipygus conquisitus group [Operclipygus bicolorsp. n., Operclipygus conquisitus (Lewis, 1902), Operclipygus friburgius (Marseul, 1864)], Operclipygus impuncticollis group [Operclipygus bickhardtisp. n., Operclipygus britannicussp. n., Operclipygus impuncticollis (Hinton, 1935)], Operclipygus panamensis group [Operclipygus crenatus (Lewis, 1888), Operclipygus panamensis (Wenzel & Dybas, 1941)], Operclipygus sejunctus group [Operclipygus depressus (Hinton, 1935), Operclipygus itoupesp. n., Operclipygus juninensissp. n., Operclipygus peckisp. n., Operclipygus punctiventersp. n., Operclipygus sejunctus (Schmidt, 1896) comb. n., Operclipygus setiventrisPageBreaksp. n.], Operclipygus mortavis group [Operclipygus ecitonissp. n., Operclipygus mortavissp. n., Operclipygus paraguensissp. n.], Operclipygus dytiscoides group [Operclipygus carinisternussp. n., Operclipygus crenulatussp. n., Operclipygus dytiscoidessp. n., Operclipygus quadratussp. n.], Operclipygus dubitabilis group [Operclipygus dubitabilis (Marseul, 1889), Operclipygus yasunisp. n.], Operclipygus angulifer group [Operclipygus angulifersp. n., Operclipygus impressifronssp. n.], Operclipygus dubius group [Operclipygus andinussp. n., Operclipygus dubius (Lewis, 1888), Operclipygus extraneussp. n., Operclipygus intermissussp. n., Operclipygus lunulussp. n., Operclipygus occultussp. n., Operclipygus perplexussp. n., Operclipygus remotussp. n., Operclipygus validussp. n., Operclipygus variabilissp. n.], Operclipygus hospes group [Operclipygus assimilissp. n., Operclipygus belemensissp. n., Operclipygus bulbistomasp. n., Operclipygus callifronssp. n., Operclipygus colombicussp. n., Operclipygus communissp. n., Operclipygus confertussp. n., Operclipygus confluenssp. n., Operclipygus curtistriussp. n., Operclipygus diffluenssp. n., Operclipygus fusistriussp. n., Operclipygus gratussp. n., Operclipygus hospes (Lewis, 1902), Operclipygus ibiscussp. n., Operclipygus ignifersp. n., Operclipygus impositussp. n., Operclipygus incisussp. n., Operclipygus innocuussp. n., Operclipygus inquilinussp. n., Operclipygus minutussp. n., Operclipygus novateutoniaesp. n., Operclipygus praecinctussp. n., Operclipygus prominenssp. n., Operclipygus rileyisp. n., Operclipygus subterraneussp. n., Operclipygus tenuissp. n., Operclipygus tiputinussp. n.], Operclipygus farctus group [Operclipygus atlanticussp. n., Operclipygus bidessois (Marseul, 1889), Operclipygus distinctus (Hinton, 1935), Operclipygus distractus (Schmidt, 1896) comb. n., Operclipygus farctissimussp. n., Operclipygus farctus (Marseul, 1864), Operclipygus gillisp. n., Operclipygus impressistriussp. n., Operclipygus inflatussp. n., Operclipygus latemarginatus (Bickhardt, 1920) comb. n., Operclipygus petrovisp. n., Operclipygus plicatus (Hinton, 1935) comb. n., Operclipygus prolixussp. n., Operclipygus punctifronssp. n., Operclipygus proximussp. n., Operclipygus subrufussp. n.], Operclipygus hirsutipes group [Operclipygus guianensissp. n., Operclipygus hirsutipessp. n.], Operclipygus hamistrius group [Operclipygus arquussp. n., Operclipygus campbellisp. n., Operclipygus chiapensissp. n., Operclipygus dybasisp. n., Operclipygus geometricus (Casey, 1893) comb. n., Operclipygus hamistrius (Schmidt, 1893) comb. n., Operclipygus impressicollissp. n., Operclipygus intersectussp. n., Operclipygus montanussp. n., Operclipygus nubosussp. n., Operclipygus pichinchensissp. n., Operclipygus propinquussp. n., Operclipygus quinquestriatussp. n., Operclipygus rubidus (Hinton, 1935) comb. n., Operclipygus rufescenssp. n., Operclipygus troglodytessp. n.], Operclipygus plicicollis group [Operclipygus cephalicussp. n., Operclipygus longidenssp. n., Operclipygus plicicollis (Schmidt, 1893)], Operclipygus fossipygus group [Operclipygus disconnectussp. n., Operclipygus fossipygus (Wenzel, 1944), Operclipygus foveipygus (Bickhardt, 1918), Operclipygus fungicolus (Wenzel & Dybas, 1941), Operclipygus gibbulus (Schmidt, 1889) comb. n., Operclipygus olivensissp. n., Operclipygus simplicipygussp. n., Operclipygus subdepressus (Schmidt, 1889), Operclipygus therondi (Wenzel, 1976)], Operclipygus impunctipennis group [Operclipygus chamelensissp. n., Operclipygus foveiventrissp. n., Operclipygus granulipectussp. n., Operclipygus impunctipennis (Hinton, 1935) comb. n., Operclipygus latifoveatussp. n., Operclipygus lissipygussp. n., Operclipygus maesisp. n., Operclipygus mangiferussp. n., Operclipygus marginipennissp. n., Operclipygus nicodemussp. n., Operclipygus nitidussp. n., Operclipygus pacificussp. n., Operclipygus pauperculussp. n., Operclipygus punctissipygussp. n., Operclipygus subviridissp. n., Operclipygus tripartitussp. n., Operclipygus voraxsp. n.], Operclipygus marginellus group [Operclipygus asheisp. n., Operclipygus baylessaesp. n., Operclipygus dentatussp. n., Operclipygus formicatussp. n., Operclipygus hintonisp. n., Operclipygus marginellus (J.E. LeConte, 1860) comb. n., Operclipygus orchidophilussp. n., Operclipygus selvorumsp. n., Operclipygus striatellus (Fall, 1917) comb. n.], incertae sedis: O. teapensis (Marseul, 1853) comb. n., Operclipygus punctulatussp. n., Operclipygus lama Mazur, 1988, Operclipygus florifaunensissp. n., Operclipygus bosquesecussp. n., Operclipygus arnaudi Dégallier, 1982, Operclipygus subsphaericussp. n., Operclipygus latipygussp. n., Operclipygus elongatussp. n., Operclipygus rupicolussp. n., Operclipygus punctipleurussp. n., Operclipygus falinisp. n., Operclipygus peregrinussp. n., Operclipygus brooksisp. n., Operclipygus profundipygussp. n., Operclipygus punctatissimussp. n., Operclipygus cavisternussp. n., Operclipygus siluriformissp. n., Operclipygus parallelussp. n., Operclipygus abbreviatussp. n., Operclipygus pygidialis (Lewis, 1908), Operclipygus faltistriussp. n., Operclipygus limonensissp. n., Operclipygus wenzelisp. n., Operclipygus iheringi (Bickhardt, 1917), Operclipygus angustisternus (Wenzel, 1944), Operclipygus shortisp. n. We establish the following synonymies: Phelisteroides miladae Wenzel & Dybas, 1941 and Pseudister propygidialis Hinton, 1935e = Operclipygus crenatus (Lewis, 1888); Phelister subplicatus Schmidt, 1893b = Operclipygus bidessois (Marseul, 1889). We designate lectotypes for Operclipygus sulcistrius Marseul, 1870, Phelister carinistrius Lewis, 1908, Phelister kerga Marseul, 1870, Phelister friburgius Marseul, 1864, Phelister impuncticollis Hinton, 1935, Phelister crenatus Lewis, 1888, Phelister sejunctus Schmidt, 1896, Pseudister depressus Hinton, 1935, Epierus dubius Lewis, 1888, Phelister hospes Lewis, 1902, Phelister farctus Marseul, 1864, Phelister bidessois Marseul, 1889, Phelister subplicatus Schmidt, 1893, Phelister plicatus Hinton, 1935, Phelister distinctus Hinton, 1935, Phelister distractus Schmidt, 1896, Pseudister latemarginatus Bickhardt, 1920, Phelister hamistrius Schmidt, 1893, Phelister plicicollis Schmidt, 1893, Phelister gibbulus Schmidt, 1889, Phelister subdepressus Schmidt, 1889, Phelister teapensis Marseul, 1853, Phelister pygidialis Lewis, 1908, Phelister iheringi Bickhardt, 1917, and Phelister marginellus J.E. LeConte 1860. We designate a neotype for Operclipygus conquisitus Lewis, replacing its lost type specimen.


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DOI: 10.3897/zookeys.271.4062
PubMed: 23717185
PubMed Central: 3652427

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<name sortKey="Caterino, Michael S" sort="Caterino, Michael S" uniqKey="Caterino M" first="Michael S." last="Caterino">Michael S. Caterino</name>
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<front>
<div type="abstract" xml:lang="en">
<label>Abstract</label>
<p>We revise the large Neotropical genus
<italic>Operclipygus</italic>
Marseul, in the histerid tribe
<named-content content-type="taxon-name">Exosternini</named-content>
(
<named-content content-type="taxon-name">Histeridae</named-content>
:
<named-content content-type="taxon-name">Histerinae</named-content>
). We synonymize 3 species, move 14 species from other genera, sink the genus
<italic>
<named-content content-type="taxon-name">Tribalister</named-content>
</italic>
Horn into
<italic>
<named-content content-type="taxon-name">Operclipygus</named-content>
</italic>
, and describe 138 species as new, bringing the total to 177 species of
<italic>
<named-content content-type="taxon-name">Operclipygus</named-content>
</italic>
. Keys are provided for the identification of all species, and the majority of the species are illustrated by habitus and male genitalia illustrations. The species are diverse throughout tropical South and Central America, with only a few species extending into the temperate parts of North America. The majority of species can be recognized by the presence of a distinct stria or sulcus along the apical margin of the pygidium, though it is not exclusive to the genus. Natural history details for species of
<italic>
<named-content content-type="taxon-name">Operclipygus</named-content>
</italic>
are scant, as most specimens have been collected through the use of passive flight interception traps. Many are probably generally associated with decaying vegetation and leaf litter, where they prey on small arthropods. But a small proportion are known inquilines, with social insects such as ants and termites, and also with some burrowing mammals, such as
<italic>
<named-content content-type="taxon-name">Ctenomys</named-content>
</italic>
Blainville. The genus now includes the following species groups and species:
<italic>
<named-content content-type="taxon-name">Operclipygus sulcistrius</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus lucanoides</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus schmidti</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus simplistrius</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus sulcistrius</named-content>
</italic>
Marseul, 1870],
<italic>
<named-content content-type="taxon-name">Operclipygus mirabilis</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus mirabilis</named-content>
</italic>
(Wenzel & Dybas, 1941) comb. n.,
<italic>
<named-content content-type="taxon-name">Operclipygus pustulifer</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus plaumanni</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus sinuatus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus mutuca</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus carinistrius</named-content>
</italic>
(Lewis, 1908) comb. n.,
<italic>
<named-content content-type="taxon-name">Operclipygus parensis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus schlingeri</named-content>
</italic>
<bold>sp. n.</bold>
],
<italic>
<named-content content-type="taxon-name">Operclipygus kerga</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus kerga</named-content>
</italic>
(Marseul, 1870),
<italic>
<named-content content-type="taxon-name">Operclipygus planifrons</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus punctistrius</named-content>
</italic>
<bold>sp. n.</bold>
],
<italic>
<named-content content-type="taxon-name">Operclipygus conquisitus</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus bicolor</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus conquisitus</named-content>
</italic>
(Lewis, 1902),
<italic>
<named-content content-type="taxon-name">Operclipygus friburgius</named-content>
</italic>
(Marseul, 1864)],
<italic>
<named-content content-type="taxon-name">Operclipygus impuncticollis</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus bickhardti</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus britannicus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus impuncticollis</named-content>
</italic>
(Hinton, 1935)],
<italic>
<named-content content-type="taxon-name">Operclipygus panamensis</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus crenatus</named-content>
</italic>
(Lewis, 1888),
<italic>
<named-content content-type="taxon-name">Operclipygus panamensis</named-content>
</italic>
(Wenzel & Dybas, 1941)],
<italic>
<named-content content-type="taxon-name">Operclipygus sejunctus</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus depressus</named-content>
</italic>
(Hinton, 1935),
<italic>
<named-content content-type="taxon-name">Operclipygus itoupe</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus juninensis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus pecki</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus punctiventer</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus sejunctus</named-content>
</italic>
(Schmidt, 1896) comb. n.,
<italic>
<named-content content-type="taxon-name">Operclipygus setiventris</named-content>
</italic>
<bold>
<pmc-comment>PageBreak</pmc-comment>
sp. n.</bold>
],
<italic>
<named-content content-type="taxon-name">Operclipygus mortavis</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus ecitonis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus mortavis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus paraguensis</named-content>
</italic>
<bold>sp. n.</bold>
],
<italic>
<named-content content-type="taxon-name">Operclipygus dytiscoides</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus carinisternus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus crenulatus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus dytiscoides</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus quadratus</named-content>
</italic>
<bold>sp. n.</bold>
],
<italic>
<named-content content-type="taxon-name">Operclipygus dubitabilis</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus dubitabilis</named-content>
</italic>
(Marseul, 1889),
<italic>
<named-content content-type="taxon-name">Operclipygus yasuni</named-content>
</italic>
<bold>sp. n.</bold>
],
<italic>
<named-content content-type="taxon-name">Operclipygus angulifer</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus angulifer</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus impressifrons</named-content>
</italic>
<bold>sp. n.</bold>
],
<italic>
<named-content content-type="taxon-name">Operclipygus dubius</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus andinus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus dubius</named-content>
</italic>
(Lewis, 1888),
<italic>
<named-content content-type="taxon-name">Operclipygus extraneus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus intermissus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus lunulus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus occultus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus perplexus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus remotus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus validus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus variabilis</named-content>
</italic>
<bold>sp. n.</bold>
],
<italic>
<named-content content-type="taxon-name">Operclipygus hospes</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus assimilis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus belemensis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus bulbistoma</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus callifrons</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus colombicus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus communis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus confertus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus confluens</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus curtistrius</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus diffluens</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus fusistrius</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus gratus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus hospes</named-content>
</italic>
(Lewis, 1902),
<italic>
<named-content content-type="taxon-name">Operclipygus ibiscus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus ignifer</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus impositus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus incisus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus innocuus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus inquilinus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus minutus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus novateutoniae</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus praecinctus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus prominens</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus rileyi</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus subterraneus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus tenuis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus tiputinus</named-content>
</italic>
<bold>sp. n.</bold>
],
<italic>
<named-content content-type="taxon-name">Operclipygus farctus</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus atlanticus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus bidessois</named-content>
</italic>
(Marseul, 1889),
<italic>
<named-content content-type="taxon-name">Operclipygus distinctus</named-content>
</italic>
(Hinton, 1935),
<italic>
<named-content content-type="taxon-name">Operclipygus distractus</named-content>
</italic>
(Schmidt, 1896) comb. n.,
<italic>
<named-content content-type="taxon-name">Operclipygus farctissimus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus farctus</named-content>
</italic>
(Marseul, 1864),
<italic>
<named-content content-type="taxon-name">Operclipygus gilli</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus impressistrius</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus inflatus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus latemarginatus</named-content>
</italic>
(Bickhardt, 1920) comb. n.,
<italic>
<named-content content-type="taxon-name">Operclipygus petrovi</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus plicatus</named-content>
</italic>
(Hinton, 1935) comb. n.,
<italic>
<named-content content-type="taxon-name">Operclipygus prolixus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus punctifrons</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus proximus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus subrufus</named-content>
</italic>
<bold>sp. n.</bold>
],
<italic>
<named-content content-type="taxon-name">Operclipygus hirsutipes</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus guianensis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus hirsutipes</named-content>
</italic>
<bold>sp. n.</bold>
],
<italic>
<named-content content-type="taxon-name">Operclipygus hamistrius</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus arquus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus campbelli</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus chiapensis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus dybasi</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus geometricus</named-content>
</italic>
(Casey, 1893) comb. n.,
<italic>
<named-content content-type="taxon-name">Operclipygus hamistrius</named-content>
</italic>
(Schmidt, 1893) comb. n.,
<italic>
<named-content content-type="taxon-name">Operclipygus impressicollis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus intersectus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus montanus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus nubosus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus pichinchensis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus propinquus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus quinquestriatus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus rubidus</named-content>
</italic>
(Hinton, 1935) comb. n.,
<italic>
<named-content content-type="taxon-name">Operclipygus rufescens</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus troglodytes</named-content>
</italic>
<bold>sp. n.</bold>
],
<italic>
<named-content content-type="taxon-name">Operclipygus plicicollis</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus cephalicus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus longidens</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus plicicollis</named-content>
</italic>
(Schmidt, 1893)],
<italic>
<named-content content-type="taxon-name">Operclipygus fossipygus</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus disconnectus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus fossipygus</named-content>
</italic>
(Wenzel, 1944),
<italic>
<named-content content-type="taxon-name">Operclipygus foveipygus</named-content>
</italic>
(Bickhardt, 1918),
<italic>
<named-content content-type="taxon-name">Operclipygus fungicolus</named-content>
</italic>
(Wenzel & Dybas, 1941),
<italic>
<named-content content-type="taxon-name">Operclipygus gibbulus</named-content>
</italic>
(Schmidt, 1889) comb. n.,
<italic>
<named-content content-type="taxon-name">Operclipygus olivensis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus simplicipygus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus subdepressus</named-content>
</italic>
(Schmidt, 1889),
<italic>
<named-content content-type="taxon-name">Operclipygus therondi</named-content>
</italic>
(Wenzel, 1976)],
<italic>
<named-content content-type="taxon-name">Operclipygus impunctipennis</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus chamelensis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus foveiventris</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus granulipectus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus impunctipennis</named-content>
</italic>
(Hinton, 1935) comb. n.,
<italic>
<named-content content-type="taxon-name">Operclipygus latifoveatus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus lissipygus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus maesi</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus mangiferus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus marginipennis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus nicodemus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus nitidus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus pacificus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus pauperculus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus punctissipygus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus subviridis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus tripartitus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus vorax</named-content>
</italic>
<bold>sp. n.</bold>
],
<italic>
<named-content content-type="taxon-name">Operclipygus marginellus</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus ashei</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus baylessae</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus dentatus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus formicatus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus hintoni</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus marginellus</named-content>
</italic>
(J.E. LeConte, 1860) comb. n.,
<italic>
<named-content content-type="taxon-name">Operclipygus orchidophilus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus selvorum</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus striatellus</named-content>
</italic>
(Fall, 1917) comb. n.],
<italic>incertae sedis: O. teapensis</italic>
(Marseul, 1853) comb. n.,
<italic>
<named-content content-type="taxon-name">Operclipygus punctulatus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus lama</named-content>
</italic>
Mazur, 1988,
<italic>
<named-content content-type="taxon-name">Operclipygus florifaunensis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus bosquesecus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus arnaudi</named-content>
</italic>
Dégallier, 1982,
<italic>
<named-content content-type="taxon-name">Operclipygus subsphaericus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus latipygus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus elongatus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus rupicolus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus punctipleurus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus falini</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus peregrinus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus brooksi</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus profundipygus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus punctatissimus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus cavisternus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus siluriformis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus parallelus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus abbreviatus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus pygidialis</named-content>
</italic>
(Lewis, 1908),
<italic>
<named-content content-type="taxon-name">Operclipygus faltistrius</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus limonensis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus wenzeli</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus iheringi</named-content>
</italic>
(Bickhardt, 1917),
<italic>
<named-content content-type="taxon-name">Operclipygus angustisternus</named-content>
</italic>
(Wenzel, 1944),
<italic>
<named-content content-type="taxon-name">Operclipygus shorti</named-content>
</italic>
<bold>sp. n.</bold>
We establish the following synonymies:
<italic>
<named-content content-type="taxon-name">Phelisteroides miladae</named-content>
</italic>
Wenzel & Dybas, 1941 and
<italic>
<named-content content-type="taxon-name">Pseudister propygidialis</named-content>
</italic>
Hinton, 1935e =
<italic>
<named-content content-type="taxon-name">Operclipygus crenatus</named-content>
</italic>
(Lewis, 1888);
<italic>
<named-content content-type="taxon-name">Phelister subplicatus</named-content>
</italic>
Schmidt, 1893b =
<italic>
<named-content content-type="taxon-name">Operclipygus bidessois</named-content>
</italic>
(Marseul, 1889). We designate lectotypes for
<italic>
<named-content content-type="taxon-name">Operclipygus sulcistrius</named-content>
</italic>
Marseul, 1870,
<italic>
<named-content content-type="taxon-name">Phelister carinistrius</named-content>
</italic>
Lewis, 1908,
<italic>
<named-content content-type="taxon-name">Phelister kerga</named-content>
</italic>
Marseul, 1870,
<italic>
<named-content content-type="taxon-name">Phelister friburgius</named-content>
</italic>
Marseul, 1864,
<italic>
<named-content content-type="taxon-name">Phelister impuncticollis</named-content>
</italic>
Hinton, 1935,
<italic>
<named-content content-type="taxon-name">Phelister crenatus</named-content>
</italic>
Lewis, 1888,
<italic>
<named-content content-type="taxon-name">Phelister sejunctus</named-content>
</italic>
Schmidt, 1896,
<italic>
<named-content content-type="taxon-name">Pseudister depressus</named-content>
</italic>
Hinton, 1935,
<italic>
<named-content content-type="taxon-name">Epierus dubius</named-content>
</italic>
Lewis, 1888,
<italic>
<named-content content-type="taxon-name">Phelister hospes</named-content>
</italic>
Lewis, 1902,
<italic>
<named-content content-type="taxon-name">Phelister farctus</named-content>
</italic>
Marseul, 1864,
<italic>
<named-content content-type="taxon-name">Phelister bidessois</named-content>
</italic>
Marseul, 1889,
<italic>
<named-content content-type="taxon-name">Phelister subplicatus</named-content>
</italic>
Schmidt, 1893,
<italic>
<named-content content-type="taxon-name">Phelister plicatus</named-content>
</italic>
Hinton, 1935,
<italic>
<named-content content-type="taxon-name">Phelister distinctus</named-content>
</italic>
Hinton, 1935,
<italic>
<named-content content-type="taxon-name">Phelister distractus</named-content>
</italic>
Schmidt, 1896,
<italic>
<named-content content-type="taxon-name">Pseudister latemarginatus</named-content>
</italic>
Bickhardt, 1920,
<italic>
<named-content content-type="taxon-name">Phelister hamistrius</named-content>
</italic>
Schmidt, 1893,
<italic>
<named-content content-type="taxon-name">Phelister plicicollis</named-content>
</italic>
Schmidt, 1893,
<italic>
<named-content content-type="taxon-name">Phelister gibbulus</named-content>
</italic>
Schmidt, 1889,
<italic>
<named-content content-type="taxon-name">Phelister subdepressus</named-content>
</italic>
Schmidt, 1889,
<italic>
<named-content content-type="taxon-name">Phelister teapensis</named-content>
</italic>
Marseul, 1853,
<italic>
<named-content content-type="taxon-name">Phelister pygidialis</named-content>
</italic>
Lewis, 1908,
<italic>
<named-content content-type="taxon-name">Phelister iheringi</named-content>
</italic>
Bickhardt, 1917, and
<italic>
<named-content content-type="taxon-name">Phelister marginellus</named-content>
</italic>
J.E. LeConte 1860. We designate a neotype for
<italic>
<named-content content-type="taxon-name">Operclipygus conquisitus</named-content>
</italic>
Lewis, replacing its lost type specimen.</p>
</div>
</front>
<back>
<div1 type="bibliography">
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</div1>
</back>
</TEI>
<pmc article-type="research-article">
<pmc-dir>properties open_access</pmc-dir>
<front>
<journal-meta>
<journal-id journal-id-type="nlm-ta">Zookeys</journal-id>
<journal-id journal-id-type="iso-abbrev">Zookeys</journal-id>
<journal-id journal-id-type="publisher-id">ZooKeys</journal-id>
<journal-title-group>
<journal-title>ZooKeys</journal-title>
</journal-title-group>
<issn pub-type="ppub">1313-2989</issn>
<issn pub-type="epub">1313-2970</issn>
<publisher>
<publisher-name>Pensoft Publishers</publisher-name>
</publisher>
</journal-meta>
<article-meta>
<article-id pub-id-type="pmid">23717185</article-id>
<article-id pub-id-type="pmc">3652427</article-id>
<article-id pub-id-type="doi">10.3897/zookeys.271.4062</article-id>
<article-categories>
<subj-group subj-group-type="heading">
<subject>Article</subject>
</subj-group>
</article-categories>
<title-group>
<article-title>A systematic revision of
<italic>Operclipygus</italic>
Marseul (Coleoptera, Histeridae, Exosternini) </article-title>
</title-group>
<contrib-group>
<contrib contrib-type="author">
<name>
<surname>Caterino</surname>
<given-names>Michael S.</given-names>
</name>
<xref ref-type="aff" rid="A1">1</xref>
<uri content-type="lsid" xlink:type="simple">urn:lsid:zoobank.org:author:F687B1E2-A07D-4F28-B1F5-4A0DD17B6490</uri>
</contrib>
<contrib contrib-type="author">
<name>
<surname>Tishechkin</surname>
<given-names>Alexey K.</given-names>
</name>
<uri content-type="lsid" xlink:type="simple">urn:lsid:zoobank.org:author:341C5592-E307-43B4-978C-066999A6C8B5</uri>
</contrib>
</contrib-group>
<aff id="A1">
<label>1</label>
Department of Invertebrate Zoology, Santa Barbara Museum of Natural History, 2559 Puesta del Sol, Santa Barbara, CA 93105 USA</aff>
<author-notes>
<corresp>Corresponding author: Michael S. Caterino (
<email xlink:type="simple">mcaterino@sbnature2.org</email>
)</corresp>
<fn fn-type="edited-by">
<p>Academic editor: M. Fikácek</p>
</fn>
</author-notes>
<pub-date pub-type="collection">
<year>2013</year>
</pub-date>
<pub-date pub-type="epub">
<day>20</day>
<month>2</month>
<year>2013</year>
</pub-date>
<issue>271</issue>
<fpage>1</fpage>
<lpage>401</lpage>
<history>
<date date-type="received">
<day>28</day>
<month>9</month>
<year>2012</year>
</date>
<date date-type="accepted">
<day>16</day>
<month>1</month>
<year>2013</year>
</date>
</history>
<permissions>
<copyright-statement>Michael S. Caterino, Alexey K. Tishechkin</copyright-statement>
<license license-type="creative-commons-attribution" xlink:href="http://creativecommons.org/licenses/by/3.0">
<license-p>This is an open access article distributed under the terms of the Creative Commons Attribution License 3.0 (CC-BY), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.</license-p>
</license>
</permissions>
<self-uri content-type="lsid" xlink:type="simple">urn:lsid:zoobank.org:pub:541A802C-21F2-4A14-9428-F5F46FFE4760</self-uri>
<abstract>
<label>Abstract</label>
<p>We revise the large Neotropical genus
<italic>Operclipygus</italic>
Marseul, in the histerid tribe
<named-content content-type="taxon-name">Exosternini</named-content>
(
<named-content content-type="taxon-name">Histeridae</named-content>
:
<named-content content-type="taxon-name">Histerinae</named-content>
). We synonymize 3 species, move 14 species from other genera, sink the genus
<italic>
<named-content content-type="taxon-name">Tribalister</named-content>
</italic>
Horn into
<italic>
<named-content content-type="taxon-name">Operclipygus</named-content>
</italic>
, and describe 138 species as new, bringing the total to 177 species of
<italic>
<named-content content-type="taxon-name">Operclipygus</named-content>
</italic>
. Keys are provided for the identification of all species, and the majority of the species are illustrated by habitus and male genitalia illustrations. The species are diverse throughout tropical South and Central America, with only a few species extending into the temperate parts of North America. The majority of species can be recognized by the presence of a distinct stria or sulcus along the apical margin of the pygidium, though it is not exclusive to the genus. Natural history details for species of
<italic>
<named-content content-type="taxon-name">Operclipygus</named-content>
</italic>
are scant, as most specimens have been collected through the use of passive flight interception traps. Many are probably generally associated with decaying vegetation and leaf litter, where they prey on small arthropods. But a small proportion are known inquilines, with social insects such as ants and termites, and also with some burrowing mammals, such as
<italic>
<named-content content-type="taxon-name">Ctenomys</named-content>
</italic>
Blainville. The genus now includes the following species groups and species:
<italic>
<named-content content-type="taxon-name">Operclipygus sulcistrius</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus lucanoides</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus schmidti</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus simplistrius</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus sulcistrius</named-content>
</italic>
Marseul, 1870],
<italic>
<named-content content-type="taxon-name">Operclipygus mirabilis</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus mirabilis</named-content>
</italic>
(Wenzel & Dybas, 1941) comb. n.,
<italic>
<named-content content-type="taxon-name">Operclipygus pustulifer</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus plaumanni</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus sinuatus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus mutuca</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus carinistrius</named-content>
</italic>
(Lewis, 1908) comb. n.,
<italic>
<named-content content-type="taxon-name">Operclipygus parensis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus schlingeri</named-content>
</italic>
<bold>sp. n.</bold>
],
<italic>
<named-content content-type="taxon-name">Operclipygus kerga</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus kerga</named-content>
</italic>
(Marseul, 1870),
<italic>
<named-content content-type="taxon-name">Operclipygus planifrons</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus punctistrius</named-content>
</italic>
<bold>sp. n.</bold>
],
<italic>
<named-content content-type="taxon-name">Operclipygus conquisitus</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus bicolor</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus conquisitus</named-content>
</italic>
(Lewis, 1902),
<italic>
<named-content content-type="taxon-name">Operclipygus friburgius</named-content>
</italic>
(Marseul, 1864)],
<italic>
<named-content content-type="taxon-name">Operclipygus impuncticollis</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus bickhardti</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus britannicus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus impuncticollis</named-content>
</italic>
(Hinton, 1935)],
<italic>
<named-content content-type="taxon-name">Operclipygus panamensis</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus crenatus</named-content>
</italic>
(Lewis, 1888),
<italic>
<named-content content-type="taxon-name">Operclipygus panamensis</named-content>
</italic>
(Wenzel & Dybas, 1941)],
<italic>
<named-content content-type="taxon-name">Operclipygus sejunctus</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus depressus</named-content>
</italic>
(Hinton, 1935),
<italic>
<named-content content-type="taxon-name">Operclipygus itoupe</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus juninensis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus pecki</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus punctiventer</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus sejunctus</named-content>
</italic>
(Schmidt, 1896) comb. n.,
<italic>
<named-content content-type="taxon-name">Operclipygus setiventris</named-content>
</italic>
<bold>
<pmc-comment>PageBreak</pmc-comment>
sp. n.</bold>
],
<italic>
<named-content content-type="taxon-name">Operclipygus mortavis</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus ecitonis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus mortavis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus paraguensis</named-content>
</italic>
<bold>sp. n.</bold>
],
<italic>
<named-content content-type="taxon-name">Operclipygus dytiscoides</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus carinisternus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus crenulatus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus dytiscoides</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus quadratus</named-content>
</italic>
<bold>sp. n.</bold>
],
<italic>
<named-content content-type="taxon-name">Operclipygus dubitabilis</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus dubitabilis</named-content>
</italic>
(Marseul, 1889),
<italic>
<named-content content-type="taxon-name">Operclipygus yasuni</named-content>
</italic>
<bold>sp. n.</bold>
],
<italic>
<named-content content-type="taxon-name">Operclipygus angulifer</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus angulifer</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus impressifrons</named-content>
</italic>
<bold>sp. n.</bold>
],
<italic>
<named-content content-type="taxon-name">Operclipygus dubius</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus andinus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus dubius</named-content>
</italic>
(Lewis, 1888),
<italic>
<named-content content-type="taxon-name">Operclipygus extraneus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus intermissus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus lunulus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus occultus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus perplexus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus remotus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus validus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus variabilis</named-content>
</italic>
<bold>sp. n.</bold>
],
<italic>
<named-content content-type="taxon-name">Operclipygus hospes</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus assimilis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus belemensis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus bulbistoma</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus callifrons</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus colombicus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus communis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus confertus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus confluens</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus curtistrius</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus diffluens</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus fusistrius</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus gratus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus hospes</named-content>
</italic>
(Lewis, 1902),
<italic>
<named-content content-type="taxon-name">Operclipygus ibiscus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus ignifer</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus impositus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus incisus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus innocuus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus inquilinus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus minutus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus novateutoniae</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus praecinctus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus prominens</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus rileyi</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus subterraneus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus tenuis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus tiputinus</named-content>
</italic>
<bold>sp. n.</bold>
],
<italic>
<named-content content-type="taxon-name">Operclipygus farctus</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus atlanticus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus bidessois</named-content>
</italic>
(Marseul, 1889),
<italic>
<named-content content-type="taxon-name">Operclipygus distinctus</named-content>
</italic>
(Hinton, 1935),
<italic>
<named-content content-type="taxon-name">Operclipygus distractus</named-content>
</italic>
(Schmidt, 1896) comb. n.,
<italic>
<named-content content-type="taxon-name">Operclipygus farctissimus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus farctus</named-content>
</italic>
(Marseul, 1864),
<italic>
<named-content content-type="taxon-name">Operclipygus gilli</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus impressistrius</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus inflatus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus latemarginatus</named-content>
</italic>
(Bickhardt, 1920) comb. n.,
<italic>
<named-content content-type="taxon-name">Operclipygus petrovi</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus plicatus</named-content>
</italic>
(Hinton, 1935) comb. n.,
<italic>
<named-content content-type="taxon-name">Operclipygus prolixus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus punctifrons</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus proximus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus subrufus</named-content>
</italic>
<bold>sp. n.</bold>
],
<italic>
<named-content content-type="taxon-name">Operclipygus hirsutipes</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus guianensis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus hirsutipes</named-content>
</italic>
<bold>sp. n.</bold>
],
<italic>
<named-content content-type="taxon-name">Operclipygus hamistrius</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus arquus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus campbelli</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus chiapensis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus dybasi</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus geometricus</named-content>
</italic>
(Casey, 1893) comb. n.,
<italic>
<named-content content-type="taxon-name">Operclipygus hamistrius</named-content>
</italic>
(Schmidt, 1893) comb. n.,
<italic>
<named-content content-type="taxon-name">Operclipygus impressicollis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus intersectus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus montanus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus nubosus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus pichinchensis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus propinquus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus quinquestriatus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus rubidus</named-content>
</italic>
(Hinton, 1935) comb. n.,
<italic>
<named-content content-type="taxon-name">Operclipygus rufescens</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus troglodytes</named-content>
</italic>
<bold>sp. n.</bold>
],
<italic>
<named-content content-type="taxon-name">Operclipygus plicicollis</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus cephalicus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus longidens</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus plicicollis</named-content>
</italic>
(Schmidt, 1893)],
<italic>
<named-content content-type="taxon-name">Operclipygus fossipygus</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus disconnectus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus fossipygus</named-content>
</italic>
(Wenzel, 1944),
<italic>
<named-content content-type="taxon-name">Operclipygus foveipygus</named-content>
</italic>
(Bickhardt, 1918),
<italic>
<named-content content-type="taxon-name">Operclipygus fungicolus</named-content>
</italic>
(Wenzel & Dybas, 1941),
<italic>
<named-content content-type="taxon-name">Operclipygus gibbulus</named-content>
</italic>
(Schmidt, 1889) comb. n.,
<italic>
<named-content content-type="taxon-name">Operclipygus olivensis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus simplicipygus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus subdepressus</named-content>
</italic>
(Schmidt, 1889),
<italic>
<named-content content-type="taxon-name">Operclipygus therondi</named-content>
</italic>
(Wenzel, 1976)],
<italic>
<named-content content-type="taxon-name">Operclipygus impunctipennis</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus chamelensis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus foveiventris</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus granulipectus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus impunctipennis</named-content>
</italic>
(Hinton, 1935) comb. n.,
<italic>
<named-content content-type="taxon-name">Operclipygus latifoveatus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus lissipygus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus maesi</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus mangiferus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus marginipennis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus nicodemus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus nitidus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus pacificus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus pauperculus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus punctissipygus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus subviridis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus tripartitus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus vorax</named-content>
</italic>
<bold>sp. n.</bold>
],
<italic>
<named-content content-type="taxon-name">Operclipygus marginellus</named-content>
</italic>
group [
<italic>
<named-content content-type="taxon-name">Operclipygus ashei</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus baylessae</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus dentatus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus formicatus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus hintoni</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus marginellus</named-content>
</italic>
(J.E. LeConte, 1860) comb. n.,
<italic>
<named-content content-type="taxon-name">Operclipygus orchidophilus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus selvorum</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus striatellus</named-content>
</italic>
(Fall, 1917) comb. n.],
<italic>incertae sedis: O. teapensis</italic>
(Marseul, 1853) comb. n.,
<italic>
<named-content content-type="taxon-name">Operclipygus punctulatus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus lama</named-content>
</italic>
Mazur, 1988,
<italic>
<named-content content-type="taxon-name">Operclipygus florifaunensis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus bosquesecus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus arnaudi</named-content>
</italic>
Dégallier, 1982,
<italic>
<named-content content-type="taxon-name">Operclipygus subsphaericus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus latipygus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus elongatus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus rupicolus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus punctipleurus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus falini</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus peregrinus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus brooksi</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus profundipygus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus punctatissimus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus cavisternus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus siluriformis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus parallelus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus abbreviatus</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus pygidialis</named-content>
</italic>
(Lewis, 1908),
<italic>
<named-content content-type="taxon-name">Operclipygus faltistrius</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus limonensis</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus wenzeli</named-content>
</italic>
<bold>sp. n.</bold>
,
<italic>
<named-content content-type="taxon-name">Operclipygus iheringi</named-content>
</italic>
(Bickhardt, 1917),
<italic>
<named-content content-type="taxon-name">Operclipygus angustisternus</named-content>
</italic>
(Wenzel, 1944),
<italic>
<named-content content-type="taxon-name">Operclipygus shorti</named-content>
</italic>
<bold>sp. n.</bold>
We establish the following synonymies:
<italic>
<named-content content-type="taxon-name">Phelisteroides miladae</named-content>
</italic>
Wenzel & Dybas, 1941 and
<italic>
<named-content content-type="taxon-name">Pseudister propygidialis</named-content>
</italic>
Hinton, 1935e =
<italic>
<named-content content-type="taxon-name">Operclipygus crenatus</named-content>
</italic>
(Lewis, 1888);
<italic>
<named-content content-type="taxon-name">Phelister subplicatus</named-content>
</italic>
Schmidt, 1893b =
<italic>
<named-content content-type="taxon-name">Operclipygus bidessois</named-content>
</italic>
(Marseul, 1889). We designate lectotypes for
<italic>
<named-content content-type="taxon-name">Operclipygus sulcistrius</named-content>
</italic>
Marseul, 1870,
<italic>
<named-content content-type="taxon-name">Phelister carinistrius</named-content>
</italic>
Lewis, 1908,
<italic>
<named-content content-type="taxon-name">Phelister kerga</named-content>
</italic>
Marseul, 1870,
<italic>
<named-content content-type="taxon-name">Phelister friburgius</named-content>
</italic>
Marseul, 1864,
<italic>
<named-content content-type="taxon-name">Phelister impuncticollis</named-content>
</italic>
Hinton, 1935,
<italic>
<named-content content-type="taxon-name">Phelister crenatus</named-content>
</italic>
Lewis, 1888,
<italic>
<named-content content-type="taxon-name">Phelister sejunctus</named-content>
</italic>
Schmidt, 1896,
<italic>
<named-content content-type="taxon-name">Pseudister depressus</named-content>
</italic>
Hinton, 1935,
<italic>
<named-content content-type="taxon-name">Epierus dubius</named-content>
</italic>
Lewis, 1888,
<italic>
<named-content content-type="taxon-name">Phelister hospes</named-content>
</italic>
Lewis, 1902,
<italic>
<named-content content-type="taxon-name">Phelister farctus</named-content>
</italic>
Marseul, 1864,
<italic>
<named-content content-type="taxon-name">Phelister bidessois</named-content>
</italic>
Marseul, 1889,
<italic>
<named-content content-type="taxon-name">Phelister subplicatus</named-content>
</italic>
Schmidt, 1893,
<italic>
<named-content content-type="taxon-name">Phelister plicatus</named-content>
</italic>
Hinton, 1935,
<italic>
<named-content content-type="taxon-name">Phelister distinctus</named-content>
</italic>
Hinton, 1935,
<italic>
<named-content content-type="taxon-name">Phelister distractus</named-content>
</italic>
Schmidt, 1896,
<italic>
<named-content content-type="taxon-name">Pseudister latemarginatus</named-content>
</italic>
Bickhardt, 1920,
<italic>
<named-content content-type="taxon-name">Phelister hamistrius</named-content>
</italic>
Schmidt, 1893,
<italic>
<named-content content-type="taxon-name">Phelister plicicollis</named-content>
</italic>
Schmidt, 1893,
<italic>
<named-content content-type="taxon-name">Phelister gibbulus</named-content>
</italic>
Schmidt, 1889,
<italic>
<named-content content-type="taxon-name">Phelister subdepressus</named-content>
</italic>
Schmidt, 1889,
<italic>
<named-content content-type="taxon-name">Phelister teapensis</named-content>
</italic>
Marseul, 1853,
<italic>
<named-content content-type="taxon-name">Phelister pygidialis</named-content>
</italic>
Lewis, 1908,
<italic>
<named-content content-type="taxon-name">Phelister iheringi</named-content>
</italic>
Bickhardt, 1917, and
<italic>
<named-content content-type="taxon-name">Phelister marginellus</named-content>
</italic>
J.E. LeConte 1860. We designate a neotype for
<italic>
<named-content content-type="taxon-name">Operclipygus conquisitus</named-content>
</italic>
Lewis, replacing its lost type specimen.</p>
</abstract>
<kwd-group>
<label>Keywords</label>
<kwd>
<pmc-comment>PageBreak</pmc-comment>
Histeridae</kwd>
<kwd>Histerinae</kwd>
<kwd>Exosternini</kwd>
<kwd>
<italic>Operclipygus</italic>
</kwd>
<kwd>myrmecophily</kwd>
<kwd>Neotropical region</kwd>
</kwd-group>
</article-meta>
</front>
</pmc>
</record>

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