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<record><TEI><teiHeader><fileDesc><titleStmt><title xml:lang="en">The largest type study of <italic>Agaricales</italic> species to date: bringing identification and nomenclature of <italic>Phlegmacium</italic> (<italic>Cortinarius</italic>) into the DNA era</title><author><name sortKey="Liimatainen, K" sort="Liimatainen, K" uniqKey="Liimatainen K" first="K." last="Liimatainen">K. Liimatainen</name><affiliation><nlm:aff id="A1">Department of Biosciences, Plant Biology, P.O. Box 65, FI-00014 University of Helsinki, Finland;</nlm:aff></affiliation></author><author><name sortKey="Niskanen, T" sort="Niskanen, T" uniqKey="Niskanen T" first="T." last="Niskanen">T. Niskanen</name><affiliation><nlm:aff id="A1">Department of Biosciences, Plant Biology, P.O. Box 65, FI-00014 University of Helsinki, Finland;</nlm:aff></affiliation></author><author><name sortKey="Dima, B" sort="Dima, B" uniqKey="Dima B" first="B." last="Dima">B. Dima</name><affiliation><nlm:aff id="A1">Department of Biosciences, Plant Biology, P.O. Box 65, FI-00014 University of Helsinki, Finland;</nlm:aff></affiliation></author><author><name sortKey="Kytovuori, I" sort="Kytovuori, I" uniqKey="Kytovuori I" first="I." last="Kytövuori">I. Kytövuori</name><affiliation><nlm:aff id="A2">Botanical Museum, P.O. Box 7, FI-00014 University of Helsinki, Finland.</nlm:aff></affiliation></author><author><name sortKey="Ammirati, J F" sort="Ammirati, J F" uniqKey="Ammirati J" first="J. F." last="Ammirati">J. F. Ammirati</name><affiliation><nlm:aff id="A3">Department of Biology, Box 351800, University of Washington, Seattle, WA 98195-1800, USA.</nlm:aff></affiliation></author><author><name sortKey="Fr Slev, T G" sort="Fr Slev, T G" uniqKey="Fr Slev T" first="T. G." last="Fr Slev">T. G. Fr Slev</name><affiliation><nlm:aff id="A4">Natural History Museum of Denmark, Center for Geogenetics, University of Copenhagen, Øster Voldgade 5–7, 1350 Københaven K, Denmark.</nlm:aff></affiliation></author></titleStmt><publicationStmt><idno type="wicri:source">PMC</idno><idno type="pmid">25737596</idno><idno type="pmc">4312940</idno><idno type="url">http://www.ncbi.nlm.nih.gov/pmc/articles/PMC4312940</idno><idno type="RBID">PMC:4312940</idno><idno type="doi">10.3767/003158514X684681</idno><date when="2014">2014</date><idno type="wicri:Area/Pmc/Corpus">000239</idno><idno type="wicri:explorRef" wicri:stream="Pmc" wicri:step="Corpus" wicri:corpus="PMC">000239</idno></publicationStmt><sourceDesc><biblStruct><analytic><title xml:lang="en" level="a" type="main">The largest type study of <italic>Agaricales</italic> species to date: bringing identification and nomenclature of <italic>Phlegmacium</italic> (<italic>Cortinarius</italic>) into the DNA era</title><author><name sortKey="Liimatainen, K" sort="Liimatainen, K" uniqKey="Liimatainen K" first="K." last="Liimatainen">K. Liimatainen</name><affiliation><nlm:aff id="A1">Department of Biosciences, Plant Biology, P.O. Box 65, FI-00014 University of Helsinki, Finland;</nlm:aff></affiliation></author><author><name sortKey="Niskanen, T" sort="Niskanen, T" uniqKey="Niskanen T" first="T." last="Niskanen">T. Niskanen</name><affiliation><nlm:aff id="A1">Department of Biosciences, Plant Biology, P.O. Box 65, FI-00014 University of Helsinki, Finland;</nlm:aff></affiliation></author><author><name sortKey="Dima, B" sort="Dima, B" uniqKey="Dima B" first="B." last="Dima">B. Dima</name><affiliation><nlm:aff id="A1">Department of Biosciences, Plant Biology, P.O. Box 65, FI-00014 University of Helsinki, Finland;</nlm:aff></affiliation></author><author><name sortKey="Kytovuori, I" sort="Kytovuori, I" uniqKey="Kytovuori I" first="I." last="Kytövuori">I. Kytövuori</name><affiliation><nlm:aff id="A2">Botanical Museum, P.O. Box 7, FI-00014 University of Helsinki, Finland.</nlm:aff></affiliation></author><author><name sortKey="Ammirati, J F" sort="Ammirati, J F" uniqKey="Ammirati J" first="J. F." last="Ammirati">J. F. Ammirati</name><affiliation><nlm:aff id="A3">Department of Biology, Box 351800, University of Washington, Seattle, WA 98195-1800, USA.</nlm:aff></affiliation></author><author><name sortKey="Fr Slev, T G" sort="Fr Slev, T G" uniqKey="Fr Slev T" first="T. G." last="Fr Slev">T. G. Fr Slev</name><affiliation><nlm:aff id="A4">Natural History Museum of Denmark, Center for Geogenetics, University of Copenhagen, Øster Voldgade 5–7, 1350 Københaven K, Denmark.</nlm:aff></affiliation></author></analytic><series><title level="j">Persoonia : Molecular Phylogeny and Evolution of Fungi</title><idno type="ISSN">0031-5850</idno><idno type="eISSN">1878-9080</idno><imprint><date when="2014">2014</date></imprint></series></biblStruct></sourceDesc></fileDesc><profileDesc><textClass></textClass></profileDesc></teiHeader><front><div type="abstract" xml:lang="en"><p><italic>Cortinarius</italic> is a species-rich and morphologically challenging genus with a cosmopolitan distribution. Many names have not been used consistently and in some instances the same species has been described two or more times under separate names. This study focuses on subg. <italic>Phlegmacium</italic> as traditionally defined and includes species from boreal and temperate areas of the northern hemisphere. Our goals for this project were to: i) study type material to determine which species already have been described; ii) stabilize the use of Friesian and other older names by choosing a neo- or epitype; iii) describe new species that were discovered during the process of studying specimens; and iv) establish an accurate ITS barcoding database for <italic>Phlegmacium</italic> species. A total of 236 types representing 154 species were studied. Of these 114 species are described only once whereas 40 species had one ore more synonyms. Of the names studied only 61 were currently represented in GenBank. Neotypes are proposed for 21 species, and epitypes are designated for three species. In addition, 20 new species are described and six new combinations made. As a consequence ITS barcodes for 175 <italic>Cortinarius</italic> species are released.</p></div></front><back><div1 type="bibliography"><listBibl><biblStruct><analytic><author><name sortKey="Abarenkov, K" uniqKey="Abarenkov K">K Abarenkov</name></author><author><name sortKey="Nilsson, Rh" uniqKey="Nilsson R">RH Nilsson</name></author><author><name sortKey="Larsson, K H" uniqKey="Larsson K">K-H Larsson</name></author></analytic></biblStruct><biblStruct><analytic><author><name sortKey="Ammirati, Jf" uniqKey="Ammirati J">JF Ammirati</name></author></analytic></biblStruct><biblStruct><analytic><author><name sortKey="Ammirati, Jf" uniqKey="Ammirati J">JF Ammirati</name></author><author><name sortKey="Hughes, Kw" uniqKey="Hughes K">KW Hughes</name></author><author><name sortKey="Liimatainen, K" uniqKey="Liimatainen K">K Liimatainen</name></author></analytic></biblStruct><biblStruct><analytic><author><name sortKey="Bidaud, A" uniqKey="Bidaud A">A Bidaud</name></author><author><name 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<front><journal-meta><journal-id journal-id-type="nlm-ta">Persoonia</journal-id><journal-id journal-id-type="iso-abbrev">Persoonia</journal-id><journal-id journal-id-type="publisher-id">Persoonia</journal-id><journal-title-group><journal-title>Persoonia : Molecular Phylogeny and Evolution of Fungi</journal-title></journal-title-group><issn pub-type="ppub">0031-5850</issn><issn pub-type="epub">1878-9080</issn><publisher><publisher-name>Naturalis Biodiversity Center & Centraallbureau voor Schimmelcultures</publisher-name></publisher></journal-meta><article-meta><article-id pub-id-type="pmid">25737596</article-id><article-id pub-id-type="pmc">4312940</article-id><article-id pub-id-type="doi">10.3767/003158514X684681</article-id><article-categories><subj-group subj-group-type="heading"><subject>Research Article</subject></subj-group></article-categories><title-group><article-title>The largest type study of <italic>Agaricales</italic> species to date: bringing identification and nomenclature of <italic>Phlegmacium</italic> (<italic>Cortinarius</italic>) into the DNA era</article-title></title-group><contrib-group><contrib contrib-type="author"><name><surname>Liimatainen</surname><given-names>K.</given-names></name><xref ref-type="aff" rid="A1"><sup>1</sup></xref><xref ref-type="corresp" rid="COR1"></xref></contrib><contrib contrib-type="author"><name><surname>Niskanen</surname><given-names>T.</given-names></name><xref ref-type="aff" rid="A1"><sup>1</sup></xref></contrib><contrib contrib-type="author"><name><surname>Dima</surname><given-names>B.</given-names></name><xref ref-type="aff" rid="A1"><sup>1</sup></xref></contrib><contrib contrib-type="author"><name><surname>Kytövuori</surname><given-names>I.</given-names></name><xref ref-type="aff" rid="A2"><sup>2</sup></xref></contrib><contrib contrib-type="author"><name><surname>Ammirati</surname><given-names>J.F.</given-names></name><xref ref-type="aff" rid="A3"><sup>3</sup></xref></contrib><contrib contrib-type="author"><name><surname>Frøslev</surname><given-names>T.G.</given-names></name><xref ref-type="aff" rid="A4"><sup>4</sup></xref></contrib></contrib-group><aff id="A1"><label>1</label>Department of Biosciences, Plant Biology, P.O. Box 65, FI-00014 University of Helsinki, Finland;</aff><aff id="A2"><label>2</label>Botanical Museum, P.O. Box 7, FI-00014 University of Helsinki, Finland.</aff><aff id="A3"><label>3</label>Department of Biology, Box 351800, University of Washington, Seattle, WA 98195-1800, USA.</aff><aff id="A4"><label>4</label>Natural History Museum of Denmark, Center for Geogenetics, University of Copenhagen, Øster Voldgade 5–7, 1350 Københaven K, Denmark.</aff><author-notes><corresp id="COR1">corresponding author e-mail: <email>kare.liimatainen@helsinki.fi.</email>.</corresp></author-notes><pub-date pub-type="epub"><day>8</day><month>9</month><year>2014</year></pub-date><pub-date pub-type="ppub"><month>12</month><year>2014</year></pub-date><volume>33</volume><fpage>98</fpage><lpage>140</lpage><history><date date-type="received"><day>2</day><month>7</month><year>2013</year></date><date date-type="accepted"><day>24</day><month>2</month><year>2014</year></date></history><permissions><copyright-statement>© 2014 Naturalis Biodiversity Center & Centraalbureau voor Schimmelcultures</copyright-statement><copyright-year>2014</copyright-year><license license-type="open-access" xlink:href="http://creativecommons.org/licenses/by-nc-nd/3.0/legalcode"><license-p>You are free to share - to copy, distribute and transmit the work, under the following conditions:</license-p><license-p>Attribution: You must attribute the work in the manner specified by the author or licensor (but not in any way that suggests that they endorse you or your use of the work).</license-p><license-p>Non-commercial: You may not use this work for commercial purposes.</license-p><license-p>No derivative works: You may not alter, transform, or build upon this work.</license-p><license-p>For any reuse or distribution, you must make clear to others the license terms of this work, which can be found at <ext-link ext-link-type="uri" xlink:href="http://creativecommons.org/licenses/by-nc-nd/3.0/legalcode">http://creativecommons.org/licenses/by-nc-nd/3.0/legalcode</ext-link>. Any of the above conditions can be waived if you get permission from the copyright holder. Nothing in this license impairs or restricts the author’s moral rights.</license-p></license></permissions><abstract abstract-type="executive-summary"><p><italic>Cortinarius</italic> is a species-rich and morphologically challenging genus with a cosmopolitan distribution. Many names have not been used consistently and in some instances the same species has been described two or more times under separate names. This study focuses on subg. <italic>Phlegmacium</italic> as traditionally defined and includes species from boreal and temperate areas of the northern hemisphere. Our goals for this project were to: i) study type material to determine which species already have been described; ii) stabilize the use of Friesian and other older names by choosing a neo- or epitype; iii) describe new species that were discovered during the process of studying specimens; and iv) establish an accurate ITS barcoding database for <italic>Phlegmacium</italic> species. A total of 236 types representing 154 species were studied. Of these 114 species are described only once whereas 40 species had one ore more synonyms. Of the names studied only 61 were currently represented in GenBank. Neotypes are proposed for 21 species, and epitypes are designated for three species. In addition, 20 new species are described and six new combinations made. As a consequence ITS barcodes for 175 <italic>Cortinarius</italic> species are released.</p></abstract><kwd-group><kwd>Basidiomycota</kwd><kwd>diversity</kwd><kwd>DNA barcoding</kwd><kwd>ITS</kwd><kwd>taxonomy</kwd><kwd>typification</kwd></kwd-group></article-meta></front><body><sec id="s1"><title>INTRODUCTION</title><p>Studies using DNA sequence data have shown that the diversity of fungi far exceeds earlier expectations with many common species of macrofungi still to be described and named (<xref rid="R85" ref-type="bibr">Schmit & Mueller 2007</xref>). In addition, it is not always easy to determine which species have been described and which are new to science, unless the type specimens of existing names can be carefully studied, including DNA sequencing.</p><p>For many species of macrofungi the names have been difficult or even impossible to interpret. The most difficult ones are the older names with brief descriptions and usually without type material. But even if type material exists it can be very difficult to be certain of the identification based only on morphology, especially in challenging genera like <italic>Cortinarius</italic> where there is considerable convergence in morphology, colouration, and microscopic features. Furthermore, the literature is often difficult to obtain, making it hard to get information on available names and their application by earlier taxonomists. Consequently, many names have not been used consistently and in some cases the same species has been described two or more times under separate names. In instances where there is no type material available, a neotype (or a lectotype if collections of the author are available) is required to stabilize the use of the name. Finally, old type collections that are considered historical materials may not be available for study or DNA sequencing requiring the selection of an epitype.</p><p>The development of molecular techniques has provided a more unambiguous tool to identify species. Currently the most commonly used locus in species level taxonomy is the nuclear ribosomal internal transcribed spacer (ITS), which has been proposed as a universal barcode marker for fungi (<xref rid="R86" ref-type="bibr">Schoch et al. 2012</xref>). The region is present in several chromosomes and is arranged in tandem repeats that are thousands of copies long (<xref rid="R22" ref-type="bibr">Burnett 2003</xref>). Due to the high copy number the region usually is easy to amplify and sequence, even from very old specimens (<xref rid="R57" ref-type="bibr">Larsson & Jacobsson 2004</xref>). In <italic>Cortinarius</italic> the ITS was proposed as a species-identifier sequence already in 2007 by Frøslev et al. and in 2008 by Ortega et al. It has also been shown that in the majority of the cases ITS is suitable for species delimitation in <italic>Cortinarius</italic>. The results of the multi-gene phylogenetic study based on ITS, <italic>rpb1</italic>, and <italic>rpb2</italic> regions by <xref rid="R33" ref-type="bibr">Frøslev et al. (2005)</xref> showed that inference from ITS alone is indicative of the species level phylogenetic delimitations of multi-gene analyses. Furthermore, the delimitations inferred from ITS usually correlate with the morphospecies (<xref rid="R32" ref-type="bibr">Frøslev et al. 2007</xref>, <xref rid="R78" ref-type="bibr">Ortega et al. 2008</xref>, <xref rid="R73" ref-type="bibr">Niskanen et al. 2012b</xref>).</p><p><italic>Cortinarius </italic>is the largest genus of <italic>Agaricales</italic> with a cosmopolitan distribution and over 2 000 described species (<xref rid="R56" ref-type="bibr">Kirk et al. 2008</xref>). <italic>Cortinarius</italic> species are important ectomycorrhizal fungi associated with different trees and shrubs, belonging to the order <italic>Fagales</italic>, families <italic>Caesalpiniaceae</italic>,<italic> Cistaceae</italic>, <italic>Dipterocarpaceae</italic>, <italic>Myrtaceae</italic>, <italic>Pinaceae</italic>, <italic>Rhamnaceae</italic>, <italic>Rosaceae</italic> and <italic>Salicaceae</italic> as well as a few herbaceous plants in the <italic>Cyperaceae</italic>. Owing to their often narrow ecological preferences and sensitivity to environmental change, many <italic>Cortinarius</italic> species have been used as indicator species for valuable natural environments, e.g. in Sweden and Denmark (<xref rid="R92" ref-type="bibr">Vesterholt 1991</xref>, <xref rid="R42" ref-type="bibr">Hallingbäck & Aronsson 1998</xref>, <xref rid="R30" ref-type="bibr">Frøslev & Jeppesen 2011</xref>). Lately it also was suggested that they have a key role in the carbon cycling of boreal forests (<xref rid="R11" ref-type="bibr">Bödeker et al. 2011</xref>).</p><p>Many of the major studies in <italic>Cortinarius</italic> have dealt with North American and especially European species, while the species of southern hemisphere are somewhat less studied (<xref rid="R66" ref-type="bibr">Moser & Horak 1975</xref>, <xref rid="R36" ref-type="bibr">Garnica et al. 2002</xref>, <xref rid="R23" ref-type="bibr">Cleland 1976</xref> (1935), <xref rid="R39" ref-type="bibr">Gasparini & Soop 2008</xref>). In Europe the most extensive studies have been done by <xref rid="R26" ref-type="bibr">Fries (e.g., 1821</xref>, <xref rid="R27" ref-type="bibr">1836–1838</xref>, <xref rid="R28" ref-type="bibr">1851</xref>) from Sweden, <xref rid="R46" ref-type="bibr">Henry (e.g., 1951</xref>, <xref rid="R47" ref-type="bibr">1958</xref>, <xref rid="R48" ref-type="bibr">1981</xref>, <xref rid="R50" ref-type="bibr">1986</xref>) and <xref rid="R5" ref-type="bibr">Bidaud et al. (e.g., 1992</xref>, <xref rid="R9" ref-type="bibr">2010</xref>) from France, <xref rid="R60" ref-type="bibr">Moser (e.g. 1960</xref>, <xref rid="R61" ref-type="bibr">1969–1970</xref>, <xref rid="R62" ref-type="bibr">1983</xref>) from Central Europe, <xref rid="R51" ref-type="bibr">Høiland (1984)</xref>, <xref rid="R14" ref-type="bibr">Brandrud (e.g., 1996</xref>, <xref rid="R15" ref-type="bibr">1998</xref>), <xref rid="R16" ref-type="bibr">Brandrud et al. (e.g., 1990</xref>, <xref rid="R20" ref-type="bibr">2012</xref>) and <xref rid="R70" ref-type="bibr">Niskanen et al. (e.g., 2009</xref>, <xref rid="R71" ref-type="bibr">2011a</xref>, <xref rid="R72" ref-type="bibr">2013a</xref>) mainly from northern Europe, <xref rid="R31" ref-type="bibr">Frøslev et al. (e.g., 2006</xref>, <xref rid="R32" ref-type="bibr">2007</xref>) from northern and Central Europe, <xref rid="R79" ref-type="bibr">Orton (1955</xref>, <xref rid="R80" ref-type="bibr">1958</xref>) from Great Britain, and <xref rid="R78" ref-type="bibr">Ortega et al. (2008)</xref> and <xref rid="R90" ref-type="bibr">Suárez-Santiago et al. (2009)</xref> from mediterranean area. Selected papers of some of the major contributors to <italic>Cortinarius</italic> systematics in North America include <xref rid="R81" ref-type="bibr">Peck (1873</xref>; also see <xref rid="R40" ref-type="bibr">Gilbertson 1962</xref>), <xref rid="R53" ref-type="bibr">Kauffman (1918</xref>, <xref rid="R54" ref-type="bibr">1923</xref>, <xref rid="R55" ref-type="bibr">1932</xref>), <xref rid="R87" ref-type="bibr">Smith (1939</xref>, <xref rid="R88" ref-type="bibr">1942</xref>, <xref rid="R89" ref-type="bibr">1944</xref>), <xref rid="R2" ref-type="bibr">Ammirati (1972)</xref>, <xref rid="R3" ref-type="bibr">Ammirati et al. (2013)</xref>, <xref rid="R37" ref-type="bibr">Garnica et al. (2009)</xref>, <xref rid="R58" ref-type="bibr">Liu et al. (1997)</xref>, <xref rid="R63" ref-type="bibr">Moser & Ammirati (1996</xref>, <xref rid="R64" ref-type="bibr">1999</xref>), <xref rid="R67" ref-type="bibr">Moser et al. (1995)</xref>, <xref rid="R12" ref-type="bibr">Bojantchev (2011a</xref>, <xref rid="R13" ref-type="bibr">b</xref>) and <xref rid="R74" ref-type="bibr">Niskanen et al. (2013b</xref>, <xref rid="R75" ref-type="bibr">c</xref>).</p><p>Very little is known about the distribution of <italic>Cortinarius </italic>species on a larger scale or the differences in the species composition between the continents, although species in the southern hemisphere are distinct from those in the northern hemisphere (<xref rid="R82" ref-type="bibr">Peintner et al. 2004</xref>, <xref rid="R38" ref-type="bibr">Garnica et al. 2005</xref>). However, recent molecular studies on <italic>Cortinarius</italic> have shed some light on these questions for North America and Europe (<xref rid="R68" ref-type="bibr">Moser & Peintner 2002</xref>, <xref rid="R59" ref-type="bibr">Matheny & Ammirati 2006</xref>, <xref rid="R37" ref-type="bibr">Garnica et al. 2009</xref>, <xref rid="R35" ref-type="bibr">2011</xref>, <xref rid="R43" ref-type="bibr">Harrower et al. 2011</xref>, <xref rid="R3" ref-type="bibr">Ammirati et al. 2013</xref>, <xref rid="R76" ref-type="bibr">Niskanen et al. 2011b</xref>, <xref rid="R77" ref-type="bibr">2012c</xref>, <xref rid="R74" ref-type="bibr">2013b</xref>, <xref rid="R75" ref-type="bibr">c</xref>). These studies show several patterns of species distributions. There are species common to North America and Europe, especially those species from more northern and montane conifer forests, i.e. <italic>Cortinarius aureofulvus</italic> M.M. Moser s.auct., <italic>C. napus</italic> Fr. and <italic>C. pinophilus</italic> Soop, but also endemic species occur both in western North America, eastern North America and Europe, i.e. <italic>C. elegantio-occidentalis</italic>Garnica & Ammirati in western North America, <italic>C. hesleri </italic>Ammirati, Niskanen, Liimat. & Matheny in eastern North America and <italic>C. puniceus </italic>P.D. Orton in Europe. <italic>Cortinarius </italic>species composition is somewhat similar between eastern North America and Europe, but there appears to be less similarity between Europe and western North America.</p><p>Different infrageneric classification systems have been proposed for <italic>Cortinarius</italic>, i.e. <xref rid="R16" ref-type="bibr">Brandrud et al. (1990)</xref> divided the genus in four subgenera: <italic>Cortinarius</italic>,<italic> Myxacium</italic>,<italic> Phlegmacium</italic> and <italic>Telamonia</italic>. Recent molecular analyses have shown that <italic>Cortinarius</italic> is monophyletic, but also that many of the infrageneric groups such as subg. <italic>Phlegmacium</italic> are not monophyletic (<xref rid="R82" ref-type="bibr">Peintner et al. 2004</xref>, <xref rid="R38" ref-type="bibr">Garnica et al. 2005</xref>). Nonetheless, several authors have treated subg. <italic>Phlegmacium</italic> in similar ways (<xref rid="R62" ref-type="bibr">Moser 1983</xref>, <xref rid="R6" ref-type="bibr">Bidaud et al. 1994</xref>, <xref rid="R20" ref-type="bibr">Brandrud et al. 2012</xref>) including in it species with viscid to glutinous pileus and dry, often bulbous stipe, or dry-capped, stout species with a yellow KOH reaction, i.e. the species of sect. <italic>Phlegmacioides</italic> (<xref rid="R69" ref-type="bibr">Niskanen et al. 2012a</xref>).</p><p>Molecular techniques have been in use for more than a decade, and the sequencing of ITS regions even from older <italic>Cortinarius</italic> specimens is possible. Furthermore, type studies are essential for a stable and consistent application of names in <italic>Cortinarius</italic> where currently a large percentage of the <italic>Cortinarius </italic>sequences are incorrectly named or without a name in the public sequence databases (e.g., <xref rid="R71" ref-type="bibr">Niskanen et al. 2011a</xref>). While several papers present sequences for individuals or groups of species, for example <xref rid="R37" ref-type="bibr">Garnica et al. (2009)</xref> and <xref rid="R71" ref-type="bibr">Niskanen et al. (2011a)</xref>, the only large study is that of <xref rid="R32" ref-type="bibr">Frøslev et al. (2007)</xref> where 52 types of <italic>Cortinarius</italic> sect. <italic>Calochroi</italic> were sequenced. The present study focuses on <italic>Cortinarius</italic> subg. <italic>Phlegmacium</italic> as traditionally defined and includes species from boreal and temperate areas of the northern hemisphere. Our goals for this project are to: i) study type material to determine which taxa have already been described; ii) stabilize the use of Friesian and other older names by choosing a neo- or epitype; iii) describe new species that were discovered during the process of studying specimens; and iv) establish an accurate ITS barcoding database for <italic>Phlegmacium</italic> species.</p></sec><sec sec-type="materials|methods" id="s2"><title>MATERIALS AND METHODS</title><sec id="s2a"><title>Taxon sampling</title><p>The type specimens of species of subg. <italic>Phlegmacium</italic> published over many years by J.F. Ammirati, A. Bidaud, T.E. Brandrud, G. Chevassut, J. Favre, R. Henry, P.A. Karsten, C.H. Kauffman, R. Kühner, K.H. McKnight, M.M. Moser, P. Moënne-Loccoz, P. Reumaux, A.H. Smith and H.D. Thiers were sampled as well as relevant collections published and illustrated in <xref rid="R16" ref-type="bibr">Brandrud et al. (1990</xref>, <xref rid="R17" ref-type="bibr">1992</xref>, <xref rid="R18" ref-type="bibr">1994</xref>, <xref rid="R19" ref-type="bibr">1998</xref>). The species of sections <italic>Calochroi</italic>, <italic>Fulvi</italic> and <italic>Riederi </italic>described from Europe were generally excluded. The first two have been treated by <xref rid="R32" ref-type="bibr">Frøslev et al. (e.g., 2007</xref>) and the latter will be treated by Brandrud et al. in the near future. In some instances type material could not be acquired from herbaria, for example, the <italic>Cortinarius</italic> type collections of C.H. Peck were only recently available on loan from the New York State Museum and will be included in a later study.</p><p>Our aim was to have at least two sequences per species in our study. Therefore, in addition to sequences from type specimens, either our own unpublished sequences or additional sequences retrieved from the sequence databases GenBank and UNITE were included. Unpublished sequences were also supplied by D. Bojantchev, K. Hughes and A. Taylor. Information on the sequences of type specimens is available in <xref ref-type="table" rid="T1">Table 1</xref> and information on other sequences included in the phylogenetic analysis is available in <xref ref-type="table" rid="T2">Table 2</xref>. Herbarium acronyms follow Index Herbariorum (<xref rid="R91" ref-type="bibr">Thiers 2013</xref>).</p></sec><sec id="s2b"><title>Molecular analyses</title><p>DNA was extracted from a few milligrams of dried material (a piece of lamella) with the NucleoSpin Plant kit (Macherey-Nagel, Düren, Germany), or with various CTAB protocols in Brandrud’s and Frøslev’s specimens (see <xref rid="R33" ref-type="bibr">Frøslev et al. 2005</xref>, <xref rid="R32" ref-type="bibr">2007</xref>). Primers ITS 1F and ITS 4 (<xref rid="R93" ref-type="bibr">White et al. 1990</xref>, <xref rid="R34" ref-type="bibr">Gardes & Bruns 1993</xref>) were used to amplify ITS regions. The same primer pairs were used in direct sequencing. For problematic material the primer combinations ITS 1F/ITS 2 and ITS 3/ITS 4 were also used. PCR amplifications were performed in a 25 μL reaction mix with about 70 ng extracted DNA, 1 U Phusion High-Fidelity DNA polymerase and 1× HF buffer (Finnzymes), 200 mM of each dNTP and 0.5 μM of each primer. The PCR reactions were run on a MBS 0.2 G Thermal Cycler (Thermo Hybaid) with the following settings: denaturation for 30 s at 98 °C, followed by 35 cycles of denaturation for 10 s at 98 °C, annealing for 30 s at 50 °C and extension for 30 s at 72 °C. The PCR products were purified using an ExoSAP-IT purification kit (Amersham Biosciences). Sequencing was performed on both strands using a BigDye Terminator v. 1.1 Sequencing kit (Applied Biosystems). Reactions were performed in 10 μL with 1 μL of PCR product, 1.3 mM of primer (ITS 1F or ITS 4), 1 μL 5X sequencing buffer, and 1 μL of Terminator Ready Reaction Mix. Reactions were run for 1 min at 96 °C, followed by 30 cycles of 30 s at 96 °C, 15 s at 50 °C and 4 min at 60 °C. Unincorporated dye terminators and primers were removed by Sephadex G-50 DNA Grade Fine (Amersham Biosciences) purification system, and the reactions were analysed by ABI 3730 DNA Analyzer (Applied Biosystems) automatic sequencer. Sequences were assembled and edited with Sequencer 4.1 (Gene Codes, Ann Arbor, Michigan, USA). A total of 405 new ITS sequences were produced for this study. Collections and GenBank sequences used for the phylogenetic analysis are given in <xref ref-type="table" rid="T1">Table 1</xref> and <xref ref-type="table" rid="T2">2</xref>. The alignment of 461 ITS sequences was produced with the MUSCLE program (<xref rid="R24" ref-type="bibr">Edgar 2004</xref>) under default settings and followed by manual adjustments in BioEdit (<ext-link ext-link-type="uri" xlink:href="http://www.mbio.ncsu.edu/BioEdit/bioedit.html">www.mbio.ncsu.edu/BioEdit/bioedit.html</ext-link>). The alignment is 812 nucleotides long (including gaps) and is available at TreeBASE under S14832 (<ext-link ext-link-type="uri" xlink:href="http://www.treebase.org/treebase-web/home.html">http://www.treebase.org/treebase-web/home.html</ext-link>).</p><p>Bayesian inference (BI) was performed with MrBayes v. 3.1.1 (<xref rid="R84" ref-type="bibr">Ronquist & Huelsenbeck 2003</xref>). The best substitution model for alignment was estimated by both the Akaike information criterion and the Bayesian information criterion with jModelTest 0.1.1 (<xref rid="R83" ref-type="bibr">Posada 2008</xref>). GTR model was chosen. Two independent runs with four chains in each were performed 6 000 000 generations with sampling every 100th generation. All trees sampled before stationarity were discarded with a 25 % safety margin (burn-in of 15 000 trees, 1 500 000 generations). Sampled trees from both runs were combined in a 50 % majority rule consensus phylogram with posterior probabilities (PP). The analyses were run with computer clusters of the CSC, IT Center for Science, Espoo, Finland.</p></sec><sec id="s2c"><title>Morphological study</title><p>Morphological descriptions are based on material collected by the authors and D. Bojantchev, T.E. Brandrud, T.S. Jeppesen, E. Sesli and A.F. Taylor including specimens in all stages of development. Microscopic characteristics were observed from dried material mounted in Melzer’s reagent (MLZ). Measurements were made in MLZ with an ocular micrometer using 100× oil-immersion lens. Basidiospores were measured from the veil or top of the stipe, 20 spores from one basidiocarp unless otherwise indicated. The length and width were measured for each spore, and their length/width ratios (Q value) were calculated. The average of all measurements is marked in <italic>italic</italic>. The lamellar trama and basidia also were examined and the pileipellis structure was studied from scalps from the pileus centre.</p></sec></sec><sec id="s3"><title>RESULTS</title><sec id="s3a"><title>Phylogenetic analyses</title><p>The 50 % majority rule phylogram resulting from the BI analysis is shown in <xref ref-type="fig" rid="F1">Fig. 1</xref>. Most species are supported by 1.00 PP (or slightly less). Six species received support below 0.95 PP: <italic>C. castaneicolor</italic>, <italic>C. collocandoides</italic>, <italic>C. flavescentipes </italic>(=<italic> C. balteatocumatilis </italic>s.auct.), <italic>C. herpeticus</italic>, <italic>C. infractiflavus</italic> and <italic>C. multiformis</italic>. Three species, <italic>C. balteatoalutaceus</italic>, <italic>C. pseudocephalixus</italic> and <italic>C. subfoetens</italic> did not form monophyletic groups in our phylogenetic analysis. Most of these nine species represent species that differ from their closest relatives by less than 10 substitutions and indel positions in ITS regions, but they all, however, have low intraspecific variation.</p><p>In a majority of the species, the intraspecific variation is from 0 to 1 substitutions or indel positions. In some species it is 2 substitutions and/or indel positions. Similarly, in a majority of the species, the interspecific difference is more than 10 substitutions and indel positions, but in some species it is only four, i.e. in the species pair <italic>C.claricolor</italic>/<italic>C. rex-claricolorum</italic>. All the species, however, have a clear barcoding gap between intra- and interspecific variation. In the following species the intraspecific variation was more than two substitutions and/or indel positions, but no clear grouping was obtained in the analysis of ITS sequences or the number of specimens was too low for making conclusions:<italic> C. albescens</italic>/<italic>C. gentianeus</italic>/<italic>C. volvatus</italic> complex, <italic>C. aureofulvus </italic>s.auct./<italic>C. orichalceus </italic>var.<italic> xanthocephalus</italic> complex,<italic> C. herpeticus</italic>/<italic>C. violaceonitens</italic> complex, <italic>C. misermontii</italic>/<italic>C. olidoamarus </italic>var.<italic> valentinus</italic>/<italic>C. subaccedens</italic>/<italic>C. van-campiae</italic> complex, <italic>C. pallidirimosus</italic>, <italic>C. porphyropus</italic> and <italic>C. scaurus</italic>.</p><p>A total of 236 types representing 154 species were successfully sequenced. Of these, 114 species are described only once whereas 40 species had one or more synonyms. The species with the largest number of synonyms are <italic>C.largus</italic> (14 synonyms), <italic>C. pseudonaevosus</italic> (6 synonyms), <italic>C. talus</italic> (5 synonyms), <italic>C. crassus</italic> (4 synonyms) and <italic>C.varius</italic> (4 synonyms). All the names of the types are listed in alphabetical order in <xref ref-type="table" rid="T1">Table 1</xref>, followed by the current name.</p><p>Several infrageneric groups with three or more species were supported by 1.00 PP: /Calochroi & Fulvi, /Claricolores, /Cyanites, /Dionysae (s.lat. 0.56 PP), /Glaucopodes, /Infracti, /Multiformes, /Phlegmacium, /Purpurascentes and /Scauri. In addition, clade /Arguti (0.98 PP) received some support and clades /Elastici & Percomes (0.94 PP, s.l. 0.53 PP), /Phlegmacioides (0.77 PP) and /Variosimiles (0.74 PP) low support.</p></sec><sec id="s3b"><title>Taxonomy</title><p>Neo- and epitypifications</p><p>Neotypes for 15 species described by Fries and six species described by Albertini, Batsch, Britzelmayr, Persoon, Schaeffer and Schweinitz are proposed as well as epitypes for three species described by Batsch, Cordier and Schaeffer in the 19th century. These include names that have been commonly used in Europe during the last 20 years (<xref rid="R16" ref-type="bibr">Brandrud et al. 1990</xref>, <xref rid="R17" ref-type="bibr">1992</xref>, <xref rid="R18" ref-type="bibr">1994</xref>, <xref rid="R19" ref-type="bibr">1998</xref>, <xref rid="R52" ref-type="bibr">Jeppesen et al. 2012</xref>) and where the current use of the names is not in contradiction with the protologue. Citations of illustrations and descriptions of the species are provided. If our observations on the species deviate from those of <xref rid="R16" ref-type="bibr">Brandrud et al. (1990</xref>, <xref rid="R17" ref-type="bibr">1992</xref>, <xref rid="R18" ref-type="bibr">1994</xref>, <xref rid="R19" ref-type="bibr">1998</xref>) and/or <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012)</xref>, they are presented in comments under each species. For <italic>C. cyanites </italic>and <italic>C. fraudulosus</italic> full descriptions are provided since the current use of the name included several closely related species. Synonyms are based on DNA studies of the type specimens and the information on the types is presented in <xref ref-type="table" rid="T1">Table 1</xref>. The reasonings for synonymy are presented in the discussion.</p><p><bold><italic>Cortinarius balteatus</italic></bold> (Fr.) Fr., Epicr. Syst. Mycol.: 257. 1838</p><p><italic>Basionym</italic>. <italic>Agaricus balteatus</italic> Fr., Observ. Mycol. 2: 138. 1818.</p><p>=<italic> Cortinarius subbalteatus</italic> Kühner, Bull. Mens. Soc. Linn. Lyon 24, 2: 40. 1955.</p><p>=<italic> Cortinarius balteatotomentosus</italic> Rob. Henry ex Rob. Henry, Bull. Soc. Mycol. France 101, 1: 4. 1985.</p><p>=<italic> Cortinarius subopimus</italic> Bidaud, Atlas des Cortinaires 7: 231. 1995.</p><p><italic>Neotype</italic>. S<sc>WEDEN</sc>, Ångermanland, Säbrå, Överdal, under <italic>Picea</italic> on cultivated area, 3 Aug. 1990, <italic>Brandrud et al.</italic> CFP940 (S), designated here. MycoBank MBT176298. GenBank KF732262.</p><p>Illustrations — <xref rid="R18" ref-type="bibr">Brandrud et al. (1994</xref>: pl. C60), <xref rid="R29" ref-type="bibr">Fries (1867–1884</xref>: pl. 142).</p><p>Descriptions of the species — <xref rid="R18" ref-type="bibr">Brandrud et al. (1994</xref>: pl. C60), <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012</xref>: 814).</p><p>Notes — <italic>Cortinarius balteatotomentosus</italic> was first described in 1958 without indicating a type, nonetheless Index Fungorum and MycoBank report it as valid. The validation has been performed later by <xref rid="R49" ref-type="bibr">Henry (1985)</xref> giving the holotype no. 306.</p><p><bold><italic>Cortinarius caerulescens</italic></bold> (Schaeff.) Fr., Epicr. Syst. Mycol.: 265. 1838</p><p><italic>Basionym</italic>. <italic>Agaricus caerulescens</italic> Schaeff., Fung. Bavar. Palat. 4: 17. 1774.</p><p><italic>Lectotype</italic>. J.C. Schaeffer, Fung. Bav. I, t. 34, f. I, II, III (1762) (designated in Cortin. Fl. Photogr. II (Swedish version): 11, 1992).</p><p><italic>Epitype</italic>. B<sc>ELGIUM</sc>, Brabant, Tervuren, in beech forest on calcareous soil, 23 Sept. 1989, <italic>Brandrud et al.</italic> CFP853 (S), designated here. MycoBank MBT176395. GenBank KF732271.</p><p>Illustrations — <xref rid="R17" ref-type="bibr">Brandrud et al. (1992</xref>: pl. B51), also see MycoBank.</p><p>Descriptions of the species — <xref rid="R17" ref-type="bibr">Brandrud et al. (1992</xref>: pl. B51), <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012</xref>: 793).</p><p><bold><italic>Cortinarius claricolor</italic></bold> (Fr.) Fr., Epicr. Syst. Mycol.: 257. 1838</p><p><italic>Basionym</italic>. <italic>Agaricus multiformis</italic> δ <italic>claricolor</italic> Fr., Observ. Mycol. 2: 65. 1818.</p><p>= <italic>Cortinarius pseudoturmalis</italic> Bidaud & Moënne-Locc., Atlas des Cortinaires 19: 1503. 2010.</p><p><italic>Neotype</italic>. S<sc>WEDEN</sc>, Ångermanland, Stigsjö, Uland, Långmyrberget, in spruce forest with blueberry, 9 Aug. 1988, <italic>Brandrud et al.</italic> CFP691 (S), designated here. MycoBank MBT176396. GenBank KF732283.</p><p>Illustrations — <xref rid="R9" ref-type="bibr">Bidaud et al. (2010</xref>: pl. 759), <xref rid="R17" ref-type="bibr">Brandrud et al. (1992</xref>: pl. B48), <xref rid="R29" ref-type="bibr">Fries (1867–1884</xref>: pl. 141).</p><p>Descriptions of the species — <xref rid="R17" ref-type="bibr">Brandrud et al. (1992</xref>: pl. B48), <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012</xref>: 821).</p><p><bold><italic>Cortinarius cumatilis</italic></bold> Fr., Epicr. Syst. Mycol.: 269. 1838</p><p><italic>Neotype</italic>. S<sc>WEDEN</sc>, Närke, Hidinge, Garphyttans National Park, N of the road, grass-herb forest with <italic>Corylus</italic>, <italic>Populus tremula</italic>, <italic>Ulmus</italic>, <italic>Fagus</italic>, <italic>Quercus</italic> and <italic>Picea</italic>, 150 m asl, 20 Sept. 1998, <italic>I. Kytövuori</italic> 98-2164 (H; NY isoneotype), designated here. MycoBank MBT176397. GenBank KF732293.</p><p>Illustrations — <xref rid="R16" ref-type="bibr">Brandrud et al. (1990</xref>: pl. A47), <xref rid="R29" ref-type="bibr">Fries (1867–1884</xref>: pl. 146).</p><p>Descriptions of the species — <xref rid="R16" ref-type="bibr">Brandrud et al. (1990</xref>: pl. A47), <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012</xref>: 824).</p><p><bold><italic>Cortinarius cyanites</italic></bold> Fr., Epicr. Syst. Mycol.: 279. 1838</p><p>=<italic> Cortinarius pseudocyanites</italic> var. <italic>paucus</italic> Reumaux, Atlas des Cortinaires 15: 1032. 2005.</p><p>=<italic> Cortinarius subcyanites</italic> Bidaud, Atlas des Cortinaires 15: 1032. 2005.</p><p><italic>Neotype</italic>. S<sc>WEDEN</sc>, Uppland, Uppsala, Stadsskogen, mixed forest, 26 Aug. 2005, <italic>A. Taylor</italic> 2005069 (S; UPS isoneotype), designated here. MycoBank MBT176398. UNITE No. UDB002193, GenBank KF732355.</p><p>Illustrations — Bidaud et al. (2005: pl. 534), <xref rid="R29" ref-type="bibr">Fries (1867–1884</xref>: pl. 152).</p><p><italic>Pileus</italic> 3–9 cm broad, convex with persistently incurved margin, plano-convex when old, innately fibrillose, greyish blue to greyish brown at centre, greyish blue towards the margin, first viscid but soon dry. <italic>Lamellae</italic> emarginate, medium spaced to almost crowded, first dark violet, later brownish violet. <italic>Stipe</italic> 4.5–10 cm long, 1–1.5 cm thick at apex, 1.5–4 cm at base, clavate, blue to greyish blue. <italic>Universal veil</italic> greyish to brownish grey, forming complete and incomplete fibrillose girdles on stipe. <italic>Context</italic> bluish white in pileus, stronger blue adjacent to lamellae, bluish in stipe, becoming vinaceous red on exposure, marbled hygrophanous. <italic>Odour</italic> pleasant, fruity (according to <xref rid="R4" ref-type="bibr">Bidaud et al. 2005</xref>). <italic>Exsiccata</italic>: pileus pale bluish grey to lilac grey, when young with the same tint as <italic>C. traganus</italic>, often with a pale brownish tint at the centre, weakly fibrillose, stipe bluish grey, brownish at the base.</p><p><italic>Spores</italic> 8.8–<italic>9.8</italic>–10.9 × 5.2–<italic>5.7</italic>–6.3 μm, av. = 9.5–10.0 × 5.5–5.8 μm, Q = 1.59–1.72–1.84, Qav. = 1.66–1.77 (6 specimens, 240 spores), amygdaloid, with a fairly narrow apex, moderately verrucose, some with few small golden yellow guttules, fairly faintly dextrinoid. <italic>Basidia</italic> 32–41 × 7–9 μm (80 basidia), 4-spored, narrowly clavate, with blood red guttules, many with yellowish contents. <italic>Lamellar trama hyphae</italic> yellowish, smooth, with abundant small to large to worm-like blood red guttules. <italic>Stipe apex hyphae</italic> almost colourless to yellowish, smooth, with abundant small to large to worm-like, blood red guttules, the guttulate layer thick. On the surface few bands of entangled, ochraceous hyphae mostly without guttules. <italic>Pileipellis</italic>: epicutis very weakly gelatinous, uppermost hyphae 3–10 μm wide, very pale ochraceous brownish, mostly very sparsely, spot-like incrusted, often with scanty to abundant small blood red guttules, lower down 3–9 μm wide, almost colourless, smooth hyphae with abundant small to large to worm-like, blood red guttules. Hypoderm not developed.</p><p>Ecology & Distribution — In mixed forests of coniferous and deciduous trees, host unknown.</p><p>Notes — We studied the <italic>C. cyanites</italic> species complex and recognized three different species, <italic>C. cyanites </italic>s.str.,<italic>C.boreicyanites</italic> and <italic>C. violaceorubens</italic>. All the species were well supported in our phylogenetic analysis and differ from one another by more than 20 substitutions and indel positions. The most distinct of the species is <italic>C. violaceorubens</italic>. It has a dark, violaceous-brownish pileus, the exsiccata are dirty violaceous grey to violaceous brown, and the spores are largest of the group (av. 9.9–11.0 × 6.3–6.6 μm). It grows in <italic>Picea</italic> dominated forests, often on rich soil and is known from France, Germany, Sweden and Finland. Sister species <italic>C.cyanites</italic> and <italic>C. boreicyanites</italic> both have smaller spores and paler exsiccata. Of these, <italic>C. boreicyanites</italic> is so far only known from the middle boreal zone of Sweden, Finland and Scotland while <italic>C. cyanites</italic> has a more southern distribution extending from South Finland and middle Sweden to France. In the protologue of <italic>C. cyanites</italic> Fries described a species with a “pileo pallide coeruleo”. In addition, the picture of <italic>C. cyanites</italic> in <xref rid="R29" ref-type="bibr">Fries (1867–1884</xref>: pl. 152) illustrates a species with a bluish grey pileus. Attached at the base of the stipe there are leaves of <italic>Quercus</italic> and <italic>Betula</italic> and spruce needles. Based on the protologue, illustration, and ecology and distribution of the three species we conclude that the species best fitting the description of Fries is the one described here as <italic>C. cyanites</italic> and we here propose collection <italic>A. Taylor 2005069</italic> as neotype for the species.</p><p><bold><italic>Cortinarius fraudulosus</italic></bold> Britzelm., Ber. Naturhist. Vereins Augsburg 4: 122. 1885</p><p><italic>Neotype</italic>.G<sc>ERMANY</sc>, Baden-Württemberg, Freudenstadt, Heiligenbronn, gently sloping grass-herb conifer forest on calcareous soil, <italic>Abies alba</italic>,<italic> Picea abies</italic>,<italic> Fagus sylvatica</italic>, 5 Oct. 1995, <italic>I. Kytövuori</italic> 95-1852, H7019563 (H; NY isoneotype), designated here. MycoBank MBT176399. GenBank KF732518.</p><p>Illustration — <xref rid="R1" ref-type="bibr">Abarenkov et al. (2010</xref>: photo under the accession no. UDB011906).</p><p><italic>Pileus</italic> 4–8 cm broad, hemispherical to convex, then plano-convex, fibrillose, white to ochraceous white when young, with age becoming pale brownish. <italic>Lamellae</italic> emarginate, almost distant, first white to very pale brownish grey, later pale brown. <italic>Stipe</italic> 5–10 cm long, 1–2 cm thick at apex, 1.5–3.5 cm wide at base, clavate to almost bulbose, sometimes slightly rooting, whitish, with handling and with age becomes brownish. <italic>Universal veil</italic> white, forming distinct girdles on stipe, sometimes floccose. <italic>Context</italic> white. <italic>Odour</italic> not recorded. <italic>Exsiccata</italic> pale greyish brown.</p><p><italic>Spores</italic> 12.9–<italic>13.7</italic>–15.0 × 7.3–<italic>8.1</italic>–8.8 μm, Q = 1.60–<italic>1.70</italic>–1.78 (1 specimen, 20 spores), amygdaloid, with a narrow apex, fairly strongly verrucose, warts anastomosing, not high, some with dark intracellular granules, moderately dextrinoid. <italic>Basidia</italic> 42–53 × 10–12 μm (20 basidia), 4-spored, clavate, otherwise colourless but with a few dark granular bodies. <italic>Lamellar trama hyphae</italic> sand brown, full of small dark granules or particles. <italic>Stipe apex hyphae</italic> yellow brown to red brown, entangled, otherwise colourless, but with small to large to long bodies of red brown to red blackish to black granules, uppermost hyphae 5–10 μm wide. <italic>Pileipellis</italic>: epicutis fairly weakly gelatinous, hyphae 5–15 μm wide, ochraceous brownish, finely, densely incrusted, with scanty red brown granules. Hypoderm present. In the overall view red brown and unstructured when seen from above.</p><p>Ecology & Distribution — In montane and hemiboreal conifer forests, calcicolous. Known from Germany, Italy and Estonia. Fruiting in autumn.</p><p>Notes — The material of <italic>C. fraudulosus</italic> formed two well supported groups in our phylogenetic analysis. One group consisted of specimens collected from Germany, Italy and Estonia while the other group included mainly specimens from northern Europe. Since <italic>C. fraudulosus</italic> Britzelm. is described from Germany, the neotype is chosen among the Central/southern European clade and the other is described below as a new species <italic>C. subfraudulosus</italic>.</p><p><bold><italic>Cortinarius glaucopus</italic></bold> (Schaeff.: Fr.) Gray, Nat. Arr. Brit. Pl. 1: 629. 1821</p><p><italic>Basionym</italic>. <italic>Agaricus glaucopus</italic> Schaeff., Fung. Bavar. Palat. 4: 23. 1774: sanctioned in Fr., Syst. Mycol. 1: 224. 1821.</p><p>= <italic>Cortinarius glaucopoides</italic> Kauffman, Papers from the Michigan Academy of Science, Arts and Letters 1: 133. 1923.</p><p><italic>Neotype</italic>. S<sc>WEDEN</sc>, Medelpad, Alnö, Ås brygga, in dry spruce forest on calcareous soil, 21 Sept. 1988, <italic>Brandrud et al.</italic> CFP786 (S), designated here. MycoBank MBT176400. GenBank KF732315.</p><p>Illustration — <xref rid="R18" ref-type="bibr">Brandrud et al. (1994</xref>: pl. C30).</p><p>Description of the species — <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012</xref>: 802).</p><p>Notes <italic>— </italic>The description of <italic>Agaricus glaucopus</italic> in <xref rid="R26" ref-type="bibr">Fries (1821)</xref> is short but fits our species. In addition, an unpublished plate of <italic>C. glaucopus</italic> painted with the supervision of Fries (S, 0318) exists. It represents a species with red brown pileus and bluish lamellae. It is very similar to the photograph of collection CFP786 in <xref rid="R18" ref-type="bibr">Brandrud et al. (1994)</xref>, which we propose as a neotype for the species. Our observations of <italic>C.glaucopus</italic> are consistent with those of <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012)</xref>, only the length of the spores is slightly different: our measurements 7.3–<italic>8.1</italic>–9 × 4.5–<italic>5.0</italic>–5.5 μm, Q = 1.54–<italic>1.63</italic>–1.76, <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012)</xref>: 7–8.5 × 4.5–5.5 μm.</p><p><bold><italic>Cortinarius herpeticus</italic></bold> Fr., Epicr. Syst. Mycol.: 268. 1838</p><p>= <italic>Cortinarius thalliopurpurascens </italic>Rob. Henry, Doc. Mycol. 25 (no. 97): 48. 1995.</p><p><italic>Neotype</italic>. S<sc>WEDEN</sc>, Ångermanland, Säbrå, Hällenyland, under <italic>Picea</italic> on cultivated area, 20 July 1990, <italic>Brandrud et al.</italic> CFP936 (S), designated here. MycoBank MBT176401. GenBank KF732321.</p><p>Illustration — <xref rid="R18" ref-type="bibr">Brandrud et al. (1994</xref>: pl. C08, as <italic>C. scaurus</italic> var. <italic>herpeticus</italic>).</p><p>Descriptions of the species — <xref rid="R18" ref-type="bibr">Brandrud et al. (1994</xref>: pl. C08), <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012</xref>: 784).</p><p>Notes — An unpublished plate of <italic>C. herpeticus</italic> (S, 0324) painted with the supervision of Fries exists. The basidiomes in the illustration are similar to the ones in the photo of collection CFP936 (<xref rid="R18" ref-type="bibr">Brandrud et al. 1994</xref>), which we propose as a neotype for the species. <italic>Cortinarius herpeticus</italic> is distinguished from <italic>C. scaurus</italic> by a stouter appearance, paler colours, more strongly ornamented spores and above all the lack of sepia-coloured pigments in the epicutis. <italic>Cortinarius violaceonitens</italic> is fairly similar, but is separated by the spores, which are narrower (9.3–<italic>10.0</italic>–10.7 × 5.4–<italic>5.9</italic>–6.3 μm), amygdaloid-fusoid, with a shallow suprahilar depression, a low ventral humb and blunt apex, and very strongly verrucose surface especially at the apex. The spores of <italic>C. herpeticus</italic> are 9.1–<italic>10.3</italic> 11.6 ×5.7–<italic>6.4</italic>–7.0 μm, narrowly oblong-ellipsoid, and moderately to strongly verrucose.</p><p><bold><italic>Cortinarius infractus</italic></bold> (Pers.: Fr.) Fr., Epicr. Syst. Mycol.: 261. 1838</p><p><italic>Basionym</italic>. <italic>Agaricus infractus</italic> Pers., Observ. Mycol. 2: 42. 1800 (1799): sanctioned in Fr., Syst. Mycol. 1: 223. 1821.</p><p>= <italic>Cortinarius amarocaerulescens</italic> Bidaud, Atlas des Cortinaires 18: 1376. 2009.</p><p> = <italic>Cortinarius infractus</italic> var. <italic>aeruginosus</italic> Reumaux, Atlas des Cortinaires 18: 1376. 2009.</p><p><italic>Neotype</italic>. S<sc>WEDEN</sc>, Bohuslän, Tossene, Anneröd, beech forest, medium rich soil, 15 Sept. 1986, <italic>Brandrud et al.</italic> CFP495 (S), designated here. MycoBank MBT176402. GenBank KF732325.</p><p>Illustrations — <xref rid="R8" ref-type="bibr">Bidaud et al. (2009</xref>: pl. 739), <xref rid="R16" ref-type="bibr">Brandrud et al. (1990</xref>: pl. A09).</p><p>Description of the species — <xref rid="R16" ref-type="bibr">Brandrud et al. (1990</xref>: pl. A09).</p><p><bold><italic>Cortinarius largus</italic></bold> Fr., Epicr. Syst. Mycol.: 259. 1838</p><p>= <italic>Cortinarius cephalixolargus</italic> Rob. Henry, Bull. Trimestriel Soc. Mycol. France 93, 3: 323. 1977.</p><p>= <italic>Cortinarius clarus</italic> Reumaux, Atlas des Cortinaires 8: 291. 1996.</p><p>= <italic>Cortinarius claviceps</italic> Reumaux, Atlas des Cortinaires 8: 291. 1996.</p><p>= <italic>Cortinarius congeminus</italic> Moënne-Locc. & Reumaux, Atlas des Cortinaires 7: 228. 1995.</p><p>= <italic>Cortinarius cupreoviolaceus</italic> Bidaud & Reumaux, Atlas des Cortinaires 8: 292. 1996.</p><p>= <italic>Cortinarius largusiellus</italic> Reumaux, Atlas des Cortinaires 8: 293. 1996.</p><p>= <italic>Cortinarius lilacinicolor</italic> Reumaux, Atlas des Cortinaires 8: 294. 1996.</p><p>= <italic>Cortinarius lintrisporus</italic> Reumaux, Doc. Mycol. 27, no. 106: 53. 1997.</p><p>= <italic>Cortinarius lividoviolaceus</italic> Rob. Henry, Doc. Mycol. 17, no. 68: 27. 1987.</p><p>= <italic>Cortinarius occultus</italic> Moënne-Locc. & Reumaux, Atlas des Cortinaires 8: 295. 1996.</p><p>= <italic>Cortinarius patibilis</italic> var. <italic>scoticus</italic> Brandrud, Edinburgh J. Bot. 54, 1: 114. 1997.</p><p>= <italic>Cortinarius paracrassus</italic> Reumaux, Atlas des Cortinaires 7: 230. 1995.</p><p>= <italic>Cortinarius paracyanopus</italic> Moënne-Locc. & Reumaux, Atlas des Cortinaires 8: 296. 1996.</p><p>= <italic>Cortinarius subspadiceus</italic> Reumaux, Atlas des Cortinaires 8: 298. 1996.</p><p><italic>Neotype</italic>. F<sc>INLAND</sc>, Varsinais-Suomi, Turku, Ruissalo, deciduous forest of <italic>Quercus robur</italic>,<italic> Corylus avellana </italic>and some <italic>Betula</italic> on mull soil, 3 Sept. 2008, <italic>K. Liimatainen & T. Niskanen</italic> 08-060, H6001957 (H; NY isoneotype), designated here. MycoBank MBT176403. GenBank AB859985.</p><p>Illustration — <xref rid="R19" ref-type="bibr">Brandrud et al. (1998</xref>: pl. D22).</p><p>Descriptions of the species — <xref rid="R15" ref-type="bibr">Brandrud (1998)</xref>, <xref rid="R19" ref-type="bibr">Brandrud et al. (1998</xref>: pl. D22), <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012</xref>: 815).</p><p>Notes — Based on morphological and molecular data <italic>C. largus</italic> seems like a uniform species. The collection CFP1085 (<xref rid="R19" ref-type="bibr">Brandrud et al. 1998</xref>), however, is from France and therefore not ideal as a type for a species described from Sweden. We do not have our own, well-documented specimen from Sweden and therefore we propose the collection <italic>K. Liimatainen & T. Niskanen</italic> 08-060 from hemiboreal deciduous forest from south western Finland as a neotype for the species.</p><p><bold><italic>Cortinarius multiformis</italic></bold> Fr., Epicr. Syst. Mycol.: 263. 1838</p><p><italic>Neotype</italic>. S<sc>WEDEN</sc>, Ångermanland, Häggdånger, Sjö, spruce forest with blueberry, 21 Aug. 1986, <italic>Brandrud et al.</italic> CFP445 (S), designated here. MycoBank MBT176404. GenBank KF732350.</p><p>Illustration — <xref rid="R16" ref-type="bibr">Brandrud et al. (1990</xref>: pl. A45).</p><p>Descriptions of the species — <xref rid="R16" ref-type="bibr">Brandrud et al. (1990</xref>: pl. A45), <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012</xref>: 808).</p><p>Notes — The species as neotypified here fits best the original description of the species and the unpublished plate of <italic>C. multiformis</italic> (S0350). The reminiscent sister species <italic>C. talimultiformis</italic>, which has been mixed with <italic>C. multiformis</italic>, has white fibrils on the pileus, and less dextrinoid spores. Our observations of <italic>C. multiformis</italic> are in concordance with those of <xref rid="R16" ref-type="bibr">Brandrud et al. (1990)</xref> and <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012</xref>: 808), except that our spore measurements are somewhat larger, 8.6–<italic>9.6</italic>–10.4 × 5.2–<italic>5.7</italic>–6.1 μm, Q = 1.58–<italic>1.70</italic>–1.79 than those of <xref rid="R16" ref-type="bibr">Brandrud et al. (1990)</xref> and <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012)</xref> 8–9.5 × 5–5.5 μm.</p><p><bold><italic>Cortinarius pansa </italic></bold>(Fr.) Sacc., Syll. Fung. 5: 901. 1887</p><p><italic>Basionym</italic>. <italic>Agaricus pansa</italic> Fr., Observ. Mycol. (Havniae) 2: 67. 1818.</p><p><italic>Neotype</italic>. F<sc>INLAND</sc>, Varsinais-Suomi, Kemiö, Pederså, at small, abandoned limestone quarries, spruce heath forest, roadside, 35 m asl, 21 Sept. 1990,<italic> I. Kytövuori</italic> 90-1826 (H; isoneotype NY), designated here. MycoBank MBT176405. GenBank KF732522.</p><p>Illustration — <xref rid="R29" ref-type="bibr">Fries (1867–1884</xref>: pl. 145).</p><p>Description of the species — <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012</xref>: 803).</p><p>Notes — The description and illustration of <italic>C. pansa</italic> published by <xref rid="R25" ref-type="bibr">Fries (1818</xref>, <xref rid="R29" ref-type="bibr">1867–1884</xref>) fit well with the species presented in <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012)</xref>. The spore measurements given in <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012)</xref> (6–)6.5–7.5 × 4–5 μm differ somewhat from ours 6.8–<italic>7.5</italic>–8.2 × 4.3–<italic>4.6</italic>–5.0 μm, Q = 1.52–<italic>1.64</italic>–1.82. The species was previously included in <italic>C. glaucopus</italic> but differs from it by more red brown pileus, smaller, less verrucose spores, and habitat often on roadsides, yards, parks and plantations.</p><p><bold><italic>Cortinarius percomis</italic></bold> Fr., Epicr. Syst. Mycol.: 260. 1838</p><p><italic>Neotype</italic>. F<sc>INLAND</sc>, Varsinais-Suomi, Karjaa, Kohagen, herb-rich <italic>Picea abies</italic> forest with some <italic>Corylus avellana</italic>,<italic> Quercus robur</italic>,<italic> Betula</italic> and <italic>Populus tremula</italic>, 2 Sept. 2008, <italic>K. Liimatainen & T. Niskanen </italic>08-041 (H; isoneotype NY), designated here. MycoBank MBT176406. GenBank KF732380.</p><p>Illustrations — <xref rid="R18" ref-type="bibr">Brandrud et al. (1994</xref>: pl. C56), <xref rid="R29" ref-type="bibr">Fries (1867–1884</xref>: pl. 143).</p><p>Descriptions of the species — <xref rid="R18" ref-type="bibr">Brandrud et al. (1994</xref>: pl. C56), <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012</xref>: 812).</p><p>Notes — Based on morphological and molecular data <italic>C. percomis</italic> seems like a uniform species. The collection CFP1104 (<xref rid="R18" ref-type="bibr">Brandrud et al. 1994</xref>), however, is from France and therefore not ideal as a type for a species described from Sweden. We do not have our own, well-documented specimen from Sweden and therefore we propose the collection <italic>K. Liimatainen & T. Niskanen</italic> 08-041 from hemiboreal <italic>Picea abies</italic> dominated forest from south western Finland as a neotype for the species. An identical ITS sequence of the species from a specimen collected from Sweden, however, exists in UNITE (UDB000726).</p><p><bold><italic>Cortinarius porphyropus</italic></bold> (Alb. & Schwein.) Fr., Epicr. Syst. Mycol.: 271. 1838</p><p><italic>Basionym</italic>. <italic>Agaricus porphyropus</italic> Alb. & Schwein., Consp. Fungorum Lusat.: 153. 1805.</p><p>= <italic>Cortinarius porphyropus</italic> var. <italic>porphyrophorus</italic> Reumaux, Atlas des Cortinaires 18: 1378. 2009.</p><p><italic>Neotype</italic>. S<sc>WEDEN</sc>, Jämtland, Ragunda sn, Ragunda, in birch forest on rich soil, 20 Aug. 1988, <italic>Brandrud et al.</italic> CFP717 (S), designated here. MycoBank MBT176407. GenBank KF732387.</p><p>Illustration — <xref rid="R17" ref-type="bibr">Brandrud et al. (1992</xref>: pl. B55).</p><p>Descriptions of the species — <xref rid="R17" ref-type="bibr">Brandrud et al. (1992</xref>: pl. B55), <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012</xref>: 819).</p><p>Notes — <italic>Cortinarius porphyropus</italic> is described from Germany. Based on our studies it is a uniform and widespread species occurring at least in Europe and North America (<xref ref-type="fig" rid="F1">Fig. 1</xref>). The most representative collection CFP717 from Sweden is here proposed as a neotype for the species.</p><p><bold><italic>Cortinarius praestans</italic></bold> (Cordier) Gillet, Hyménomycètes: 475. 1874</p><p><italic>Basionym</italic>. <italic>Agaricus praestans</italic> Cordier, Champ. France, Discom.: 98. 1870.</p><p><italic>Holotype</italic>. Pl. XXI (Cordier, Champ. France, 1870).</p><p><italic>Epitype</italic>. F<sc>RANCE</sc>, Ain, Oyonnax SE, Commune d’Echallon, by the road from St-Germain-de-Joux to Echallon, N of the crossing to Plagne, E sloping, rich <italic>Fagus</italic> forest with <italic>Picea abies</italic>, 540 m asl, 27 Oct. 1994, <italic>P. & I. Kytövuori </italic>94-1861 (H; isoepitype NY). MycoBank MBT176411. GenBank KF732389.</p><p>Illustration — <xref rid="R16" ref-type="bibr">Brandrud et al. (1990</xref>: pl. A42).</p><p>Descriptions of the species — <xref rid="R16" ref-type="bibr">Brandrud et al. (1990</xref>: pl. A42), <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012</xref>: 823).</p><p>Notes — <italic>Cortinarius praestans</italic> is one of the most distinctive <italic>Cortinarius</italic> species and is easy to recognize by its large basidiomata and spores, and habitat with thermophilous deciduous trees. It is described from France, but identical sequences of the species exist from Estonia, Germany, Italy and Sweden (<xref ref-type="fig" rid="F1">Fig. 1</xref>, <xref ref-type="table" rid="T2">Table 2</xref>).</p><p><bold><italic>Cortinarius purpurascens</italic></bold> Fr., Epicr. Syst. Mycol.: 265. 1838</p><p>= <italic>Cortinarius eumarginatus</italic> Rob. Henry ex Bidaud, Carteret & Reumaux, Atlas des Cortinaires 18, 1, 2: 1378. 2009.</p><p><italic>Neotype</italic>. S<sc>WEDEN</sc>, Närke, Hidinge, Garphyttans National Park, S of the road, fairly rich spruce grass-herb forest with <italic>Corylus</italic>, <italic>Populus tremula</italic>, <italic>Betula </italic>and<italic> Quercus</italic>, 150 m asl, 20 Sept. 1998, <italic>I. Kytövuori</italic> 98-2121 (H; isoneotype NY), designated here. MycoBank MBT176419. GenBank KF732406.</p><p>Illustration — <xref rid="R8" ref-type="bibr">Bidaud et al. (2009</xref>: pl. 743).</p><p>Description of the species — <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012</xref>: 802).</p><p>Notes — Based on the data we have so far, the species is known from northern to southern Europe (<xref ref-type="fig" rid="F1">Fig. 1</xref>).</p><p><bold><italic>Cortinarius russus</italic></bold> Fr., Epicr. Syst. Mycol.: 261. 1838</p><p><italic>Neotype</italic>. S<sc>WEDEN</sc>, Ångermanland, Säbrä, Överdal, in dry spruce forest on rich soil, 3 Aug. 1990, <italic>Brandrud et al.</italic> CFP941 (S), designated here. MycoBank MBT176412. GenBank KF732416.</p><p>Illustration — <xref rid="R18" ref-type="bibr">Brandrud et al. (1994</xref>: pl. C35, C44).</p><p>Descriptions of the species — <xref rid="R18" ref-type="bibr">Brandrud et al. (1994</xref>: pl. C35), <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012</xref>: 817).</p><p><bold><italic>Cortinarius scaurus</italic></bold> (Fr.: Fr.) Fr., Epicr. Syst. Mycol.: 268. 1838</p><p><italic>Basionym</italic>. <italic>Agaricus scaurus</italic> Fr., Observ. Mycol. 2: 75. 1818.</p><p>= <italic>Cortinarius parolivascens</italic> Moënne-Locc. & Reumaux, Atlas des Cortinaires 18: 1375. 2009.</p><p>= <italic>Cortinarius scaurus</italic> var. <italic>notandus</italic> Bidaud, Atlas des Cortinaires 18: 1375. 2009.</p><p>= <italic>Cortinarius scaurus</italic> f. <italic>phaeophyllus</italic> M.M. Moser, Fungi non Delineati 15: 18. 2001.</p><p><italic>Neotype</italic>. S<sc>WITZERLAND</sc>, Bern, Fribourg, Rechthalten, on the border of a peat-bog with <italic>Pinus strobus</italic>, 29 Sept. 1991, <italic>Brandrud et al.</italic> CFP1074 (S), designated here. MycoBank MBT176413. GenBank KF732423.</p><p>Illustrations — <xref rid="R18" ref-type="bibr">Brandrud et al. (1994</xref>: pl. C21), <xref rid="R29" ref-type="bibr">Fries (1867–1884</xref>: pl. 146).</p><p>Descriptions of the species — <xref rid="R18" ref-type="bibr">Brandrud et al. (1994</xref>: pl. C21), <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012</xref>: 783).</p><p>Notes — <italic>Cortinarius scaurus</italic> is a widespread species and to date known from eastern North America and Europe (<xref ref-type="fig" rid="F1">Fig. 1</xref>). The most representative collection CFP1074 from Switzerland is proposed as a neotype and the ITS sequence of the specimen is identical to the UNITE sequence UDB001068 from Femsjö, Sweden.</p><p><bold><italic>Cortinarius serarius</italic></bold> Fr., Epicr. Syst. Mycol.: 269. 1838</p><p><italic>Neotype</italic>. S<sc>WEDEN</sc>, Ångermanland, Häggdånger, Torrom, in dry spruce forest on rich soil, 11 Aug. 1990, <italic>Brandrud et al.</italic> CFP959 (S), designated here. MycoBank MBT176414. GenBank KF732428.</p><p>Illustration — <xref rid="R18" ref-type="bibr">Brandrud et al. (1994</xref>: pl. C25).</p><p>Descriptions of the species — <xref rid="R18" ref-type="bibr">Brandrud et al. (1994</xref>: pl. C25), <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012</xref>: 824).</p><p><bold><italic>Cortinarius subpurpurascens</italic></bold> (Batsch) Fr., Epicr. Syst. Mycol.: 265. 1838</p><p><italic>Basionym</italic>. <italic>Agaricus subpurpurascens</italic> Batsch, Elench. Fung., cont. prim.: 71. 1786.</p><p>= <italic>Cortinarius largoides</italic> Rob. Henry ex Bidaud, Carteret & Reumaux, Atlas des Cortinaires 18: 1378. 2009.</p><p>= <italic>Cortinarius subinops</italic> Reumaux, Atlas des Cortinaires 18: 1379. 2009.</p><p><italic>Holotype</italic>. Batsch, Elench. Fung., cont. prim. tab. 16: 74. 1786.</p><p><italic>Epitype</italic>. F<sc>INLAND</sc>, Varsinais-Suomi, Turku, Ruissalo, deciduous forest of <italic>Quercus robur</italic>, <italic>Corylus avellana</italic> and some <italic>Betula</italic> on mull soil, 3 Sept. 2008, <italic>K. Liimatainen & T. Niskanen</italic> 08-059 (H; isoepitype NY), designated here. MycoBank MBT176420. GenBank KF732449.</p><p>Illustrations — <xref rid="R8" ref-type="bibr">Bidaud et al. (2009</xref>: pl. 749, 750).</p><p>Description of the species — <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012</xref>: 819).</p><p>Notes — Based on the data we have so far the species is known from northern to southern Europe (<xref ref-type="fig" rid="F1">Fig. 1</xref>). A representative collection <italic>Liimatainen & Niskanen</italic> 08-059 from South Finland is here proposed as an epitype of the species.</p><p><bold><italic>Cortinarius subtortus </italic></bold>(Pers.) Fr., Epicr. Syst. Mycol. (Upsaliae): 273. 1838</p><p><italic>Basionym</italic>. <italic>Agaricus subtortus</italic> Pers., Syn. Meth. Fung. (Göttingen) 2: 284. 1801, sanctioned in Fr., Syst. Mycol. 1: 222. 1821.</p><p><italic>Neotype</italic>. S<sc>WEDEN</sc>, Inre Småland, Femsjö, Prästskogen, partly paludified spruce forest with some deciduous tree species, 175 m asl, 18 Sept. 1987, <italic>I. Kytövuori</italic> 87-1510 (H; isoneotype NY), designated here. MycoBank MBT176415. GenBank KF732454.</p><p>Description of the species — <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012</xref>: 811).</p><p><bold><italic>Cortinarius talus</italic></bold> Fr., Epicr. Syst. Mycol.: 263. 1838</p><p>= <italic>Cortinarius aurantionapus</italic> Bidaud & Reumaux, Atlas des Cortinaires 16: 1096. 2006.</p><p>= <italic>Cortinarius crenulatus</italic> Rob. Henry ex Bidaud & Reumaux, Atlas des Cortinaires 16: 1097. 2006.</p><p>= <italic>Cortinarius ochropudorinus</italic> Rob. Henry ex Bidaud & Reumaux, Atlas des Cortinaires 16: 1097. 2006.</p><p>=<italic>Cortinarius pseudominor</italic> Rob. Henry ex Reumaux, Atlas des Cortinaires 16: 1098. 2006.</p><p>= <italic>Cortinarius pseudotalus</italic> Rob. Henry ex Bidaud & Reumaux, Atlas des Cortinaires 16: 1098. 2006.</p><p><italic>Neotype</italic>. S<sc>WEDEN</sc>, Jämtland, Ragunda sn, Ragunda, in birch forest on rich soil (<italic>Betula</italic>,<italic> Populus</italic>,<italic> Pinus</italic>), 26 Aug. 1989, <italic>Brandrud et al.</italic> CFP832 (S), designated here. MycoBank MBT176416. GenBank KF732457.</p><p>Illustrations — <xref rid="R17" ref-type="bibr">Brandrud et al. (1992</xref>: pl. B47), <xref rid="R29" ref-type="bibr">Fries (1867–1884</xref>: pl. 145).</p><p>Descriptions of the species — <xref rid="R17" ref-type="bibr">Brandrud et al. (1992</xref>: pl. B47), <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012</xref>: 811).</p><p>Notes — Our observations of the species are in concordance with those of <xref rid="R17" ref-type="bibr">Brandrud et al. (1992)</xref> and <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012)</xref>, except for the length of the spores, which according to our measurements is 7.3–<italic>8.0</italic>–8.8 ×4.5–<italic>5.0</italic>–5.2 μm and in <xref rid="R17" ref-type="bibr">Brandrud et al. (1992)</xref> and <xref rid="R52" ref-type="bibr">Jeppesen et al.(2012)</xref> 7.5–9.5 × 4.5–5.5 μm.</p><p><bold><italic>Cortinarius turmalis</italic></bold> Fr., Epicr. Syst. Mycol.: 257. 1838</p><p><italic>Neotype</italic>. S<sc>WEDEN</sc>, Medelpad, Borgsjö, Julåsen, in herbaceous spruce forest, 20 Aug. 1988, <italic>Brandrud et al.</italic> CFP716 (S), designated here. MycoBank MBT176417. GenBank KF732464.</p><p>Illustration — <xref rid="R18" ref-type="bibr">Brandrud et al. (1994</xref>: pl. C31).</p><p>Descriptions of the species — <xref rid="R18" ref-type="bibr">Brandrud et al. (1994</xref>: pl. C31), <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012</xref>: 821).</p><p><bold><italic>Cortinarius variecolor</italic></bold> (Pers.: Fr.) Fr., Epicr. Syst. Mycol.: 259. 1838</p><p><italic>Basionym</italic>. <italic>Agaricus variecolor</italic> Pers., Syn. Meth. Fung. 2: 280. 1801. Sanctioned in Fr., Syst. Mycol. 1: 222. 1821.</p><p>= <italic>Cortinarius muricinicolor</italic> Moënne-Locc., Atlas des Cortinaires 8: 295. 1996.</p><p>= <italic>Cortinarius piriodolens</italic> Moënne-Locc., Atlas des Cortinaires 8: 296. 1996.</p><p><italic>Neotype</italic>. S<sc>WEDEN</sc>, Gotland, Viklau, Tjaukle, in spruce forest on calcareous soil, 29 Sept. 1990, <italic>Brandrud et al.</italic> CFP1021 (S), designated here. MycoBank MBT176418. GenBank KF732466.</p><p>Illustrations — <xref rid="R17" ref-type="bibr">Brandrud et al. (1992</xref>: pl. B20), <xref rid="R29" ref-type="bibr">Fries (1867–1884</xref>: pl. 144).</p><p>Descriptions of the species — <xref rid="R15" ref-type="bibr">Brandrud (1998)</xref>, <xref rid="R17" ref-type="bibr">Brandrud et al. (1992</xref>: pl. B20), <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012</xref>: 815).</p></sec></sec><sec id="s4"><title>NEW SPECIES AND COMBINATIONS</title><sec id="s4a"><title>Species with an isolated position</title><p><bold><italic>Cortinarius cremeiamarescens</italic></bold> Kytöv., Liimat. & Niskanen, <italic>sp. nov</italic>. — MycoBank MB805786; <xref ref-type="fig" rid="F2">Fig. 2</xref>a, <xref ref-type="fig" rid="F3">3</xref>a</p><p><italic>Etymology</italic>. The name refers to the colour of the basidiomata and bitter taste of pileus cuticle.</p><p>= <italic>Cortinarius caesiostramineus</italic> Rob. Henry sensu <xref rid="R16" ref-type="bibr">Brandrud et al. 1990</xref>, <xref rid="R52" ref-type="bibr">Jeppesen et al. 2012</xref>, p.p.</p><p><italic>Type</italic>. S<sc>WEDEN</sc>, Gotland, Alskog and När parish, Ollajvs Nature Reserve, mesic to damp spruce forest with some <italic>Pinus</italic>,<italic> Quercus</italic> and <italic>Corylus</italic>, 27 Sept. 2011, <italic>I. Kytövuori</italic> 11-014 (holotype H; isotype NY). GenBank KF732493.</p><p><italic>Pileus</italic> 3.5–5.5 cm broad, hemispherical to convex, then expanded, very finely innately fibrillose, cream-coloured to pale ochraceous yellow. <italic>Lamellae</italic> emarginate, almost crowded, pale greyish brown. <italic>Stipe</italic> 6.5–9.5 cm long, 0.7–1.4 cm thick at apex, 1–2.3 cm wide at base, with slightly marginate bulb, at first white, becoming pale brownish yellow with age. <italic>Universal veil</italic> white, sparse, at bulb margin. <italic>Context</italic> white. <italic>Odour</italic> indistinct. <italic>Taste:</italic> pileus cuticle bitter. <italic>Exsiccata</italic>: pileus ochraceous clay-colour to ochraceous yellow to warm ochraceous brown especially at the centre, stipe pale grey to brown.</p><p>In MLZ: <italic>Spores</italic> 7.0–<italic>7.8</italic>–8.8 × 4.3–<italic>4.7</italic>–5.0 μm, av. = 7.5–8.3 × 4.6–4.9 μm, Q = 1.54–<italic>1.67</italic>–1.87, Qav. = 1.59–1.74 (9 specimens, 260 spores, <xref ref-type="fig" rid="F3">Fig. 3</xref>a), citriform to narrowly fusoid, beaked, thin-walled, fairly finely, densely, and often sharply verrucose, slightly to moderately dextrinoid. <italic>Basidia</italic> 24–34 × 6.5–8 μm (80 basidia), 4-spored, narrowly clavate, very thin-walled, colourless, more or less filled with blood red drops. <italic>Lamellar trama hyphae</italic> yellow, filled with small blood red drops, larger globose and worm-like guttules fairly scanty (especially more scanty than in <italic>C. gentianeus</italic>). <italic>Stipe apex hyphae</italic> almost colourless to yellow, smooth, full of small golden yellow to blood red drops, larger globose and worm-like guttules fewer. <italic>Pileipellis</italic>: epicutis strongly gelatinous, hyphae 2–6 μm wide, very thin-walled and difficult to define, filled with small to medium-sized blood red guttules, mostly evenly distributed in the hyphae. Hypoderm present, pale yellow. In <italic>C. gentianeus</italic> the epicutis hyphae are full of very long, worm-like, foamy guttules and the blood red layer is much thicker than in <italic>C. cremeiamarescens</italic>.</p><p>Ecology & Distribution — In hemiboreal and southern boreal conifer-dominated forests on rich to calcareous soil. Known from southern Europe and western North America, British Columbia and Alaska. Fruits from late August to late October.</p><p><italic>Other specimens examined</italic>. F<sc>INLAND</sc>, Varsinais-Suomi, Parainen, Lemlahdensaari, Fallskogen, <italic>Pinus sylvestris</italic> heath forest with some <italic>Picea abies</italic> on sandy soil, by the calcareous dust road, 20 Oct. 2006, <italic>K. Liimatainen & T. Niskanen</italic> 06-273; Uusimaa, Espoo, Luukkaa outdoor recreation area, N of Haukkalampi, mesic, partly grass-herb-spruce forest with some <italic>Populus tremula</italic>,<italic> Betula</italic> and <italic>Pinus sylvetris</italic>, 9 Sept. 2004, <italic>K. Liimatainen & T. Niskanen</italic> 04-768, H6029461; Kirkkonummi, Dorgarn, Meiko-Trehörningen nature reserve, semi-open spruce forest with some hardwood bushes, 27 Sept. 2007, <italic>I. Kytövuori</italic> 07-1722, H6001575 (H); Siuntio, Lappträsk, grass-herb-spruce forest with some <italic>Pinus</italic>,<italic> Betula</italic>,<italic> Populus</italic> and <italic>Corylus</italic>, 24 Aug. 2000, <italic>I. Kytövuori</italic> 00-027 (H); Etelä-Savo, Mäntyharju, Juolasvesi, Hietaniemi, Sojonkangas, fairly rich spruce-pine forest with abundant <italic>Betula</italic> and <italic>Populus tremula</italic>, 29 Sept. 1994, <italic>I. Kytövuori</italic> 94-1177b, H6035734 (H). – S<sc>WEDEN</sc>, Västergötland, Undenäs, c. 1.5 km NNW of Sätra bruk, herb-rich <italic>Picea abies</italic> forest with some <italic>Betula</italic>,<italic> Populus</italic> and <italic>Pinus</italic>, 6 Sept. 2003, <italic>T. Niskanen et al.</italic> 03-1255, H7018363 (H); F03-1163, H7018395 (H); Närke. Hidinge, Garphyttans Nationalpark, herb-rich <italic>Picea abies</italic> forest with <italic>Corylus</italic>,<italic> Populus tremula</italic>,<italic> Betula</italic> and <italic>Quercus</italic>, 26 Sept. 2004, <italic>T. Niskanen</italic><italic>et al.</italic> 03-1255, H7017775 (H).</p><p><italic>Additional specimens</italic>. C<sc>ANADA</sc>, British Columbia, Interior Cedar Hemlock Forest, mycorrhizal root tip of <italic>Betula papyrifera</italic>, isolate UBCOCS153F, GenBank EF218754. – USA, Alaska, Delta Junction, source: boreal forest, soil 0–20 cm, GenBank JN889840.</p><p>Notes — <italic>Cortinarius cremeiamarescens</italic> was previously confused taxonomically with <italic>C. gentianeus</italic> Rob. Henry (= <italic>C. caesiostramineus</italic> sensu <xref rid="R52" ref-type="bibr">Jeppesen et al. 2012</xref> p.p.). However, the latter species is larger, typically has paler, more whitish or greyish exsiccata and larger (7.7–<italic>8.5</italic>–9.3 × 4.8–<italic>5.2</italic>–5.4 μm, Q = 1.54–<italic>1.65</italic>–1.76), amygdaloid to citriform, less thin-walled, more dextrinoid, and more verrucose spores, and much more abundant large, blood red, foamy, worm-like guttules in the pileipellis and lamellar trama. <italic>Cortinarius cremeiamarescens</italic> formed a well-supported clade in our phylogenetic analysis (1.00 PP). The ITS sequences of the species are identical and it differs from its sister species <italic>C. gentianeus</italic> by 17 substitutions and indel positions. Further relationships with other species of <italic>Cortinarius</italic> were not resolved in our analysis.</p><p><bold><italic>Cortinarius flavivelatus</italic></bold> Kytöv., Liimat. & Niskanen, <italic>sp. nov.</italic> — MycoBank MB805863; <xref ref-type="fig" rid="F2">Fig. 2</xref>b, <xref ref-type="fig" rid="F3">3</xref>b</p><p><italic>Etymology</italic>. The name refers to the yellow universal veil.</p><p><italic>Type</italic>. S<sc>WEDEN</sc>, Norrbotten, Pajala, Junosuando, Nature Reserve Area between Sarvikero and Tulemajoki, dryish <italic>Picea abies</italic> heath forest with <italic>Pinus</italic>,<italic> Betula</italic>, and with open meadows, 15 Aug. 1998, <italic>I. Kytövuori</italic> 98-885 (holotype H; isotype NY). GenBank KF732528.</p><p><italic>Pileus</italic> 5–8 cm broad, hemispherical to convex, then plano-convex, viscid, finely innately fibrillose, olive brown to ochraceous brown to brown at the centre, lighter at the margin, with hygrophanous streaks. <italic>Lamellae</italic> emarginate, crowded, first distinctly pale bluish, later pale brown with a bluish tint. <italic>Stipe</italic> 6–10 cm long, 1–1.8 cm thick at apex, 1.5–2.5 cm wide at base, clavate to almost cylindrical, whitish, with a bluish tint at the apex. <italic>Universal veil</italic> yellow, forming girdles on stipe, somewhat viscid. <italic>Context</italic> white in pileus and lower part of the stipe, bluish at stipe apex. <italic>Odour</italic> indistinct. <italic>KOH-reaction</italic> negative in all parts. <italic>Exsiccata</italic>: pileus warm yellowish to reddish brown, stipe whitish.</p><p>In MLZ: <italic>Spores</italic> 9.1–<italic>9.8</italic>–10.9 × 5.0–<italic>5.4</italic>–5.9 μm, Q = 1.67–<italic>1.82</italic>–1.98 (1 specimen, 60 spores, <xref ref-type="fig" rid="F3">Fig. 3</xref>b), amygdaloid-fusoid to slightly, narrowly citriform, with a shallow suprahilar depression and somewhat beaked apex, fairly finely, separately verrucose, slightly dextrinoid. <italic>Basidia</italic> 27–38 × 7.5–8 μm (40 basidia), 4-spored, clavate, pale sand brown, with few dark red brown granules, almost hyaline when mature. <italic>Lamellar trama hyphae</italic>: with abundant dark granules and chips, sometimes in small mounds. <italic>Stipe apex hyphae</italic> pale yellowish sand brown with small brown granules, outermost hyphae entangled, narrow, with more or less abundant blackish red granules. <italic>Pileipellis</italic>: epicutis strongly gelatinous, uppermost hyphae 5–10 μm wide, ochre brown, finely to strongly spirally incrusted, mostly not granulose, lower down dark-granulose. Hypoderm well developed, red brown. The upper part of the hypoderm and the transition hyphae towards the epicutis strongly incrusted and with mounds of blackish brown granules. Evenly distributed small granules present (sometimes absent) between the mounds.</p><p>Ecology & Distribution — In northern boreal coniferous forests. Known from Sweden, Norrbotten. No sequences of this species exist in public databases.</p><p>Notes — Based on morphology and molecular data <italic>C. flavivelatus</italic> is a sister species of <italic>C. pini </italic>Brandrud. <italic>Cortinarius pini</italic>, however, has white, sometimes ochraceous white universal veil and much larger spores (10.7–<italic>11.7</italic>–12.9 × 6.3–<italic>6.8</italic>–7.3 μm). In ITS regions the difference between the species is 17 substitutions and indel positions.</p><p><bold><italic>Cortinarius kytoevuorii</italic></bold> Niskanen & Liimat., <italic>sp. nov</italic>. — MycoBank MB805865; <xref ref-type="fig" rid="F2">Fig. 2</xref>c, <xref ref-type="fig" rid="F3">3</xref>c</p><p><italic>Etymology</italic>. The species is named in honour of Ilkka Kytövuori, a mycologist from Finland.</p><p><italic>Type</italic>. F<sc>INLAND</sc>, Koillismaa, Kuusamo, Oulanka, Ampumavaara, S slope, old, grass-herb <italic>Picea abies</italic> forest with some <italic>Betula</italic>,<italic> Pinus sylvestris</italic> and <italic>Populus tremula</italic>, on calcareous soil, 17 Sept. 2005, <italic>T. Niskanen & K. Liimatainen</italic> 05-158, H6029355 (holotype H; isotype NY). GenBank KF732529.</p><p><italic>Pileus</italic> 6–9 cm broad, hemispherical to convex, then expanded, finely innately fibrillose, yellow brown to brown, with hygrophanous streaks. <italic>Lamellae</italic> emarginate, almost crowded to medium spaced, at first pale brownish grey, becoming more brown with age. <italic>Stipe</italic> 6–9 cm long, 1.2–1.5 cm thick at apex, 2–2.5 cm wide at base, with fairly narrow, marginate bulb, at first white, becoming pale brownish yellow with age. <italic>Context</italic> whitish to pale yellow. <italic>Odour</italic> indistinct. <italic>KOH reactions</italic>: in pileipellis brown (no reddish tints); in context, mycelium and bulb margin negative. <italic>Exsiccata</italic>: pileus dull, dark red brown overall, stipe almost con-colorous with pileus.</p><p>In MLZ: <italic>Spores</italic>: 7.5–<italic>8.5</italic>–9.5 × 5.0–<italic>5.3</italic>–5.4 μm, Q = 1.54–<italic>1.61</italic>–1.72 (1 specimen, 60 spores, <xref ref-type="fig" rid="F3">Fig. 3</xref>c), amygdaloid-ellipsoid, very strongly, separately, ± sharply verrucose (<italic>C. porphyropus</italic> like but more dextrinoid), slightly to moderately dextrinoid. <italic>Basidia</italic>: 24–32 × 7.5–9 μm (20 basidia), 4-spored, clavate. <italic>Lamellar trama hyphae</italic>: pale pellucid yellowish, guttulate, smooth. <italic>Stipe apex hyphae</italic> colourless to pale straw-coloured, smooth, guttulate. <italic>Pileipellis</italic>: epicutis in overall view orange, (very) weakly gelatinous, hyphae 5–10 μm wide, finely to strongly spirally incrusted, many of the uppermost ones abundantly with intercellular, unevenly distributed, orange red granules, granule mounds and concretions. Large-celled hypoderm present, yellowish and with scanty small orange granules in the upper part, lower down colourless.</p><p>Ecology & Distribution — In coniferous forests, on calcareous soil. Fruits in autumn.</p><p>Notes — <italic>Cortinarius kytoevuorii</italic> is reminiscent of <italic>C. glaucopus</italic> but is more slender, has a yellow brown to brown pileus, and lacks bluish tints in basidiomata. It is most easily recognised by the orange red granules and concretions in the uppermost hyphae of the pileipellis when mounted in MLZ. The sister species <italic>C. subrugulosus</italic> Bidaud & Armada has a more southern distribution. The northernmost known collection is from Sweden, Öland under <italic>Fagus</italic> (<italic>Kytövuori</italic> 98-2578 (H)). Furthermore, the spores are shorter (7.3–<italic>7.9</italic>–8.6 × 5.0–<italic>5.3</italic>–5.7 μm), relatively broader (Qav. = 1.5), less verrucose, and more strongly dextrinoid, and in the pileipellis small yellow to blood-red guttules are seen in MLZ. In our phylogenetic analysis <italic>C. kytoevuorii</italic> formed a well-supported clade (1.00 PP) with <italic>C. subrugulosus</italic>, but further relationships were not solved. The difference in ITS regions between the two sister species is 11 substitutions and indel positions.</p><p><bold><italic>Cortinarius ochribubalinus</italic></bold> Kytöv., Liimat. & Niskanen, <italic>sp. nov.</italic> — MycoBank MB805866; <xref ref-type="fig" rid="F3">Fig. 3</xref>d</p><p><italic>Etymology</italic>. The name refers to the colour of the basidiomata.</p><p><italic>Type</italic>. F<sc>INLAND</sc>, Uusimaa, Espoo, Nuuksio, the open-air territory of Pirttimäki, between the main road and the lake, opposite the parking area, fairly rich grass-herb forest with <italic>Populus tremula</italic>,<italic> Betula</italic>,<italic> Alnus incana</italic>,<italic> Quercus</italic>,<italic> Corylus</italic>,<italic> Prunus padus</italic>,<italic> Salix</italic> spp., and some old pines and young spruce, 2 Sept. 1993, <italic>I. Kytövuori</italic> 93-641, H6032734 (holotype H; isotype NY). GenBank KF732530.</p><p><italic>Pileus</italic> 5–8 cm broad, convex, soon plano-convex, sometimes with a broad umbo, very finely fibrillose, centre ochraceous, whitish towards margin. <italic>Lamellae</italic> emarginate, medium spaced, at first very pale brownish grey, later pale brown. <italic>Stipe</italic> 6–10 cm long, 0.8–1.3 cm thick at apex, 1.5–2 cm at base, clavate, at first white, becoming pale brownish yellow with age. <italic>Universal veil</italic> white, on pileus margin thin, forming thin belts on the stipe. <italic>Context</italic> white. <italic>Odour</italic> pleasant. <italic>Exsiccata</italic>: pileus warm yellowish brownish at centre, pale leather-coloured to almost whitish at margin, stipe concolorous with the pileus margin.</p><p>In MLZ: <italic>Spores</italic> 12.5–<italic>13.4</italic>–14.3 × 7.5–<italic>7.9</italic>–8.2 μm, Q = 1.57–<italic>1.69</italic>–1.84 (1 specimen, 60 spores, <xref ref-type="fig" rid="F3">Fig. 3</xref>d), amygdaloid, strongly verrucose, most strongly so at the apex, very much like those in <italic>C. riederi</italic> group or <italic>C. violaceus</italic>, moderately dextrinoid. <italic>Basidia</italic> 39–48 × 9–12 μm (20 basidia), light sand brown, with large to small blood black granules. <italic>Lamellar trama hyphae</italic> with blood black substance and/or same-coloured granules, dark chips almost lacking. <italic>Stipe apex hyphae</italic> yellow, with colourless guttules, coloured granules lacking, but outermost hyphae sand brown to somewhat redder, with few to abundant blood red, small granules. <italic>Velum/Cortina hyphae</italic> sand brown to somewhat redder, with few to abundant blood red, small granules. <italic>Pileipellis</italic>: epicutis somewhat gelatinous, hyphae 3–10 μm wide, near the surface ochraceous brown, finely, densely incrusted, mostly not granulose, lower down strongly granulose with small to large blackish brown granules in mounds and long, sausage-shaped clusters and concretions. Hypoderm well developed, red brown, hyphae mostly granulose in the upper part.</p><p>Ecology & Distribution — With deciduous trees, possible hosts <italic>Populus</italic>,<italic> Corylus</italic> or <italic>Quercus</italic>. Known from South Finland. No sequences of this species exist in public databases.</p><p>Notes — Based on our molecular studies <italic>C. ochribubalinus</italic> does not have any known, close relatives. It holds an isolated position in the phylogenetic tree and differs in ITS regions by more than 20 substitutions and indel positions from the closest species <italic>C. patrickensis</italic> (M.M. Moser) Niskanen, Liimat., Kytöv., Bojantchev & Ammirati. Morphologically, it is reminiscent of species in /Arguti. The spores are large and similar to those of <italic>C. fraudulosus</italic> and <italic>C. subfraudulosus</italic>, but more strongly verrucose. The smell of the lamellae is also different, pleasant in <italic>C. ochrobubalinus</italic> and often earthy-raphanoid in <italic>C. fraudulosus</italic> and <italic>C. subfraudulosus</italic>. In addition, the universal veil is sparse where as <italic>C. fraudulosus</italic> and <italic>C. subfraudulosus </italic>have more abundant sometimes even floccose universal veil remnants on stipe.</p></sec></sec><sec id="s5"><title>/ARGUTI</title><p><bold><italic>Cortinarius patrickensis</italic></bold> (M.M. Moser) Niskanen, Liimat.,</p><p>Kytöv., Bojantchev & Ammirati, <italic>comb. nov.</italic> — MycoBank MB805883</p><p><italic>Basionym</italic>. <italic>Cortinarius fraudulosus </italic>var.<italic> patrickensis</italic> M.M. Moser, Mycotaxon 74, 1: 10. 2000.</p><p><italic>Type</italic>. USA, California, Humboldt Co., Patrick’s Point State Park prope Trinidad, in coniferous forest (<italic>Picea sitchensis</italic>, <italic>Pseudotsuga menziesii</italic>), 25 Nov. 1995, <italic>M. Moser</italic> 95/617 (holotype IB). GenBank KF732307.</p><p><italic>Other specimens examined</italic>. S<sc>WEDEN</sc>, Medelpad, Borgsjö, Julåsen, SE sloping, partly paludified, rich grass-herb-spruce forest with some pines and birches, 15 Sept. 1995, <italic>I. Kytövuori</italic> 95-1400, H7019584 (H). – USA, Six Rivers Nat’l forest, off Hwy 299, 1004 m asl, mixed forest (<italic>Notholithocarpus densiflorus</italic>, <italic>Quercus kelloggii</italic>, <italic>Pinus</italic> spp., <italic>Pseudotsuga menziesii</italic>, etc.), 20 Nov. 2010, <italic>D. Bojantchev</italic> DBB39406 (UC).</p><p>Notes — <italic>Cortinarius patrickensis</italic> is a typical member of /Arguti, it has a whitish to pale brown basidiomata and large spores, and the placement is also supported by molecular data. It was first described as a variety of <italic>C. fraudulosus</italic> but based on our genetic and morphological data it should be treated as a species. In our phylogenetic analysis <italic>C. patrickensis</italic> grouped together with <italic>C. rosargutus</italic> Chevassut & Rob. Henry (0.86 PP). Also with the pairwise comparison the closest species is <italic>C. rosargutus</italic> from which it differs by 5 substitutions and indel positions in ITS regions. Both taxa, <italic>C. patrickensis</italic> and <italic>C. rosargutus</italic>, include several identical or almost identical sequences and have a clear barcoding gap (no overlap between intra- and interspecific variation). A description of the species is provided in <xref rid="R65" ref-type="bibr">Moser & Ammirati (2000)</xref>. Typical for the species are amygdaloid, large spores 12.2–<italic>13.3</italic>–14.7 × 7.0–<italic>7.7</italic>–8.4 μm, Q = 1.62–<italic>1.73</italic>–1.84, and green apple-like to strong green corn-like odour. The spores are narrower than those of <italic>C. rosargutus</italic> (Qav. = 1.60). <italic>Cortinarius patrickensis</italic> grows in coniferous forests, often on calcareous soil. It was previously only known from California, USA but is here reported for the first time from Europe in Sweden. Species might be occasional in suitable habitats, at least in Europe and North America, but has most likely been misidentified as either <italic>C. fraudulosus</italic> or <italic>C. rosargutus</italic>.</p><p><bold><italic>Cortinarius subfraudulosus</italic></bold> Kytöv., Liimat. & Niskanen, <italic>sp. </italic><italic>nov</italic>. — MycoBank MB805867; <xref ref-type="fig" rid="F3">Fig. 3</xref>e</p><p><italic>Etymology</italic>. The name refers to the affinity of <italic>C. fraudulosus</italic>.</p><p><italic>Type</italic>. N<sc>ORWAY</sc>, Oppland, Lunner, Skøyenåsen, <italic>Picea abies</italic> forest with <italic>Corylus</italic> on calcareous soil, 3 Sept. 2011, <italic>I. Kytövuori</italic> 11-006 (holotype H; isotype NY). GenBank KF732564.</p><p>Illustrations — <xref rid="R16" ref-type="bibr">Brandrud et al. (1990</xref>: pl. A07 as <italic>C. argutus</italic> subsp. <italic>fraudulosus</italic>), <xref rid="R1" ref-type="bibr">Abarenkov et al. (2010</xref>: photo under the accession no. UDB011261).</p><p><italic>Pileus</italic> 4–10 cm broad, hemispherical to convex, then plano-convex, finely fibrillose, white to ochraceous white when young, with age becoming pale brownish. <italic>Lamellae</italic> emarginate, medium spaced to almost distant, first white to very pale brownish grey, later pale brown. <italic>Stipe</italic> 5–10 cm long, 1–2 cm thick at apex, 1.5–3 cm wide at base, clavate to slightly bulbous, sometimes slightly rooting, whitish, becoming brownish with handling and with age. <italic>Universal veil</italic> white, forming distinct girdles on stipe, sometimes floccose. <italic>Context</italic> white, but becomes very slowly black when bruised. <italic>Odour</italic> weak, a combination of earthy and raphanoid. <italic>Exsiccata</italic>: pileus dirty greyish to yellow brown at the centre, whitish to yellowish brown towards the margin, stipe greyish white to pale brown.</p><p>In MLZ: <italic>Spores</italic> 12.0–<italic>13.6</italic>–15.0 × 7.0–<italic>7.8</italic>–8.4 μm, av. = 12.5–14.4 × 7.6–8.2 μm, Q = 1.57–<italic>1.72</italic>–1.88, Qav. = 1.64–1.75 (4 specimens, 100 spores, <xref ref-type="fig" rid="F3">Fig. 3</xref>e), amygdaloid (to weakly citriform), with a narrow apex, moderately to fairly strongly verrucose, warts anastomosing, not high, moderately dextrinoid. <italic>Basidia</italic> 38–55 × 10–12 μm (2 specimens, 60 basidia), almost colourless to pale sand brownish, clear or with few small, brown granules. <italic>Lamellar trama hyphae</italic> full of dark granules or chips or blood black particles. <italic>Stipe apex hyphae</italic>yellow to reddish brown, outermost ones with abundant large red brown to blood blackish granule masses or opaque particles. <italic>Velum/Cortina hyphae</italic> with abundant large red brown to blood blackish granule masses or opaque particles. <italic>Pileipellis</italic>: epicutis fairly weakly gelatinous, hyphae at the surface 3–10 μm wide, finely incrusted, ochraceous brownish, mostly not granulose, lower down somewhat granulose or not. In the transition between epicutis and hypoderm up to 15 μm wide, dark red brown, strongly incrusted hyphae with large, blackish granula mounds, intracellular concretions and/or red brown, 3–15 μm wide extracellular pigment mounds on the hyphae, with few evenly distributed small granules. Hypoderm well developed, red brownish to very dark red brown, somewhat granulose or not.</p><p>Ecology & Distribution — In hemiboreal and southern boreal coniferous forests, calcicolous. Known from Fennoscandia and Estonia. Fruiting in autumn, in September.</p><p><italic>Other specimens examined</italic>. E<sc>STONIA</sc>, Hiiumaa, Kõrgessaare, Kõpu, old spruce forest with <italic>Pinus</italic>, <italic>Betula</italic>,<italic> Quercus</italic>,<italic> Populus tremula</italic> and <italic>Corylus</italic>, on calcareous soil, 15 Sept. 2001, <italic>T. Niskanen & I. Kytövuori</italic> (H); Pühalepa, Kallaste pank, spruce forest with <italic>Pinus</italic>,<italic> Betula</italic>,<italic> Populus tremula</italic> and <italic>Corylus</italic>, on calcareous soil, 23 Sept. 2008, <italic>J. Vesterholt & J. Vauras</italic> 26655F (TUR-A). – S<sc>WEDEN</sc>, Östergötland, Väversunda parish, E sloping, very rich spruce grass-herb forest with hardwood bushes, 26 Sept. 1988, <italic>I. Kytövuori</italic> 88-2016 (H); Uppland, Börstils sn, Marieberg, Täpporna, in rich spruce forest, 11 Sept. 1986, <italic>T.E. Brandrud et al.</italic> CFP481, F44801 (S); Södermanland, Mörkö parish, Oaxen, dryish spruce grass-herb forest with <italic>Pinus</italic>,<italic> Betula</italic>,<italic> Populus tremula</italic> and <italic>Salix</italic> spp., on calcareous soil, 27 Sept. 1998, <italic>I. Kytövuori</italic> 98-2244 (H), 98-2245 (H).</p><p><italic>Additional specimen</italic>. E<sc>STONIA</sc>, Saare, Kihelkonna, Mäebe, in coniferous forest, 21 Sept. 2009, <italic>V. Liiv</italic> TU106607, UNITE No. UDB011261 as <italic>C. fraudulosus</italic> (TU(M)).</p><p>Notes — The species has earlier been called <italic>C. fraudulosus</italic> in northern Europe. <italic>Cortinarius fraudulosus</italic>, however, has been described by <xref rid="R21" ref-type="bibr">Britzelmayr (1885)</xref> from Siebentischwald, Bavaria, Germany. In our phylogenetic tree the northern material and the more southern European material formed two separate, well-supported clades which differ from one another by 9 substitutions and indel positions in their ITS regions. The sequences of <italic>C. subfraudulosus</italic> have one base polymorphisms and the maximum pairwise distance is zero. Therefore, we here describe the northern species as new.</p></sec><sec id="s6"><title>/CALOCHROI & FULVI</title><p><bold><italic>Cortinarius flavipallens</italic></bold> Kytöv., Liimat. & Niskanen, <italic>sp. nov.</italic> — MycoBank MB805868; <xref ref-type="fig" rid="F2">Fig. 2</xref>d, <xref ref-type="fig" rid="F4">4</xref>a</p><p><italic>Etymology</italic>. The name refers to the colour of the pileus.</p><p><italic>Type</italic>. F<sc>INLAND</sc>, Kainuu, Kajaani, Hietalahti, Torakangas NNW of Korpitaipale, by the power line, fairly damp grass-herb-spruce forest with <italic>Pinus</italic>,<italic> Betula</italic> and <italic>Populus tremula</italic>, on calcareous soil, 13 Sept. 2008, <italic>I. Kytövuori</italic> 08-1729, H6032745 (holotype H; isotype NY). GenBank KF732554.</p><p><italic>Pileus</italic> 4–8 cm broad, hemispherical to convex, soon expanded, pale ochraceous to pale brown. <italic>Lamellae</italic> emarginate, crowded, pale grey to pale greyish brown. <italic>Stipe</italic> 4–6 cm long, 1–1.5 cm thick at apex, 2–2.5 cm wide at base, with a marginate bulb, white. <italic>Universal veil</italic> whitish to pale brown at bulb margin. <italic>Context</italic> white. <italic>Odour</italic> indistinct. <italic>KOH reactions</italic> on pileus and bulb margin red, on basal tomentum and rhizomorphs slowly and/or weakly vinaceous pink. <italic>Exsiccata</italic>: pileus pale ochre to ochre brown, darkest at the centre, paler towards the margin, stipe greyish white to pale brown.</p><p>In MLZ: <italic>Spores</italic> 9.5–<italic>10.7</italic>–11.6 × 5.4–<italic>6.0</italic>–6.3 μm, av. = 10.3–11.0 × 5.9–6.1, Q = 1.65–<italic>1.77</italic>–1.85, Qav. = 1.73–1.80 (3 specimens, 120 spores, <xref ref-type="fig" rid="F4">Fig. 4</xref>a), amygdaloid-fusoid to weakly citriform, with a shallow suprahilar depression and mostly a somewhat blunt apex, moderately verrucose, warts anastomosing or not, slightly to moderately dextrinoid. <italic>Basidia</italic> 26–38 × 8–10 μm (60 basidia), 4-spored, clavate, very pale yellowish, clear, few slightly granulose-guttulate. <italic>Lamellar trama hyphae</italic> pale yellowish, smooth, sometimes guttulate, some colourless crystals solitary or in roundish masses, not in branch-like fascicles. <italic>Pileipellis</italic>: epicutis strongly gelatinous, hyphae on the surface 3–5 μm wide, very pale ochre, smooth to very finely, densely, spirally incrusted, walls distinctly visible, also some hyphae with yellow, foamy contents. Lower down equally wide to slightly wider, strongly incrusted hyphae, often in bundles. Hypoderm absent. In KOH the gelatinized hyphae pale vinaceous pink.</p><p>Ecology & Distribution — In boreal <italic>Picea abies</italic> dominated forests on calcareous soil. So far only known from Finland but might be widely distributed, since an ITS sequence (FJ039640) from a specimen collected in British Columbia in western Canada differed only by 4 substitutions and indel positions from Finnish material and might be conspecific.</p><p><italic>Other specimens examined</italic>. F<sc>INLAND</sc>, Laatokan Karjala, Parikkala, Vaaranperä, Soininmäki, S part, Soininjoki, fairly old, SE sloping spruce forest with some <italic>Pinus</italic>,<italic> Betula</italic> and <italic>Populus</italic><italic>tremula</italic>, 17 Sept. 2009, <italic>A. Ahola</italic> H6032393 (H); Perä-Pohjanmaa, Tervola, Peura, Raemäki, in <italic>Picea</italic> dominated forest on calcareous soil, 5 Sept. 2011, <italic>T. Kekki</italic> 368 (H).</p><p>Notes — <italic>Cortinarius flavipallens</italic> with its pale ochraceous colours is in appearance between <italic>C. metarius</italic> Kauffman (= <italic>C. barbarorum</italic> Bidaud, Moënne-Locc. & Reumaux) and <italic>C. caesiocinctus</italic> Kühner, without the bright yellow pileus of the former or the bluish grey one of the latter. In addition, the KOH reaction in the mycelium is slower and weaker than in the former two species. <italic>Cortinarius piceae</italic> Frøslev, T.S. Jeppesen & Brandrud and <italic>C. corrosus</italic> Fr. differ from <italic>C. flavipallens</italic> by their negative KOH reaction in the basal mycelium. The ITS sequences of <italic>C. flavipallens</italic> were identical and it formed a well-supported clade in our phylogenetic analysis. However, the relationships with other species of /Calochroi p.p. were not resolved.</p><p><bold><italic>Cortinarius luteicolor</italic></bold> (A.H. Sm.) Ammirati, Bojantchev, Niskanen & Liimat., <italic>stat. nov. & nom. nov. </italic>— MycoBank MB805896</p><p><italic>Etymology</italic>. The name refers to the yellowish colours of the pileus and lamellae.</p><p><italic>Basionym</italic>. <italic>Cortinarius orichalceus </italic>var.<italic> olympianus </italic>f.<italic> luteifolius </italic>A.H. Sm., Lloydia 7: 185. 1944.</p><p><italic>Type</italic>. USA, Washington, Olympic Mts, near Lake Angeles, 19 Sept. 1941, <italic>A.H. Smith</italic> 16970, barcode 10389 (holotype MICH). GenBank KF732368.</p><p><italic>Other specimens examined</italic>. USA, California, Sierra Nevada, Yosemite Nat’l Park, Hwy 120, 2433 m asl, <italic>Pinus contorta</italic>, <italic>Pseudotsuga menziesii</italic>, <italic>Abies concolor</italic>, <italic>A. magnifica</italic>, etc., 16 Nov. 2011, <italic>D. Bojantchev </italic>DBB46740 (UC), 14 Nov. 2010, <italic>D. Bojantchev</italic> DBB38211 (UC).</p><p><italic>Additional specimens</italic>. C<sc>ANADA</sc>, British Columbia, VMS13, GenBank FJ-717511 as ‘<italic>Cortinarius</italic> sp.’ (UBC). – USA, Oregon, Clackamas Co., Bull Run Watershed, <italic>Tsuga heterophylla</italic>,<italic> Pseudotsuga menziesii</italic>, 11 Nov. 1995, M.M. 95/500/ <italic>J.F. Ammirati</italic> 11701, GenBank EU057024 as ‘sp.’; Washington, Chelan Co., Chatter Creek, <italic>Tsuga heterophylla</italic>, <italic>Pseudotsuga menziesii</italic>, 6 Oct. 2007, <italic>J. Birkebak</italic> JMB10-06-2007-03 (UBC), GenBank HM068562 as ‘<italic>C.</italic><italic>cupreorufus</italic>’.</p><p>Notes — ITS sequence analysis shows <italic>C. luteicolor</italic> as a well-delimited species within clade /Laeticolores. Morphologically, it is not similar to any of the species we have studied. Initially, the species was described as a form of <italic>C.</italic><italic>orichalceus</italic> var. <italic>olympianus</italic> (= <italic>C. pseudocupreorufus </italic>(A.H. Sm.) Niskanen, Liimat. & Ammirati). In the ITS regions it, however, differs from <italic>C. pseudocupreorufus</italic> by more than 25 substitutions and indel positions. Furthermore, <xref rid="R89" ref-type="bibr">Smith (1944)</xref> noticed several differences among these taxa: “This form (= <italic>C. luteicolor</italic>) differs from the typical form (= <italic>C. pseudocupreorufus</italic>) in lacking the faint lilac tint in the apex of stipe, in the pileus not becoming dark vinaceous red but instead merely dull cinnamon brown when fresh, and in the brighter yellow lamellae. The pilei of the herbarium specimens are also paler, but the bulb is deep purplish as in typical material of the variety (= <italic>C. pseudocupreorufus</italic>)”. Based on morphology and molecular data we conclude that <italic>C. luteicolor</italic> should be treated as a species. Description of the species is presented in <xref rid="R89" ref-type="bibr">Smith (1944)</xref>. Typical for the species are at first rich dull yellow or yellow tinged pileus gradually becoming dull cinnamon, pale yellow gills, and subalmond-shaped spores (9–11.5 6–7 μm). <italic>Cortinarius luteicolor</italic> grows in coniferous forests (<italic>Pseudotsuga</italic>,<italic> Tsuga</italic>) and is currently known from Pacific northwest of North America, from California and Washington, USA, and British Columbia, Canada.</p><p><bold><italic>Cortinarius pseudocupreorufus</italic></bold> (A.H. Sm.) Niskanen, Liimat. & Ammirati, <italic>stat. nov. & nom. nov.</italic> — MycoBank MB805886</p><p><italic>Etymology</italic>. The name refers to the affinity to <italic>C. cupreorufus</italic>.</p><p><italic>Basionym</italic>. <italic>Cortinarius orichalceus </italic>var.<italic> olympianus</italic> A.H. Sm., Lloydia 7, 3: 184. 1944.</p><p><italic>Type</italic>. USA, Washington, Olympic National Park, Olympic Hot Springs, under conifers, 2 Oct. 1941, <italic>A.H. Smith</italic> 17513, barcode 10390 (holotype MICH). GenBank KF732367.</p><p>Notes — In our phylogenetic analysis <italic>C. pseudocupreorufus</italic> is placed as a sister species of <italic>C. cupreorufus</italic> Brandrud from which it differs by 10 substitutions and indel positions in ITS regions. The spores of <italic>C. pseudocupreorufus</italic> are 9–11 × 5–6.5 μm and somewhat amygdaloid, while the spores of <italic>C. cupreorufus</italic> are broader, 10–11.5 × 6.5–7.5 μm, and amygdaloid-citriform. Based on morphology and molecular data we conclude that <italic>C. pseudocupreorufus</italic> should be treated as a species. Description of the species is presented in <xref rid="R89" ref-type="bibr">Smith (1944)</xref>. The species is so far only known from the type locality.</p></sec><sec id="s7"><title>/CYANITES</title><p><bold><italic>Cortinarius boreicyanites</italic></bold> Kytöv., Liimat., Niskanen & A.F.S. Taylor, <italic>sp. nov</italic>. — MycoBank MB805870; <xref ref-type="fig" rid="F4">Fig. 4</xref>b</p><p><italic>Etymology</italic>. The name refers to the affinity with <italic>C. cyanites</italic> and distribution in boreal zone.</p><p><italic>Holotype</italic>. S<sc>WEDEN</sc>, Jämtland, Ragunda sn, Ragunda, Böle, at the riverside, in birch forest on rich soil (<italic>Betula</italic>,<italic> Picea</italic>), 24 July 1990, <italic>Brandrud et al.</italic> CFP931 (S). GenBank KF732296.</p><p><italic>Pileus</italic> 4–10 cm broad, hemispherical to convex, then expanded, distinctly innately fibrillose, bluish grey when young, later pale greyish brown. <italic>Lamellae</italic> emarginate, almost crowded, greyish blue when young, later brownish violet. <italic>Stipe</italic> 5–10 cm long, 1–2 cm thick at apex, 2.5–4 cm wide at base, clavate to bulbose, greyish blue. <italic>Universal veil</italic> pale greyish brown, abundant, forming girdles on stipe. <italic>Context</italic> violet when young, later in pileus and bulb white to brownish white, marbled hygrophanous, turning vinaceous red on exposure. <italic>Odour</italic> indistinct. <italic>Exsiccata</italic>: pileus reddish brown at the centre, greyish brown at the margin, stipe pale bluish greyish, brown at the base.</p><p>In MLZ: <italic>Spores</italic> 9.1–<italic>9.8</italic>–10.4 × 5.4–<italic>5.8</italic>–6.3 μm, av. = 9.7–9.9 × 5.6–5.9 μm, Q = 1.58–<italic>1.69</italic>–1.80, Qav. = 1.66–1.73 (3 specimens, 120 spores, <xref ref-type="fig" rid="F4">Fig. 4</xref>b), amygdaloid, with a blunt apex, fairly finely to moderately verrucose, many with few small golden yellow guttules, slightly dextrinoid. <italic>Basidia</italic> 36–45 × 7–9 μm (60 basidia), 4-spored, narrowly clavate, with a long, narrow pedicel, with blood red guttules, commonly with yellow foamy contents. <italic>Lamellar trama hyphae</italic> 4–10 μm wide, colourless to yellowish brown, smooth, with abundant small to large worm-like blood red guttules. <italic>Stipe apex hyphae</italic> 3–8 μm wide, pale yellowish, smooth, with small to large blood red guttules, the outermost ones c. 3 μm wide, ochraceous brown, mostly not guttulate. <italic>Pileipellis</italic>: epicutis very weakly gelatinous, uppermost hyphae 4–8(–13) μm wide, ochraceous brown, very thinly spot-like incrusted, not or weakly guttulate with small, golden yellow guttules, lower hyphae up to 10 μm wide, brown, smooth, with abundant small to large worm-like, blood red guttules. Hypoderm not developed.</p><p>Ecology & Distribution — In boreal mixed forests of <italic>Picea</italic>,<italic> Betula</italic> and <italic>Populus </italic>in northern Europe. In Scotland also collected with <italic>Helianthemum</italic>. Fruiting in autumn.</p><p><italic>Other specimens examined</italic>. F<sc>INLAND</sc>, Pohjois-Savo, Kuopio, Vehmersalmi, submesic, mixed forest (<italic>Betula</italic>,<italic> Picea</italic>,<italic> Pinus</italic>), in grassy places, 1 Aug. 2003, <italic>T. Niskanen</italic><italic>et al.</italic> 03-112 (H). – G<sc>REAT</sc> B<sc>RITAIN</sc>, Scotland, Grampian, Braemar, Morrone Wood, with <italic>Helianthemum</italic> on rich calcareous soil, 8 Aug. 2010, <italic>A. Taylor</italic> 2010203 (UPS).</p><p>Notes — ITS sequence analysis shows <italic>C. boreicyanites</italic> as a well-delimited species within clade /Cyanites (<xref ref-type="fig" rid="F1">Fig. 1</xref>). The ITS sequences of the species are identical and differ by more than 30 substitutions and indel positions from those of <italic>C. cyanites</italic> Fr. and <italic>C. violaceorubens</italic> Moënne-Locc. & Reumaux. Typical for <italic>C. boreicyanites</italic> are a pale greyish brown pileus, exsiccata with reddish brown pileus at the centre, greyish brown at the margin, and spores on average 9.7–9.9 × 5.6–5.9 μm. The species is so far only known from the middle boreal zone of Sweden, Finland and Scotland. <italic>Cortinarius cyanites</italic> has a more southern distribution extending from the hemiboreal zone of Finland and Sweden to France. In addition, the pileus is more bluish. <italic>Cortinarius violaceorubens</italic> has a more brownish pileus, dark dirty violaceous brown exsiccata, and larger spores (av. 9.9–11.0 × 6.3–6.6 μm). It grows in <italic>Picea</italic> dominated forests, often on rich soil.</p></sec><sec id="s8"><title>/DIONYSAE s.l</title><p><bold><italic>Cortinarius boreidionysae</italic></bold> Kytöv., Liimat., Niskanen & Dima, <italic>sp. nov</italic>. — MycoBank MB805894; <xref ref-type="fig" rid="F2">Fig. 2</xref>e, <xref ref-type="fig" rid="F4">4</xref>c</p><p><italic>Etymology</italic>. The name refers to the affinity with <italic>C. dionysae</italic> and the northern distribution.</p><p><italic>Type</italic>. F<sc>INLAND</sc>, Perä-Pohjanmaa, Tervola, Peura, Raemäki, grass-herb-spruce forest with spring-fed depressions, calcareous soil, 11 Sept. 1997, <italic>I. Kytövuori</italic> 97-1220 (holotype H; isotype NY). GenBank KF732488.</p><p><italic>Pileus</italic> 5–10 cm broad, hemispherical to convex, then expanded, glutinous, innately fibrillose, mustard brown to cocoa brown with a somewhat silky shining centre and olive yellowish tint on the margin when young. <italic>Lamellae</italic> emarginate, crowded, violaceous when young, later violet grey to pale brownish grey. <italic>Stipe</italic> 3–9 cm long, 1–1.8 cm thick at apex, 2–2.5 cm wide at base, with a marginate bulb, when young pale violaceous, becoming yellowish. <italic>Universal veil</italic> yellow at bulb margin. <italic>Context</italic> in pileus white, in stipe violaceous, in bulb at first whitish, later brownish yellowish. <italic>Odour</italic> faintly farinaceous. <italic>Exsiccata</italic>: pileus uniformly orange brown, sometimes with a faint greyish tint, stipe somewhat paler.</p><p>In MLZ: <italic>Spores</italic> 8.4–<italic>9.3</italic>–10.2 × 5.2–<italic>5.8</italic>–6.1 μm, av. = 8.9–9.5 × 5.3–5.8 μm, Q = 1.54–<italic>1.67</italic>–1.80, Qav. = 1.63–1.67 (5 specimens, 160 spores, <xref ref-type="fig" rid="F4">Fig. 4</xref>c), strongly citriform, strongly beaked, moderately, sharply verrucose, often with small coloured guttules, faintly to moderately dextrinoid. <italic>Basidia</italic> 23–32 × 7–9 μm (60 basidia), 4-spored, clavate, some with brownish yellow, foamy contents. <italic>Lamellar trama hyphae</italic> yellow, granulose-guttulate, blood red guttules absent. <italic>Stipe apex hyphae</italic> yellowish to yellow, smooth, with golden yellow small drops and large, blood red, worm-like guttules abundant in the outermost hyphae. <italic>Pileipellis</italic>: epicutis strongly gelatinous, uppermost hyphae 3–11 μm wide, mostly densely, spirally incrusted with small to large blood red guttules, lower down with a thick layer of somewhat wider hyphae filled with small to large or very large, worm-like, foamy, blood red guttules. Hypoderm well developed, yellow to red brownish.</p><p>Ecology & Distribution — In northern boreal <italic>Picea abies</italic> dominated forests on calcareous soil, rare. Fruiting in autumn.</p><p><italic>Other specimens examined</italic>. F<sc>INLAND</sc>, Oulun Pohjanmaa, Kiiminki, Juuvansydänmaa, S part of Iso Juuvankangas, W of the lake Iso Juuvanjärvi, grass-herb-spruce forest with some <italic>Betula</italic>,<italic> Populus tremula</italic> and <italic>Pinus</italic>, on calcareous soil, 17 Aug. 2007, <italic>M. Toivonen & I. Kytövuori</italic> 07-245, H6000476 (H); Perä-Pohjanmaa, Tervola, Peura, Raemäki, grass-herb-spruce forest with spring-fed depressions, calcareous soil, 21 Aug. 1998, <italic>I. Kytövuori</italic> 98-1316, H6035751 (H), 21 Aug. 2007, <italic>M. Toivonen & I. Kytövuori</italic> 07-491, H6000724 (H); church village, Ala-Kirvesmaa, rich grass-herb-spruce forest with some <italic>Betula</italic>,<italic> Populus</italic> and <italic>Pinus</italic>, on calcareous soil, 24 Aug. 2007, <italic>I. Kytövuori</italic> 07-768, H6001001 (H); Petäjämaa, Kivimaa, 6 Sept. 2012, <italic>I. Kytövuori</italic> (H); Tornio, Korkiamaa, Runteli, rich grass-herb-spruce forest with hardwood bushes and some pines, slightly paludified depressions, calcareous soil, 12 Aug. 1995, <italic>I. Kytövuori</italic> 95-210, H6035732 (H), 10 Sept. 1997, <italic>I. Kytövuori</italic> 97-1170 (H), 20 Aug. 1998, <italic>I. Kytövuori</italic> 98-1279, H6035740 (H), 22 Aug. 2007, <italic>I. Kytövuori</italic> 07-626, H6000859 (H); Kuusamo, Oulanka National Park, Ampumavaara, mesic to damp, herb-rich <italic>Picea abies</italic> forest, with some <italic>Pinus</italic>,<italic> Populus</italic> and <italic>Betula</italic>, 19 Sept. 2002, <italic>T. Niskanen et al. </italic>02-876, H6033134 (H).</p><p>Notes — <italic>Cortinarius boreidionysae</italic> reminds one of <italic>C. olivaceodionysae </italic>A. Ortega, Vila & Fdez.-Brime, but the latter has paler, olive brown to yellowish brown pileus, somewhat larger, less distinctly and less regularly citriform spores, and a more southern distribution (up to the hemiboreal zone). The ITS sequences of <italic>C. boreidionysae</italic> are identical and it formed a well-supported clade in our phylogenetic analysis. It belongs to /Dionysae but further relationships were not solved. Based on ITS sequence comparison the closest species is <italic>C. olivaceodionysae </italic>from which it differs in ITS regions by 9 substitutions and indel positions, although two of them include intragenomic base polymorphisms.</p><p><italic>Cortinarius olivaceodionysae</italic> has been called <italic>C. dionysae</italic> Rob. Henry in northern Europe. <italic>Cortinarius dionysae</italic>, however, has been described from France (<xref rid="R44" ref-type="bibr">Henry 1933</xref>) as a species with a distinctly farinaceous smell and collected under <italic>Fagus</italic> – both characters in contradiction with our material of <italic>C. olivaceodionysae</italic>. To confirm the identity of <italic>C. dionysae</italic> we tried to sequence the type material but unfortunately we did not succeed. In our phylogenetic analysis all the taxa with a strong farinaceous smell, <italic>C. dionysae</italic> sensu Frøslev & Garnica, <italic>C. dionysae</italic> var. <italic>avellanus</italic> Rob. Henry ex Bidaud & Carteret and <italic>C. palazonianus</italic> Vila, A. Ortega & Fdez.-Brime, formed a well-supported subclade (1.00 PP) inside the clade /Dionysae. Of these, <italic>C. dionysae</italic> sensu Frøslev & Garnica and <italic>C. dionysae</italic> var. <italic>avellanus</italic> are potential candidates for the real <italic>C. dionysae</italic>. The former is collected under <italic>Fagus</italic>, but is so far only recorded from Germany and the spores are smaller than given in the original description. The latter is found in mixed forest in France but the spores fit well to the description. Further studies with French material are needed to confirm the identity of <italic>C. dionysae</italic>.</p><p><bold><italic>Cortinarius cruentipellis</italic></bold> Kytöv., Liimat., Niskanen & Dima, <italic>sp. nov</italic>. — MycoBank MB805872; <xref ref-type="fig" rid="F2">Fig. 2</xref>f, <xref ref-type="fig" rid="F4">4</xref>d</p><p><italic>Etymology</italic>. The name refers to the large and abundant blood red drops/guttules in the pileipellis.</p><p>= <italic>Cortinarius luteoimmarginatus</italic> Rob. Henry sensu <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012)</xref>.</p><p>= <italic>Cortinarius polymorphus</italic> Rob. Henry s.auct. p.p.</p><p><italic>Type</italic>. S<sc>WEDEN</sc>, Öland, Långlöt, Åstad, Nitares hägn, grassy pasture with <italic>Corylus</italic> and <italic>Juniperus</italic>, 13 Sept. 2003, <italic>I. Kytövuori</italic>,<italic> K. Liimatainen & T. Niskanen</italic> 03-1451 (holotype H; isotype NY). GenBank KF732539.</p><p><italic>Pileus</italic> 3–7.5 cm broad, hemispherical to convex, then expanded, unevenly wavy, olivaceous yellowish brown at centre, olivaceous to ochraceous yellow towards margin. <italic>Lamellae</italic> emarginate, almost crowded, pale grey when young, later pale greyish brown. <italic>Stipe</italic> 3–5 cm long, 0.5–1.4 cm thick at apex, 1.5–2.5 cm wide at base, with a marginate bulb, when young whitish, becoming slightly yellow with age. <italic>Universal veil</italic> ochraceous yellow at bulb margin. <italic>Context</italic> whitish. <italic>Odour</italic> indistinct. <italic>Exsiccata</italic>: pileus orange brown to dirty red brown at the centre, yellowish brown at the margin, stipe concolorous with the centre.</p><p>In MLZ: <italic>Spores</italic> 9.5–<italic>10.4</italic>–11.6 × 5.7–<italic>6.1</italic>–6.6 μm, av. = 10.3–10.5 × 6.0–6.2 μm, Q= 1.59–<italic>1.71</italic>–1.85, Qav. = 1.69–1.73 (5 specimens, 160 spores, <xref ref-type="fig" rid="F4">Fig. 4</xref>d), often strongly citriform, beaked, moderately, sharply verrucose, fairly faintly to moderately dextrinoid. <italic>Basidia</italic> 23–34 × 7.5–9.5 μm (50 basidia), 4-spored, clavate, some with yellow foamy contents. <italic>Lamellar trama hyphae</italic> yellowish, many with greenish yellowish, oily guttules. Blood red guttules absent. <italic>Stipe apex hyphae</italic> pale yellowish to greyish brownish to reddish brownish, smooth to finely or more strongly incrusted, somewhat granulose-guttulate, large blood red and smaller golden yellow guttules present in the outermost hyphae. <italic>Pileipellis</italic>: epicutis distinctly gelatinous, uppermost hyphae 4–10 μm wide, thin-walled, finely densely, spirally incrusted, mostly not guttulate, below these a thick layer of smooth to somewhat incrusted hyphae with abundant very long sausage-like guttules with blood red foamy contents. Hypoderm present, yellow brownish.</p><p>Ecology & Distribution — Very rare in temperate to hemiboreal grass-herb forests with deciduous trees, mostly <italic>Corylus</italic> and <italic>Quercus</italic>, on calcareous soil. In addition, it occurs in half-open deciduous vegetation in wooded pastures and parks. In Northwest Europe it is known in Denmark, Norway, Sweden and Estonia. Fruits in autumn.</p><p><italic>Other specimens examined</italic>. E<sc>STONIA</sc>, Hiiumaa, Pühalepa, Sarve, dryish <italic>Corylus</italic> forest with <italic>Juniperus</italic>,<italic> Quercus</italic> and <italic>Populus</italic>, stony calcareous soil, 16 Sept. 2001, <italic>I. Kytövuori & T. Niskanen</italic> (Kytövuori 01-055, H). – N<sc>ORWAY</sc>. Oslo, Bygdøy, Dronginberget south, deciduous forest, 26 Sept. 2004, <italic>T.E. Brandrud</italic> (Niskanen F04-983) (H7017763); Akershus, Asker, Spirodden, rich calcareous forest, <italic>Corylus</italic>, 8 Sept. 2011, <italic>I. Kytövuori</italic> 11-008 (H). – S<sc>WEDEN</sc>, Öland, Algutsrum, Gråbor Fornborg, rich grass-herb forest with <italic>Corylus</italic>,<italic> Quercus</italic>,<italic> Populus tremula </italic>and<italic> Picea</italic>, on sedimentary limestone, 30 Sept. 1988, <italic>I. Kytövuori</italic> 98-2503 (H); Öland, Torslunda, 1.5 km S from Borg, <italic>Corylus</italic> forest with some <italic>Picea</italic>,<italic> Betula </italic>and<italic> Quercus</italic>, 12 Sept. 2003, <italic>T. Niskanen</italic> D03-1393 & <italic>I. Kytövuori & K. Liimatainen</italic>, H7018687 (H).</p><p>Notes — <italic>Cortinarius cruentipellis</italic> is a rather small, olivaceous tinted <italic>Phlegmacium</italic> of deciduous forests with citriform spores and strongly blood red pileipellis in MLZ. Of the other deciduous forest species with red MLZ reaction in pileus cuticle, <italic>C. gracilior</italic> (M.M. Moser) M.M. Moser is small and yellowish and has exsiccata with pale stramineous pileus and whitish stipe, <italic>C. leonicolor</italic> Reumaux (= <italic>C. anserinus</italic> (Velen.) Rob. Henry s.auct.) has much larger spores, and <italic>C. subdecolorans</italic> Langl. & Reumaux (= <italic>C. multiformium</italic> Cons. & Moënne-Locc.) pileipellis hyphae filled by mostly small guttules. Based on the ITS sequence analysis <italic>C. cruentipellis</italic> is a well-supported species (1.00 PP) belonging to the clade /Dionysae (0.82 PP). It has no close known relatives, and in the ITS regions it differs by more than 30 substitutions and indel positions from the closest species.</p><p><xref rid="R52" ref-type="bibr">Jeppesen et al. (2012)</xref> used the name <italic>C. luteoimmarginatus</italic> Rob. Henry for this species. In the original description of <italic>C. luteoimmarginatus</italic> (<xref rid="R45" ref-type="bibr">Henry 1939</xref>, as <italic>C. multiformis</italic> var. <italic>luteoimmarginatus</italic>) the species is mentioned to have a clavate stipe, like the one of <italic>C. cliduchus</italic>, and this is also illustrated by Henry in <xref rid="R10" ref-type="bibr">Bidaud et al. (2012)</xref> whereas our species has a marginate bulb (<xref rid="R52" ref-type="bibr">Jeppesen et al. 2012</xref>). In addition, the spores are 11–11.5 × 5.5–6.5 μm, generally longer than those of our species and not overlapping with the average size of the spores (10.4 × 6.1 μm) or with the measurements of <xref rid="R52" ref-type="bibr">Jeppesen et al. (2012)</xref> (10–11 × 5.5–6.5 μm). In the original description no type has been designated but later <xref rid="R49" ref-type="bibr">Henry (1985)</xref> suggested a type (no. 1006) and a heterotype (no. 1014) for the species. The type of <italic>C. luteoimmarginatus</italic>, however, was not located in Paris (PC). A collection numbered as 1014 was found, but it was labelled as <italic>C. privignofulvus</italic> and represented a species of subg. <italic>Telamonia</italic>. Since the original description of <italic>C. luteoimmarginatus</italic> does not fit to our species we here describe it as new.</p></sec><sec id="s9"><title>/GLAUCOPODES</title><p><bold><italic>Cortinarius subrubrovelatus</italic></bold> (Bidaud) Kytöv., Liimat., Niskanen & Dima, <italic>comb. nov</italic>. — MycoBank MB805884</p><p><italic>Basionym</italic>. <italic>Cortinarius glaucopus</italic> var. <italic>subrubrovelatus</italic> Bidaud, Atlas des Cortinaires 17, 2: 1237. 2008.</p><p><italic>Holotype</italic>. F<sc>RANCE</sc>, Ain, Farges, in mixed forest of <italic>Populus</italic>,<italic> Corylus</italic>,<italic> Fagus</italic> and <italic>Abies</italic>, 1 Nov. 1997,<italic> A. Bidaud</italic> 97-11-431 (PC). GenBank KF732317.</p><p><italic>Other specimens examined</italic>. F<sc>INLAND</sc>, Varsinais-Suomi, Lohja, Jalassaari, herb-rich <italic>Picea abies</italic> forest with some <italic>Corylus avellana</italic>, 19 Sept. 2004, <italic>I. Kytövuori & T. Niskanen</italic> 04-881, H6031330 (H); Paimio, Huso, Kalkkimäki, below the old limestone quarries, moist spruce grass-herb forest with hardwood bushes, 6 Oct. 1985, <italic>I. Kytövuori</italic> 85-1553 (H). – G<sc>ERMANY</sc>, Bayern, Oberjoch, conifer forest with <italic>Picea abies</italic>, 4 Oct. 2001, <italic>S. Garnica</italic> 4498 (TUB 011414), GenBank AY174787.</p><p><italic>Additional specimens</italic>. E<sc>STONIA</sc>, Saare, Lümanda, in garden, with spruce and pine, <italic>V. Liiv</italic> 2009-10-29 (as ‘<italic>C. pini</italic>’ TU106663/UDB011262).</p><p>Notes — ITS sequence analysis shows <italic>C. subrubrovelatus</italic> as a well-defined species (1.00 PP) in /Glaucopodes. Typical for <italic>C. subrubrovelatus</italic> is pale brown to red brown pileus, bluish lamellae, and small, relatively broad spores (7.0–<italic>7.9</italic>–8.8 × 4.5–<italic>5.0</italic>–5.4 μm, av. = 7.7–8.0 × 4.8–5.2 μm, Q = 1.44–<italic>1.56</italic>–1.70, Qav. = 1.52–1.61 (3 specimens, 180 spores)). From its closest relatives <italic>C. subfoetens</italic> and <italic>C. glaucopus</italic> it differs by 2 and 4 substitutions and/or indel positions. <italic>Cortinarius subfoetens</italic> has larger, more fusoid and narrower, and less dextrinoid (8.2–<italic>8.9</italic>–9.7 × 5.0–<italic>5.3</italic>–5.4 μm, Q = 1.58–<italic>1.69</italic>–1.82) spores, and the spores of <italic>C. glaucopus</italic> are relatively narrower as seen in the Q-values (7.3–<italic>8.1</italic>–9.1 × 4.5–<italic>5.0</italic>–5.4 μm, Q = 1.54–<italic>1.64</italic>–1.82). The latter also has somewhat more reddish brown colours deep in the pileipellis and less incrusted upper hyphae than those of <italic>C. subrubrovelatus</italic>. The differences in the ITS region between the three species are not large but since all three groups are represented by several collections and also have morphological differences we conclude that <italic>C. subrubrovelatus</italic> should be treated as a species. <italic>Cortinarius subrubrovelatus</italic> grows in coniferous forests on calcareous soil and is known from Central Europe and hemiboreal zone of northern Europe.</p></sec><sec id="s10"><title>/ELASTICI & PERCOMES</title><p><bold><italic>Cortinarius luteiaureus</italic></bold> Kytöv., Liimat. & Niskanen, <italic>sp. nov</italic>. — MycoBank MB805895; <xref ref-type="fig" rid="F4">Fig. 4</xref>e</p><p><italic>Etymology</italic>. The name refers to the colour of the pileus.</p><p><italic>Type</italic>. F<sc>INLAND</sc>, Oulun Pohjanmaa, Kiiminki, Juuvansydänmaa, S part of Iso Juuvankangas, W of the lake Iso Juuvanjärvi, grass-herb <italic>Picea abies</italic> forest with some <italic>Betula</italic>,<italic> Populus tremula</italic> and <italic>Pinus</italic>, on calcareous soil, 17 Aug. 2007, <italic>M. Toivonen & I. Kytövuori</italic> 07-247b, H6033617 (holotype H; isotype NY). GenBank KF732568.</p><p><italic>Pileus</italic> 4–7 cm broad, convex, then plano-convex, with a low and broad umbo, viscid to glutinous, not fibrillose, yellow to brownish yellow. <italic>Lamellae</italic> emarginate, almost crowded, greyish white, later very pale greyish brown. <italic>Stipe</italic> 5–10 cm long, 1–1.5 cm thick at apex, 1.5–2 cm wide at base, almost cylindrical with clavate to subbulbous base, white. <italic>Universal veil</italic> yellow, forming girdles on stipe. <italic>Context</italic> white. <italic>Odour</italic> not recorded. <italic>Exsiccata</italic>: pileus yellow brownish, stipe whitish.</p><p>In MLZ: <italic>Spores</italic> 9.7–<italic>10.6</italic>–11.6 × 5.7–<italic>6.1</italic>–6.6 μm, Q = 1.62–<italic>1.74</italic>–1.92 (1 specimen, 60 spores, <xref ref-type="fig" rid="F4">Fig. 4</xref>e), narrowly amygdaloid, with rounded apex, moderate to fairly strongly verrucose, moderately dextrinoid, often with dark red brown angular granules in the spore. <italic>Basidia</italic> 32–44 × 8–10 μm (20 basidia), 4-spored, clavate, sand brown, with fairly small granules and chips. <italic>Lamellar trama hyphae</italic> with moderate to very large sand brown to red brown granules. <italic>Stipe apex hyphae</italic> sand brown, densely granulose, outermost ones more orange or reddish, not granulose. <italic>Pileipellis</italic>: epicutis strongly gelatinous, uppermost hyphae 2–4 μm wide, ochraceous yellow to ochraceous brown, mostly not granulose, lower down 4–10 μm wide, full of small to very large dark red brown granules. Hypoderm absent.</p><p>Ecology & Distribution — In coniferous forest (<italic>Picea</italic>) on calcareous soil. Known from northern Finland, Oulun Pohjanmaa. No sequences of this species exist in public databases.</p><p>Notes — <italic>Cortinarius luteiaureus</italic> resembles somewhat a species which we assume could be called <italic>C. rhizophorus</italic> Bidaud & Cons. but we have not studied the type material of the species yet. <italic>Cortinarius</italic> cf. <italic>rhizophorus</italic> is larger, somewhat paler, has larger spores (10.2–<italic>11.3</italic>–12.5 × 6.1–<italic>6.5</italic>–7.0 μm) and occurs in temperate to hemiboreal broad-leaved and coniferous forests. <italic>Cortinarius varius</italic> (Schaeff.: Fr.) Fr. is also somewhat similar but has blue lamellae, stout, clavate stipe, wider spores, and hypoderm in the pileipellis. <italic>Cortinarius luteiaureus</italic> differs in ITS regions by more than 30 substitutions and indel positions from the closest species found with the BLAST. Based on our phylogenetic analysis it belongs to the large /Elastici & Percomes clade.</p></sec><sec id="s11"><title>/INFRACTI</title><p><bold><italic>Cortinarius infractiflavus</italic></bold> (M.M. Moser) Kytöv., Niskanen, Liimat., Bojantchev & Ammirati, <italic>stat. nov. & nom. nov.</italic> — MycoBank MB805887</p><p><italic>Etymology</italic>. The name refers to the affinity to <italic>C. infractus</italic> and the yellowish pileus.</p><p><italic>Basionym</italic>. <italic>Cortinarius infractus </italic>var.<italic> flavus</italic> M.M. Moser, Mycotaxon 72: 313. 1999.</p><p><italic>Type</italic>. USA, Wyoming, Shoshone Natl. Forest, prope Brooks Lake Lodge, in subalpine coniferous forest (<italic>Picea engelmanii</italic>,<italic> Abies lasiocarpa</italic>), 12 Aug. 1997, <italic>M. Moser</italic> 97/169 (holotype IB). GenBank KF732327.</p><p><italic>Other specimens examined</italic>. B<sc>ULGARIA</sc>, Pirin Mt, Bansko locality, 1600 m asl, under <italic>Picea abies</italic>, 7 Oct. 2009, <italic>D. Bojantchev</italic>, DBB19634 (UC). – F<sc>INLAND</sc>, Sompion Lappi, Pelkosenniemi, Suvanto, Kalkkivaara, half-open dry <italic>Pinus sylvestris</italic> forest with <italic>Picea</italic>, <italic>Juniperus</italic>,<italic> Betula</italic>,<italic> Populus </italic>and <italic>Alnus</italic>, dolomite rock, 26 Aug. 1992, <italic>I. Kytövuori</italic> 92-1109 (H).</p><p><italic>Additional specimen</italic>. C<sc>ANADA</sc>, British Columbia SMI286, GenBank FJ039612 (UBC).</p><p>Notes — <italic>Cortinarius infractiflavus</italic> formed a well-supported clade (1.00 PP) in our phylogenetic analysis. It belongs to sect. <italic>Infracti</italic> and differs from the sister species <italic>C. infractus</italic> by 9 substitutions and indel positions in ITS regions. Morphologically, it can be distinguished from <italic>C. infractus</italic> by the yellowish colours of the pileus, paler gills, a nearly mild taste and larger spores (7.5–<italic>8.2</italic>–9.1 × 6.3–<italic>6.7</italic>–7.0 μm, Q = 1.16–<italic>1.23</italic>–1.30; <italic>C. infractus</italic> 7.3–<italic>8.0</italic>–8.8 × 5.7–<italic>6.1</italic>–6.6 μm, Q= 1.23–<italic>1.30</italic>–1.38). <italic>Cortinarius infractiflavus</italic> grows in boreal and mountainous conifer forests (<italic>Picea</italic>,<italic> Abies</italic>) and is widespread occurring from Wyoming, western USA to Finland and Bulgaria. Full description of the species can be found from <xref rid="R64" ref-type="bibr">Moser & Ammirati (1999)</xref>.</p></sec><sec id="s12"><title>/MULTIFORMES</title><p><bold><italic>Cortinarius caesiolamellatus</italic></bold>(Bidaud) Kytöv., Liimat., Niskanen, Brandrud, Frøslev & A.F.S. Taylor, <italic>comb. nov</italic>. — MycoBank MB805885</p><p><italic>Basionym</italic>. <italic>Cortinarius rufoallutus</italic> var. <italic>caesiolamellatus</italic> Bidaud, Atlas des Cortinaires 16: 1095. 2006.</p><p><italic>Type</italic>. F<sc>RANCE</sc>, Ain, Bugey, col de Bérentin, in young <italic>Picea abies</italic> plantation, on calcareous soil, 3 Oct. 1993, <italic>A. Bidaud</italic> PML4905 (holotype PC). GenBank KF732414.</p><p>= <italic>Cortinarius multiformis</italic> var. <italic>caesiophyllus</italic> Moënne-Locc., Atlas des Cortinaires 16: 1095. 2006. — <italic>Type</italic>. <italic>P. Moënne-Loccoz</italic> PML882 (holotype PC), France, Savoie, Arith, under <italic>Picea</italic> on calcareous soil, 3 June 1988. Gen-Bank KF732351.</p><p>Illustrations — Bidaud et al. (2006: pl. 581, 587).</p><p><italic>Pileus</italic> 4–8 cm broad, hemispherical to plano-convex, sometimes radially rugulose towards the margin, viscid, red brown to ochraceous brown, rarely bluish brown, becoming paler ochraceous brown with age, often bi-coloured, outer part hygrophanous and darker (grey) brown. <italic>Lamellae</italic> emarginate, almost crowded, pale grey with a bluish tint when young, later pale brown. <italic>Stipe</italic> 4–8 cm long, 0.7–1.7 cm thick at apex, cylindrical, slender, 1.5–2.5 cm wide at base, mostly bulbous but the bulb often fairly small, fibrillose (not glossy like <italic>C. multiformis</italic>), whitish but often with a bluish tint at apex when very young, often becoming pale brown with age. <italic>Universal veil</italic> white, sparse, sometimes viscid (at bulb margin). <italic>Context</italic> in the pileus fairly thin, whitish, brownish below the pileus cuticle, in the stipe with a bluish tint or not when young. <italic>Odour</italic> faint to distinct of honey in the context of the stipe base. <italic>Exsiccata</italic>: pileus greyish to reddish brown, stipe very pale.</p><p>In MLZ: <italic>Spores</italic> 8.2–<italic>9.3</italic>–10.7 × 5.0–<italic>5.6</italic>–6.3 μm, av. = 8.5–9.9 × 5.2–5.9 μm, Q = 1.51–<italic>1.65</italic>–1.85, Qav. = 1.57–1.74 (7 specimens, 240 spores), ovoid-amygdaloid to narrowly ovoid-ellipsoid, with a fairly long, blunt, tapering apex, moderately to strongly verrucose, warts wide, anastomosing, slightly to fairly strongly dextrinoid. <italic>Basidia</italic> 25–34 × 7.5–9 μm (100 basidia), 4-spored, clavate, mostly with fairly wide base. Many basidia and subhymenium with yellowish, granulose/guttulate contents. <italic>Pileipellis</italic>: epicutis with a fairly thick, gelatinous layer with some 2–3 μm wide, erectentangled, smooth, obscure, colourless hyphae. Hypoderm present, fairly thin-walled, with pale brownish parietal pigment, some small brown spots present, large yellow to yellow brown spots absent, few spirally incrusted hyphae present deep in the pileipellis.</p><p>Ecology & Distribution — In montane to middle boreal, mesic coniferous forests, often on rich soil. In Europe apparently mainly/only under <italic>Picea</italic>, but in USA also found under <italic>Pinus</italic>. Known from Central and northern Europe, and Washington, USA, and considered occasional. The fruiting period is very long, once collected in June (France) and once in November (USA).</p><p><italic>Other specimens examined</italic>. E<sc>STONIA</sc>, Saaremaa, Mustjala, Võhma, dryish pine heath forest with some spruce, on calcareous soil, 16 Sept. 1993, <italic>I. Kytövuori</italic> 93-1336 (H). – F<sc>INLAND</sc>, Uusimaa, Kunnarla, Myllyoja, grass-herb forest with some pines and hardwood trees, 22 Sept. 1994, <italic>I. Kytövuori</italic> 94-852 (H); Pohjois-Savo, Kuopio, Vehmersalmi, Putroniemi, Roikanselän ranta-Mäkijärvi E, under <italic>Picea</italic>, 30 years old forest, 26 Sept. 2009, <italic>J. Ruotsalainen</italic> 8103F, H6011464 (H). – F<sc>RANCE</sc>, Oyonnax SE, Giron, Forêt de Champfromier, rich conifer forest with <italic>Abies alba</italic>,<italic> Picea abies</italic> and <italic>Fagus</italic>, 27 Oct. 1994, <italic>P. & I. Kytövuori</italic> 94-1897 (H); Savoie, Arith, under <italic>Picea</italic> on calcareous soil, 3 June 1988, <italic>P. Moënne-Loccoz</italic> PML882 (PC). – G<sc>ERMANY</sc>, Baden-Württemberg, Thölendorf, <italic>Picea</italic>, 11 Oct. 1998, UL 98/122 (TUB 011841), GenBank AY669531 as ‘<italic>C. allutus</italic>’; Schwaben, Ehingen a.d. Donau, Kohlhau, <italic>Picea</italic> plantation on calcareous soil, 28 Sept. 2010, <italic>G. Schmidt-Stohn & T.E. Brandrud</italic>, TEB 428-10 (O, TUB). – N<sc>ORWAY</sc>, Oslo, Steinbruvannet-Røverkollen, <italic>Picea</italic> forest of somewhat richer, low-herb type, 4 Sept. 2011, <italic>T.E. Brandrud</italic> 687-11 (O). – S<sc>WEDEN</sc>, Uppland, Uppsala, Nåsten, mixed forest, 27 Aug. 2005, <italic>A. Taylor</italic> 2005085, UNITE No. UDB002202 as ‘<italic>C. allutus</italic>’ (UPS); Vänge, Fiby urskog, virgin spruce forest with <italic>Pinus</italic>,<italic> Betula</italic>,<italic> Populus tremula</italic>,<italic> Colylus</italic>,<italic> Alnus glutinosa</italic>, etc., with fairly rich depressions, decaying logs very abundant, 8 Sept. 1994, <italic>P. & I. Kytövuori</italic> 94-309 (H). – USA, Washington, Grays Harbor Co., Ocean Shore State Park, under <italic>Pinus</italic> on sandy soil, 11 Nov. 2009, <italic>J.F. Ammirati & T. Niskanen</italic> 09-201 (H).</p><p>Notes — <italic>Cortinarius caesiolamellatus</italic> and <italic>C. caesiophylloides</italic> are easily distinguished from the related species in /Multiformes by their initially bluish tinged lamellae and often also stipe apex. Based on material seen so far and available descriptions, these two blue-gilled species are hardly distinguishable macromorphologically, but <italic>C. caesiolamellatus</italic> differs in slightly larger and more strongly verrucose spores. Furthermore, there seems to be a geographical differentiation in Europe; <italic>C. caesiolamellatus</italic> apparently being mainly a Central European species, whereas <italic>C. caesiophylloides</italic> is hitherto found only in northern Europe. If bluish tints have gone, <italic>C. caesiolamellatus</italic> can also be confused with conifer associated <italic>C. rufoallutus</italic> Rob. Henry ex Bidaud & Reumaux, <italic>C. multiformis</italic> Fr. and <italic>C. talimultiformis</italic> Kytöv., Liimat., Niskanen, A.F.S Taylor & Sesli. The latter two, however, have less strongly ornamented spores and lack the yellow to yellow brown large spots deep in the pileipellis. <italic>Cortinarius rufoallutus</italic> differs from <italic>C. caesiolamellatus</italic> by stouter appearance, small amygdaloid-fusoid, finely verrucose spores and abundant strongly spirally incrusted red brown hyphae deep in the pileipellis.</p><p>In our phylogenetic analysis <italic>C. caesiolamellatus</italic> forms a well-supported group (1.00 PP) and belongs together with <italic>C. caesiophylloides</italic> and <italic>C. pallidirimosus</italic> in a strongly supported (1.00 PP) subclade in clade /Multiformes (1.00 PP). The species was originally described twice in <xref rid="R7" ref-type="bibr">Bidaud et al. (2006)</xref>, as a variety of both <italic>C. rufoallutus</italic> (var. <italic>caesiolamellatus</italic>) and <italic>C. multiformis </italic>(var.<italic> caesiophyllus</italic>). Based on the molecular and morphological data neither of those relationships is supported. <italic>Cortinarius caesiolamellatus</italic> clearly represents a distinct species and is here combined at species level.</p><p><bold><italic>Cortinarius caesiophylloides</italic></bold>Kytöv., Liimat., Niskanen, Brandrud & Frøslev, <italic>sp. nov</italic>. — MycoBank MB805873; <xref ref-type="fig" rid="F2">Fig. 2</xref>g, <xref ref-type="fig" rid="F5">5</xref>a</p><p><italic>Etymology</italic>. The name refers to the bluish tints in lamellae and affinity with <italic>C. caesiolamellatus</italic> (= <italic>Cortinarius multiformis</italic> var. <italic>caesiophyllus</italic>).</p><p><italic>Type</italic>. F<sc>INLAND</sc>, Etelä-Savo, Joutsa, Koivuranta, W of Rakkolanselkä, fairly young, mesic to damp, <italic>Picea abies</italic>-dominated forest with some <italic>Betula</italic> and <italic>Pinus sylvestris</italic>, 30 Aug. 2005, <italic>T. Niskanen</italic><italic>et al.</italic> 05-016, H6029792 (holotype H; isotype NY). GenBank KF732572.</p><p><italic>Pileus</italic> 4–8 cm broad, hemispherical to plano-convex, viscid, apricot to redbrown-coloured, with hygrophanous streaks or outer zone, hygrophanous areas somewhat darker umber to less vivid (grey) brown, becoming paler ochraceous brown to ochraceous yellow with age, sometimes almost whitish ochre at centre. <italic>Lamellae</italic> emarginate, almost crowded, pale grey with a bluish tint when young, later pale brown. <italic>Stipe</italic> 5–11 cm long, 1–1.5 cm thick at apex, 1.5–3.5 cm wide at base, with a more or less distinct marginate bulb, whitish but usually with a bluish tint at apex when very young, often becoming pale brown with age, but not as pronounced as with<italic> C. multiformis</italic>. <italic>Universal veil</italic> white, sparse to hardly visible, at bulb margin. <italic>Context</italic> white, slightly bluish at the apex of the stipe when young. <italic>Odour</italic> faint to distinct of honey in the context of the stipe base. <italic>Exsiccata</italic>: pileus pale dirty brown, stipe somewhat lighter.</p><p>In MLZ: <italic>Spores</italic> 8.6–<italic>9.7</italic>–10.9 × 5.2–<italic>5.8</italic>–6.3 μm, av. = 9.0–10.2 × 5.5–6.0 μm, Q = 1.54–<italic>1.68</italic>–1.85, Qav. = 1.63–1.73 (4 specimens, 240 spores, <xref ref-type="fig" rid="F5">Fig. 5</xref>a), amygdaloid, with fairly narrow apex, sometimes almost ellipsoid, finely to moderately verrucose, warts often rounded-confluent, slightly to fairly strongly dextrinoid; with a bit narrower apex, thinner wall and finer verrucosity than in <italic>C. multiformis</italic>. <italic>Basidia</italic> 24–37 × 7–9 μm (60 basidia), 4-spored, clavate, almost colourless. <italic>Lamellar trama hyphae</italic> very pale yellowish, smooth, concolorous guttulate. <italic>Stipe apex hyphae</italic> very pale yellow, smooth, somewhat concolorous guttulate. <italic>Pileipellis</italic>: epicutis with a clear gelatinous layer with sparse, erect-entangled, 2–3 μm wide, smooth, colourless and very obscurely seen hyphae. Hypoderm present, hyphae fairly thin-walled, with pale yellowish brown amber-like parietal pigment, small brown encrust-spots present, large yellow brown spots absent, a few spirally incrusted (zebra-striped) hyphae seen deep in the cuticle.</p><p>Ecology & Distribution — In mesic coniferous forests, presumably with <italic>Picea</italic>, mainly in richer low-herb types, sometimes on calcareous soil. Known only from Fennoscandia and considered occasional. Fruits from August to September.</p><p><italic>Other specimens examined</italic>. F<sc>INLAND</sc>, Kainuu, Paltamo, Oikarilankylä, Kivesvaara, old mesic spruce forest with some <italic>Betula</italic>,<italic> Pinus</italic> and <italic>Populus tremula</italic>, 11 Sept. 2008, <italic>I. Kytövuori</italic> 08-1554, H6032621 (H); Sompion Lappi, Pelkosenniemi, Suvanto, Kalkkivaara, steep SW slope, dry pine forest with <italic>Juniperus</italic>,<italic> Betula</italic>,<italic> Populus</italic>,<italic> Alnus incana</italic>, 26 Aug. 1992, <italic>I. Kytövuori</italic> 92-3044, H6032620 (H). – N<sc>ORWAY</sc>, Nord-Trøndelag, Stjørdal, Beistadvollen, calcareous spruce forest (on karst surfaces), 13 Aug. 2009, <italic>T.E. Brandrud</italic> 277-09 (O).</p><p>Notes — <italic>Cortinarius caesiophylloides</italic> is a typical member of sect. <italic>Multiformes</italic>. Together with <italic>C. caesiolamellatus</italic> they are the only known species of the section with bluish tinted lamellae when young. <italic>Cortinarius caesiolamellatus</italic> differs from <italic>C. caesiophylloides</italic> by more strongly ornamented spores. When bluish tints are absent they can be confused with <italic>C. multiformis</italic> and <italic>C. talimultiformis</italic>, but in the latter two there are abundant, yellow to yellow-brown, large spots deep in the pileipellis. Those are practically lacking from <italic>C. caesiophylloides</italic> and <italic>C. caesiolamellatus</italic>. In <italic>C. multiformis</italic> the honey smell is either lacking or is very weak, and the stipe becomes strong brass brown with age.</p><p><italic>Cortinarius caesiophylloides</italic> is strongly supported (1.00 PP) in our phylogenetic analysis. It belongs to /Multiformes (1.00 PP) and further, in a well-supported (1.00 PP) subclade with <italic>C. caesiolamellatus</italic> and <italic>C. pallidirimosus</italic>. The ITS sequences of <italic>C. caesiophylloides</italic> have one base polymorphism and the maximum pairwise distance is zero. The difference to <italic>C. pallidirimosus</italic> is 11 substitutions and indel positions.</p><p><bold><italic>Cortinarius melleicarneus</italic></bold> Kytöv., Liimat., Niskanen & Brandrud, <italic>sp. nov.</italic> — MycoBank MB805874; <xref ref-type="fig" rid="F5">Fig. 5</xref>b, <xref ref-type="fig" rid="F6">6</xref>a</p><p><italic>Etymology</italic>. The name refers to the colour of the pileus.</p><p><italic>Type</italic>. E<sc>STONIA</sc>, Hiiumaa, Pühalepa, Sarve, Soonlepa, deciduous forest (<italic>Corylus</italic>,<italic> Quercus</italic>) with some <italic>Pinus</italic> on calcareous soil, 16 Sept. 2001, <italic>I. Kytövuori</italic> 01-053 (holotype H; isotype NY). GenBank KF732577.</p><p><italic>Pileus</italic> 4–10 cm broad, hemispherical to convex, with rather persistently incurved margin, then expanded, sometimes somewhat silvery-silky from fine veil remnants, cream-coloured, pale yellow brown, honey brown to more grey brown, with hygrophanous streaks or patches/zones near margin; hygrophanous zones somewhat darker grey brown, almost with an olivaceous brown tinge. <italic>Lamellae</italic> emarginate, crowded, first pale greyish white, later pale greyish brown. <italic>Stipe</italic> 5–7(–9) cm long, 1.2–2 cm thick at apex, 2–3 cm wide at base, clavate or with a somewhat marginate bulb, short and robust, white. <italic>Universal veil</italic> very sparse at bulb margin, white. <italic>Context</italic> in pileus pale yellow brown, marbled hygrophanous flesh-coloured, in stipe white. <italic>Odour</italic> not recorded.</p><p>In MLZ: <italic>Spores</italic> 7.9–<italic>8.7</italic>–9.5 × 4.3–<italic>4.7</italic>–5.2 μm, av. = 8.6–8.9 × 4.7–4.8 μm, Q = 1.67–<italic>1.86</italic>–2.00, Qav. = 1.85–1.89 (2 specimens, 120 spores, <xref ref-type="fig" rid="F5">Fig. 5</xref>b), amygdaloid to fusoid, with a low suprahilar depression and blunt apex, moderately to fairly strongly, densely and coarsely, somewhat unevenly, not sharply verrucose, slightly to moderately dextrinoid. <italic>Basidia</italic> 25–31 ×7–8 μm (40 basidia), 4-spored, clavate, many granulose-guttulate. <italic>Lamellar trama hyphae</italic> pale yellow, smooth. <italic>Stipe apex hyphae</italic> very pale yellowish, smooth, coloured guttules absent. <italic>Pileipellis</italic>: epicutis gelatinous, with 3–8 μm wide, thin-walled, smooth to very finely incrusted, colourless and very obscure hyphae. Hypoderm present, with pale yellowish brown walls, few incrusted hyphae and small brown spots deep in the pileipellis.</p><p>Ecology & Distribution — With thermophilous deciduous trees (<italic>Corylus</italic>,<italic> Quercus</italic>) on calcareous soil, including near-shore sandy shell-beds. Known from Estonia and Norway.</p><p><italic>Other specimens examined</italic>. N<sc>ORWAY</sc>, Aust-Agder, Grimstad, Fevik, under <italic>Quercus</italic>,<italic> Fagus </italic>and (planted)<italic> Larix </italic>also present, on sandy soil, rather rich, probably with shell-bed deposits, 21 Sept. 1994, leg. <italic>I-L. Fonneland</italic>, det. <italic>T.E. Brandrud</italic> 86-94, O 125960 (O, sub nom. <italic>C. areni-silvae</italic>).</p><p>Notes — <italic>Cortinarius melleicarneus</italic> belongs to /Multiformes. It differs from the other species of the group by honey-coloured pileus, fusoid spores, and habitat with deciduous trees on calcareous soil. No honey-smell was noted in the two collections, and the lack of this smell might be a diagnostic character towards the usually strongly honey-smelling <italic>C. talus</italic>. <italic>Cortinarius talus</italic> has furthermore usually an innately fibrillose pileus structure and rarely possess short-stemmed, compact carpophores. <italic>Cortinarius cruentipellis </italic>grows in similar habitats, but it has ochraceous yellow universal veil on bulb margin, larger spores, and red-colouring pileipellis in MLZ. In dry conditions, <italic>C. melleicarneus</italic> might resemble <italic>C. areni-silvae</italic> (Brandrud) Brandrud which also occurs in sandy soils, and the Norwegian collection was originally determined as <italic>C. areni-silvae</italic> (based on dry material and collectors notes). ITS sequences of the two <italic>C. melleicarneus</italic> specimens are identical and it forms a clade in our analysis. It differs by 11 substitutions and indel positions from <italic>C. talimultiformis</italic>, 14 from <italic>C. talus</italic>, 15 from <italic>C. multiformis</italic>, and 17 from <italic>C. rufoallutus</italic> Rob. Henry ex Bidaud & Reumaux<italic>.</italic></p><p><bold><italic>Cortinarius pallidirimosus</italic></bold> Kytöv., Liimat. & Niskanen, <italic>sp. </italic><italic>nov</italic>. — MycoBank MB805875; <xref ref-type="fig" rid="F5">Fig. 5</xref>c, <xref ref-type="fig" rid="F6">6</xref>b</p><p><italic>Etymology</italic>. The name refers to the pale pileus with streaks.</p><p><italic>Type</italic>. F<sc>INLAND</sc>, Inarin Lappi, Utsjoki, Kevo, Tsieskuljohka, mesic heath forest with <italic>Betula</italic> and <italic>Pinus</italic>, with some moist depressions, 17 Aug. 1995, <italic>I. Kytövuori</italic> 95-585, H6035694 (holotype H; isotype NY). GenBank KF732578.</p><p><italic>Pileus</italic> 3–9 cm broad, hemispherical to convex, then expanded, viscid, very finely innately fibrillose, whitish to cream-coloured, centre brownish yellow, becoming ochraceus with age, with hygrophanous streaks. <italic>Lamellae</italic> emarginate, crowded, pale greyish brown when young, later pale brown. <italic>Stipe</italic> 6–13 cm long, 0.7–1.5 cm thick at apex, 1.5–2.5 cm wide at base, clavate to almost cylindrical, whitish, becoming very pale brown with age. <italic>Universal veil</italic> white, sparse. <italic>Context</italic> white. <italic>Odour</italic> in context honey-like. <italic>Exsiccata</italic>: pileus cream-coloured to ochre brownish, somewhat darker at the centre, lighter towards the margin, stipe somewhat lighter than pileus.</p><p>In MLZ: <italic>Spores</italic> 8.6–<italic>9.6</italic>–10.7 × 5.2–<italic>5.7</italic>–6.1 μm, av. = 9.0–10.1 × 5.5–5.9, Q = 1.54–<italic>1.70</italic>–1.85, Qav. = 1.60–1.76 (12 specimens, 320 spores, <xref ref-type="fig" rid="F5">Fig. 5</xref>c), amygdaloid to amygdaloid-ellipsoid, moderately to fairly strongly, unevenly, sometimes coarsely verrucose, warts fairly wide but not high, slightly (to moderately) dextrinoid. <italic>Basidia</italic> 24–36 × 7.5–9 μm (100 basidia), 4-spored, clavate, almost colourless. <italic>Lamellar trama hyphae</italic> pale pellucid yellowish, smooth, somewhat concolorous granulose-guttulate. Few small mounds of colourless crystals sometimes present in the lamellar trama. <italic>Stipe apex hyphae</italic> pale yellowish, smooth. <italic>Pileipellis</italic>: epicutis with a clear slime layer with 1.5–3 μm wide, smooth, colourless and very obscure hyphae. Hypoderm present, very pale brownish to almost colourless.</p><p>Ecology & Distribution — In the middle to northern boreal, mesic, mixed forests with <italic>Betula</italic>, among mosses, on oligotrophic to eutrophic soil. Often solitarily or in small groups. Known from Fennoscandia, Sakha, Russia, and Oregon, USA, and considered occasional. Fruiting in late summer to autumn.</p><p><italic>Other specimens examined</italic>. F<sc>INLAND</sc>, Kainuu, Puolanka, Väyrylä, Körölä, grass-herb-spruce forest with some pines and hardwood bushes, 15 Sept. 1997, <italic>I. Kytövuori</italic> 97-1523, H6035715 (H); Perä-Pohjanmaa, Rovaniemi, Louevaara, Tuohilaki nature reserve area (west), eutrophic, submesic to mesic <italic>Picea abies</italic> forest with <italic>Betula</italic>,<italic> Populus tremula</italic> and some <italic>Pinus sylvestris</italic>, 29 Aug. 2004, <italic>T. Niskanen & K. Liimatainen</italic> 04-470, H6029374 (H); Tervola, Louepalo, Kätkävaaran marmorilouhos, E of the western quarry, S of the track, gently E sloping forest of young <italic>Betula</italic> and <italic>Picea</italic>, with some <italic>Populus tremula</italic>,<italic> Alnus incana</italic> and <italic>Salix</italic> spp., 23 Aug. 2007, <italic>I. Kytövuori</italic> 07-692, H6000925 (H); Peura, Kaitalampi, Kaitaharju, half-open, eutrophic grass-herb-spruce forest with some pines, calcareous soil, 5 Sept. 1992, <italic>I. Kytövuori</italic> 92-1779, H6035693 (H); Koillismaa, Kuusamo, Oulanka National Park, Ampumavaara, mesic to damp, herb-rich <italic>Picea abies</italic> forest with some <italic>Pinus</italic>,<italic> Populus</italic> and <italic>Betula</italic>, 19 Sept. 2002, <italic>T. Niskanen et al. </italic>02-859, H6033129 (H); Kittilän Lappi, Kolari, Teuravuoma-Lappea (Korkealehdontie), Tappikumpu, mesic spruce-birch forest, 18 Aug. 1998, <italic>I. Kytövuori</italic> 98-1078, H6035705 (H); Sompion Lappi, Sodankylä, Tähtelä, Hassikankaannokka, mesic spruce forest with some pines and hardwood trees, 24 Aug. 1992, <italic>I. Kytövuori </italic>92-966 (H); Inarin Lappi, Utsjoki, Kevo, Tsieskuljohka, mesic heath forest with <italic>Betula</italic> and <italic>Pinus</italic>, some moist depressions, 17 Aug. 1995, <italic>I. Kytövuori</italic> 92-540, H6035704 (H). – N<sc>ORWAY</sc>, Troms, Storfjord, Skibotndalen, Lullesletta, dry pine forest with <italic>Betula</italic> and <italic>Salix</italic>, 13 Aug. 1998, <italic>I. Kytövuori</italic> 98-711 (H). – S<sc>WEDEN</sc>, Jämtland, Bodsjö, Sidsjö, Stuguberget, dryish, fairly young spruce-pine forest with some hardwood trees and bushes, some moist depressions, 4 Sept. 1997, <italic>I. Kytövuori</italic> 97-699, H7019576 (H); Frösö, Böle, Fillstabäcken, damp to mesic coniferous forest (<italic>Picea</italic>,<italic> Pinus</italic>) on calcareous soil, 2 Sept. 2003, <italic>Niskanen</italic><italic>et al.</italic> 03-1058, H7018404 (H).</p><p><italic>Additional specimens</italic>. R<sc>USSIA</sc>, Sakha, Khangalasskiy Ulus, Myachei-Sise Mts, <italic>Larix gmelinii</italic> forest with <italic>Betula platyphylla</italic>, 8 Aug. 1999, <italic>U. Peintner</italic> IB19990590 (IB), UNITE No. UDB001073. – USA, Oregon, source: mycorrhizal root tip, clone=”8_73M8, GenBank JQ393042.</p><p>Notes — <italic>Cortinarius pallidirimosus</italic> can easily be identified by the cream-coloured, hygrophanous streaked pileus, honey-like smell in context, large, amygdaloid to amygdaloid-ellipsoid spores, and habitat with <italic>Betula </italic>in the boreal zone. The other <italic>Betula</italic> associated species in sect. <italic>Multiformes</italic>, <italic>C. talus</italic> Fr., has relatively wider, ochraceous yellow pileus and smaller spores (7.3–<italic>8.0</italic>–8.8 × 4.5–<italic>5.0</italic>–5.2 μm). <italic>Cortinarius gentianeus</italic> Bidaud and <italic>C. cremeiamarescens</italic> Kytöv., Liimat. & Niskanen are reminiscent of <italic>C. pallidirimosus</italic>, but the former two have a bitter tasting pileus, indistinct odour, and smaller, citriform to narrowly fusoid spores. In addition, the lamellae and pileipellis of both species are strongly red in MLZ.</p><p>The placement of <italic>C. pallidirimosus</italic> in /Multiformes is well supported (1.00 PP) and it forms a strongly supported subclade with <italic>C. caesiolamellatus</italic> and <italic>C. caesiophylloides</italic>. From its sister species <italic>C. caesiophylloides</italic> it differs by 11 substitutions and indel positions in ITS regions. The sequences of <italic>C. pallidirimosus </italic>have one base polymorphism and six intraspecific variation sites and the maximum pairwise distance is six. The amount of variation is highly compared to many other species in the genus <italic>Cortinarius</italic> in which the variation typically is 0–2 substitutions and/or indel positions. Most likely the data includes a close sister species, but more specimens would be needed to study this complex more in detail.</p><p><bold><italic>Cortinarius talimultiformis</italic></bold>Kytöv., Liimat., Niskanen, A.F.S. Taylor & Sesli, <italic>sp. nov</italic>. — MycoBank MB805876; <xref ref-type="fig" rid="F5">Fig. 5</xref>d, <xref ref-type="fig" rid="F6">6</xref>c</p><p><italic>Etymology</italic>. The name refers to the appearance of the species, which has features from both <italic>C. multiformis</italic> and <italic>C. talus</italic>.</p><p><italic>Type</italic>. S<sc>WEDEN</sc>, Uppsala, Hässelby Park, mixed forest, 11 July 2004, <italic>A. Taylor</italic> AT2004096 (holotype UPS; isotype S) UNITE No. UDB001167. GenBank KF732583.</p><p>= <italic>Cortinarius aurantionapus</italic> var. <italic>similis</italic> Moënne-Locc., Atlas des Cortinaires 16: 1096. 2006. — <italic>Type</italic>. <italic>P. Moënne-Loccoz</italic>, PML883 (holotype PC), France, Haute-Savoie, Aviernoz, under <italic>Picea abies</italic>, on calcareous soil, 6 June 1988, .</p><p>Illustration — Bidaud et al. (2006: pl. 596).</p><p><italic>Pileus</italic> 4–12 cm broad, hemispherical to plano-convex, viscid, whitish fibrillose, especially near the margin, ochraceous yellow to red brown, with strongly hygrophanous streaks. <italic>Lamellae</italic> emarginate, crowded, pale grey when young, later pale greyish brown. <italic>Stipe</italic> 4–10 cm long, 1–2 cm thick at apex, 2–3.5 cm wide at base, clavate or with a somewhat marginate bulb, white. <italic>Universal veil</italic> white, sparse, sometimes fairly abundant at pileus margin. <italic>Context</italic> white. <italic>Odour</italic> slightly honey-like at the context of the base of the stipe. <italic>Exsiccata</italic>: pileus leather brown to yellow brown to mahogany brown, sometimes whitish variegate at the centre, stipe whitish to pale brownish.</p><p>In MLZ: <italic>Spores</italic> 8.4–<italic>9.5</italic>–10.7 × 5.0–<italic>5.5</italic>–6.1 μm, av. = 8.9–9.9 × 5.3–5.8 μm, Q = 1.58–<italic>1.73</italic>–1.87, Qav. = 1.68–1.78 (9 specimens, 320 spores, <xref ref-type="fig" rid="F5">Fig. 5</xref>d), amygdaloid to narrowly amygdaloid to amygdaloid-fusoid, with a long, narrowish apex, rather weakly to fairly strongly, lowly verrucose, slightly to moderately dextrinoid. <italic>Basidia</italic> 27–36 × 7–9 μm (80 basidia), 4-spored, clavate. <italic>Lamellar trama hyphae</italic> very pale yellowish, smooth. <italic>Stipe apex hyphae</italic> 4–8 μm wide, pale yellowish, smooth, some outermost ones 2–3 μm wide, in entangled bundles. <italic>Pileipellis</italic>: epicutis with fairly thick, clear slime layer with 1.5–2.5 μm wide, colourless, hardly visible hyphae. Hypoderm present, in its upper part large, brownish yellow pigment spots abundant.</p><p>Ecology & Distribution — In hemiboreal to boreal, mesic coniferous forests with <italic>Picea</italic> and <italic>Abies</italic>. Known from North and Central Europe, and East Black Sea Region’s mountains in Turkey, considered common. Basidiocarps occur from June to late August.</p><p><italic>Other specimens examined</italic>. F<sc>INLAND</sc>, Pohjois-Häme, Virrat, Killinkoski, <italic>Abies sibirica</italic> forest, 30 Aug. 1966, <italic>I. Kytövuori</italic>, H6032747 (H); Oulun Pohjanmaa, Kiiminki, Keskikylä, Pikkuhalmeenmaa, grass-herb-spruce forest with some <italic>Betula</italic>,<italic> Populus tremula</italic> and <italic>Pinus</italic>, on calcareous soil, 15 Aug. 2007, <italic>M. Toivonen & I. Kytövuori</italic> 07-131, H6000360 (H); Perä-Pohjanmaa, Tornio, Kalkkimaa, Kalkkimaan lehto (NW), mainly grass-herb forest with <italic>Picea</italic>,<italic> Betula</italic>,<italic> Populus tremula</italic>,<italic> Juniperus</italic> and <italic>Pinus</italic>, on calcareous soil, 22 Aug. 2007, <italic>I. Kytövuori</italic> 07-605, H6000838 (H). – F<sc>RANCE</sc>, Haute-Savoie, Aviernoz, under <italic>Picea abies</italic>, on calcareous soil, 6 June 1988, <italic>P. Moënne-Loccoz</italic>, PML883 (PC, holotype of <italic>C. aurantionapus</italic> var. <italic>similis</italic>). – G<sc>ERMANY</sc>, Baden-Württemberg, Titisee, saurer Boden unter Fichten, 10 Sept. 2002, Saar 4855, TUB 011864 (TUB), GenBank AY669532 as ‘<italic>C. allutus</italic>’ (TUB). – N<sc>ORWAY</sc>, Oppland, Gran, Lygna, mesic spruce forest with some <italic>Betula</italic> and <italic>Pinus</italic>, 13 Sept. 2010, <italic>E. Bendiksen & I. Kytövuori</italic> (H). – T<sc>URKEY</sc>, East Black Sea Region, Trabzon, Macka, Gurgenagac village, <italic>Picea orientalis</italic> forest, 7 July 2010, <italic>E. Sesli</italic>, SES 2741.</p><p>Notes — The appearance of <italic>C. talimultiformis</italic> is rather variable, but most often it is fairly low, wide and stout. The colour of the pileus varies from ochraceous yellow to mahogany brown and the form of the stipe from clavate to bulbous. Typical for the species, however, are the white fibrils on the pileus margin, narrow, prominently verrucose spores, abundant large yellow spots in the pileipellis, and habitat with <italic>Picea</italic> or <italic>Abies</italic>. The fruiting period of this species starts early and it can be found in early June. The sister species <italic>C. multiformis</italic> Fr. grows in similar habitats but has no fibrils on the pileus, the spores are moderately to strongly dextrinoid, fairly finely verrucose, and abundant large brownish yellow spots are present deep in the pileipellis. The colour of the pileus in the exsiccata is uniformly red brown. The other common species, <italic>C. talus</italic> Fr., is associated with deciduous trees, is pale ochraceous yellow, and the spores are smaller (7.3–<italic>8.0</italic>–8.8 × 4.5–<italic>5.0</italic>–5.2 μm). <italic>Cortinarius talimultiformis</italic> formed a well-supported group in our phylogenetic analysis. It is a sister species of <italic>C. multiformis</italic> from which it differs by 6 substitutions and indel positions in ITS regions.</p></sec><sec id="s13"><title>/PHLEGMACIOIDES</title><p><bold><italic>Cortinarius balteatialutaceus</italic></bold> Kytöv., Liimat. & Niskanen, <italic>sp. nov</italic>. — MycoBank MB805877; <xref ref-type="fig" rid="F6">Fig. 6</xref>d, <xref ref-type="fig" rid="F7">7</xref>a</p><p><italic>Etymology</italic>. The name refers to the colour of the pileus.</p><p><italic>Type</italic>. S<sc>WEDEN</sc>, Jämtland, Frostviken, Jormlien, Säterklumpen, <italic>Betula</italic> forest with solitary <italic>Picea</italic>, 7 Sept. 2009, <italic>P. & I. Kytövuori</italic> 09-751 (holotype H; isotype NY). GenBank KF732586.</p><p><italic>Pileus</italic> 6–12 cm broad, hemispherical to plano-convex, with a fairly persistently involute margin, first very slightly viscid, dry with age, pale brown with a whitish to very pale brown margin, becoming darker brown from centre. <italic>Lamellae</italic> emarginate, crowded, pale brownish grey when young, later pale brown. <italic>Stipe</italic> 5–7 cm long, 1.3–2.5 cm thick at apex, 1.8–3.5 cm wide at base, clavate, sometimes with an almost marginate bulb, whitish. <italic>Universal veil</italic> white, sparse. <italic>Context</italic> white. <italic>Odour</italic> indistinct. <italic>KOH-reaction</italic> in context yellow.<italic> Exsiccata</italic>: pileus leather brown at centre, paler at margin, stipe very pale brownish.</p><p>In MLZ: <italic>Spores</italic> 9.1–<italic>10.0</italic>–10.9 × 5.0–<italic>5.4</italic>–5.9 μm, av. = 9.8–10.4 × 5.3–5.5 μm, Q = 1.75–<italic>1.85</italic>–1.98, Qav. = 1.78–1.87 (5 specimens, 140 spores, <xref ref-type="fig" rid="F7">Fig. 7</xref>a), amygdaloid-fusoid, sometimes with a low suprahilar depression, moderately to fairly strongly verrucose, warts mostly not anastomosing, weakly dextrinoid, yellowish brown. Although narrow, the spores are quite different in appearance than in <italic>C. balteaticlavatus</italic>. <italic>Basidia</italic> 30–39 × 7–8.5 μm (50 basidia), clavate, pale brownish, very finely granulose at the base. <italic>Lamellar trama hyphae</italic> with moderate to large, dark rice-like granules on yellowish green ground colour. <italic>Stipe apex hyphae</italic> pale yellow to pale brown, outermost hyphae with reddish brown, granulose contents. <italic>Pileipellis</italic>: epicutis very weakly gelatinous, uppermost hyphae 3–8 μm wide, pale ochraceous to reddish brown, very finely, densely incrusted, with some small, red brown granules. Deep in the pileipellis some hyphae with fairly large, red brown to blackish brown granules and chips. The thin-walled upper hyphae lost with age and are mostly not seen in older basidiomes. Hypoderm absent.</p><p>Ecology & Distribution — Presumably with <italic>Betula</italic>, in subalpine birch forests but also in middle and northern boreal forests at least in oceanic areas. Produces fruitbodies from mid-August to mid-September. Known from Fennoscandia. No sequences of this species exist in public databases.</p><p><italic>Other specimens examined</italic>. F<sc>INLAND</sc>, Inarin Lappi, Utsjoki, Kevo, shore of the river Tsarsjoki, W of the cascade, herb-rich, temporarily flooded site on a small island, with <italic>Betula</italic> and <italic>Salix</italic> bushes, 16 Aug. 1995, <italic>J. Vauras</italic> 10452 (TUR5282). – N<sc>ORWAY</sc>, Sogn og Fjordane, Sogndal, on the E side of the main road vis-vis the Kaupanger centre, deciduous forest (<italic>Populus</italic>,<italic> Betula</italic>) with some <italic>Pinus</italic>, 10 Sept. 2000, <italic>J. Ruotsalainen</italic> 100900 (H); Sør-Trøndelag, Oppdal, Ålbu, Ålbusbakkan, sloping dryish forest with <italic>Pinus</italic>,<italic> Corylus</italic> and <italic>Betula</italic>, 5 Sept. 2010, <italic>I. Kytövuori</italic> (H); Nordland, Grane, Fagerli, rich spruce forest with few young <italic>Betula</italic>, calcareous soil, 3 Sept. 2010, <italic>I. Kytövuori</italic> (H).</p><p>Notes — <italic>Cortinarius balteatialutaceus</italic> belongs to /Balteatoalbi together with <italic>C. balteatoalbus</italic> Rob. Henry and <italic>C. balteaticlavatus</italic> Kytöv., Liimat. & Niskanen. In the ITS regions it differs by 5 substitutions and indel positions from <italic>C. balteatoalbus</italic> and 7 substitutions and indel positions from <italic>C. balteaticlavatus</italic>. The sequences of <italic>C. balteatialutaceus </italic>have one base difference and the maximum pairwise distance is one. Typical for the species of the clade, compared to many other species of /Phlegmacioides, are basidiomata totally without bluish tints even when young and narrow, rather small spores. <italic>Cortinarius balteatialutaceus</italic> is a stout, <italic>C. balteatus</italic>-like species and grows in subalpine birch forests but also in middle and northern boreal forests at least in oceanic areas. The sister species <italic>C. balteatoalbus</italic> has less verrucose spores and lamellar trama hyphae are densely small-granulose in MLZ. In addition, <italic>C. balteatoalbus</italic> has a more southern, hemiboreal to montane distribution although it also rarely occurs in the middle boreal zone. <italic>Cortinarius balteaticlavatus</italic> has a relatively longer stipe, brown pileus margin, on average smaller spores (av. = 8.5–9.3 × 4.9–5.1 μm) and it grows in mixed forests in boreal zone. <italic>Cortinarius balteatus</italic> (Fr.) Fr. and <italic>C. badiolatus</italic> (M.M. Moser) M.M. Moser occur in subalpine forests but have larger spores, <italic>C. balteatus</italic> 10–12 × 5.5–6.5 μm and <italic>C. badiolatus</italic> 10.5–12.5 × 5.5–7.5 μm. In addition, the pileus margin of <italic>C. balteatus</italic> is usually blue, at least when young. <italic>Cortinarius areni-silvae</italic> (Brandrud) Brandrud is reminiscent of <italic>C. balteatialutaceus</italic> but the former grows in dry, sandy pine heaths and has smaller spores (7.7–9.5 × 4.3–5.2 μm).</p><p><bold><italic>Cortinarius balteatibulbosus</italic></bold> Kytöv., Niskanen, Liimat., Bojantchev & A.F.S. Taylor, <italic>sp. nov</italic>. — MycoBank MB805878; <xref ref-type="fig" rid="F6">Fig. 6</xref>e, <xref ref-type="fig" rid="F7">7</xref>b</p><p><italic>Etymology</italic>. The name refers to the confusion of this species with <italic>C. balteatoalbus</italic> and to the marginate bulb.</p><p><italic>Type.</italic> F<sc>INLAND</sc>, Uusimaa, Espoo, Nuuksio, Pirttimäki, half-open cut meadow, 4 Sept. 1998, <italic>I. Kytövuori</italic> 98-1624, H6033539 (holotype H; isotype NY). GenBank KF732589.</p><p><italic>Pileus</italic> 6–13 cm broad, hemispherical to plano-convex, slightly viscid in moist weather, otherwise dry, coarsely innately fibrillose, pale brown with a whitish to very pale brown margin, rarely completely white, becoming brown with age. <italic>Lamellae</italic> emarginate, crowded, pale grey when young, later pale brown. <italic>Stipe</italic> 4–7 cm long, 1.5–2.5 cm thick at apex, 2.5–4 cm wide at base, with a more or less marginate bulb, whitish. <italic>Universal veil</italic> white, later ochraceous brown at bulb margin, sparse. <italic>Context</italic> white. <italic>Odour</italic> indistinct in lamellae, in flesh mild and pleasant. <italic>KOH-reaction</italic> in context yellow. <italic>Exsiccata</italic>: pileus brown to brownish to leather-coloured, stipe at the top whitish, at the base concolorous with the pileus.</p><p>In MLZ: <italic>Spores</italic> 9.1–<italic>9.9</italic>–10.9 × 5.2–<italic>5.7</italic>–6.1 μm, av. = 9.6–10.2 × 5.6–5.8 μm, Q = 1.58–<italic>1.74</italic>–1.89, Qav. = 1.66–1.82 (6 specimens, 120 spores, <xref ref-type="fig" rid="F7">Fig. 7</xref>b), broadly amygdaloid to citriform, fairly strongly, darkly verrucose, fairly dark-coloured, somewhat dextrinoid. <italic>Basidia</italic> 30–42 × 7–10 μm (60 basidia), clavate, brownish, finely granulose and with few larger granules at the base. <italic>Lamellar trama hyphae</italic> with moderate to large, dark rice-like granules on brownish to yellowish ground colour. <italic>Stipe apex hyphae</italic> orange brown, in outermost hyphae with abundant dark red brown, partly blackish brown contents. <italic>Pileipellis</italic>: gelatinous layer practically absent, uppermost hyphae 4–8 μm wide, thin-walled, ochraceous brown to reddish brown, mostly finely, densely, spirally incrusted, with few red brown granules. Lower hyphae somewhat wider with slightly thicker, smooth to incrusted, brown walls and dark sepia brown granular contents, granules small to large. The thin-walled upper hyphae lost with age and are mostly not seen in older basidiomata.</p><p>Ecology & Distribution — In rich mixed forests, in parks, and on yard lawns with different deciduous trees (<italic>Tilia</italic>,<italic> Quercus</italic>,<italic> Fagus</italic>,<italic> Populus</italic>,<italic> Corylus</italic>,<italic> Betula</italic>). Until now all the collections known to us are from the hemiboreal zone. Widespread in Europe, from Bulgaria to Finland. Can fruit very early in the season, in July, but also in September.</p><p><italic>Other specimens examined</italic>. B<sc>ULGARIA</sc>, Saranzi, 700 m asl, <italic>Quercus cerris</italic>, <italic>Q. petraea</italic> and <italic>Q. frainetto</italic>, 10 June 2010, <italic>D. Bojantchev</italic> DBB33060 (UC). – F<sc>INLAND</sc>, Varsinais-Suomi, Vihti, Lintumäki, Ruohoisnummentie, fairly old, fairly rich <italic>Picea abies</italic> dominated forest, 19 July 2004, <italic>H. Tuovila</italic>, H6027358 (H); Uusimaa, Espoo, Nuuksio, Pirttimäki fairly rich grass-herb forest with <italic>Populus tremula</italic>,<italic> Betula</italic>,<italic> Alnus incana</italic>,<italic> Quercus</italic>,<italic> Corylus</italic>,<italic> Prunus padus</italic>,<italic> Salix</italic> spp., and some old pines and young spruces, 2 Sept. 1993, <italic>I. Kytövuori</italic> 93-639, H6032755 (H), 4 Sept. 2004, Ukkola (H); Helsinki, Tamminiemi, on yard, under <italic>Tilia</italic>, on lawn, 3 July 2000, <italic>T. Niskanen</italic> H6032413 (H); cut lawn, under <italic>Tilia</italic>, 18 Sept. 2000, <italic>I.Kytövuori</italic> (H); Vantaa, Tikkurila, at the river, on a cut lawn under <italic>Tilia</italic>, 22 July 2004, <italic>I. Kytövuori</italic> 04-044 (H). – S<sc>WEDEN</sc>, Uppsala, Berthåga graveyard, mixed forest, 21 July 2004, <italic>A. Taylor</italic> 2004091, UNITE No. UDB001132 as ‘<italic>C. balteatoalbus</italic>’ (UPS); <italic>A. Taylor </italic>2004127, UNITE No. UDB000722 as ‘<italic>C.balteatoalbus</italic>’ (UPS); Järfälla, Järfälla Naturreservat, mixed forest of <italic>Picea abies</italic>, <italic>Quercus robur</italic>, <italic>Betula</italic>, <italic>Corylus</italic>, 7 Sept. 2010, <italic>D. Bojantchev</italic> DBB36892 (UC); Slottsbacken, under <italic>Fagus sylvatica</italic>, 2 July 2004, <italic>A. Taylor</italic> 2004045, UNITE No. UDB000711 as ‘<italic>C. balteatoalbus</italic>’ (UPS); Stenhagen, mixed forest, 10 July 2004, <italic>A. Taylor </italic>2004088, UNITE No. UDB000715 as ‘<italic>C. balteatoalbus</italic>’ (UPS).</p><p>Notes — Typical for <italic>C. balteatibulbosus</italic> are low and stout basidiomata without bluish tints, coarsely innately fibrillose pileus, marginate bulb, broadly amygdaloid to citriform, fairly strongly verrucose spores, and habitat with deciduous trees. The species of /Balteatoalbi have smooth to only finely fibrillose pileus and clavate to almost cylindrical stipe. In addition, the spores of <italic>C. balteaticlavatus</italic> Kytöv., Niskanen & Liimat. are smaller, especially shorter. <italic>Cortinarius balteatialutaceus</italic> is almost whitish and has narrower spores and a much more northern distribution. <italic>Cortinarius balteatoalbus</italic> is paler, the spores are narrower, pileipellis is more incrusted, and in MLZ the red brown small granules (large, dark ones in <italic>C. balteatibulbosus</italic>) are abundant in lamellar trama hyphae.</p><p>In our phylogenetic analysis <italic>C. balteatibulbosus</italic> forms a well supported clade in /Phlegmacioides. It groups together with <italic>C.flavescentipes</italic> Reumaux (= <italic>C. balteatocumatilis</italic> Rob. Henry ex P.D. Orton s.auct.) and <italic>C. pseudonebularis</italic> Moënne-Locc. (1.00 PP). The sequences of <italic>C. balteatibulbosus</italic> have one length polymorphism and the maximum pairwise distance is zero. In ITS regions it differs by 19 substitutions and indel positions from <italic>C. flavescentipes</italic>. The species was previously called <italic>C. balteatoalbus</italic> in northern Europe but our studies revealed that <italic>C. balteatoalbus</italic> is a completely different species and therefore this one is described as new.</p><p><bold><italic>Cortinarius balteaticlavatus</italic></bold> Kytöv., Liimat. & Niskanen, <italic>sp. nov</italic>. — MycoBank MB805879; <xref ref-type="fig" rid="F6">Fig. 6</xref>f, <xref ref-type="fig" rid="F7">7</xref>c</p><p><italic>Etymology</italic>. The name refers to the affinity of <italic>C. balteatus</italic> and clavate stipe.</p><p><italic>Type</italic>. F<sc>INLAND</sc>, Pohjois-Häme, Virrat, Hauhuu, Sikosaari, Salmela, on the grassy roadside with young <italic>Picea</italic>,<italic> Betula</italic>,<italic> Populus tremula</italic>,<italic> Alnus incana</italic> and <italic>Salix</italic> spp., 23 Aug. 1996, <italic>I. Kytövuori</italic> 96-595, H6032412 (holotype H; isotype NY). GenBank KF732596.</p><p><italic>Pileus</italic> 5–9 cm broad, hemispherical to plano-convex, with an involute margin when young, dry, sand brown to brown, completely without bluish tints. <italic>Lamellae</italic> emarginate, crowded, pale brownish grey when young, later brown. <italic>Stipe</italic> 6–11 cm long, 1.2–2 cm thick at apex, 1.5–2.5 cm wide at base, clavate, whitish. <italic>Universal veil</italic> whitish to ochraceous, fairly sparse. <italic>Context</italic> thick, white to very pale marbled brownish. <italic>Odour</italic> indistinct. <italic>KOH-reaction</italic> in context yellow. <italic>Exsiccata</italic>: pileus uniformly pale brown to brown, stipe whitish at the top, pale brown at the base.</p><p>In MLZ: <italic>Spores</italic> 8.2–<italic>8.9</italic>–10.0 × 4.5–<italic>5.0</italic>–5.4 μm, av. = 8.5–9.3 × 4.9–5.1 μm, Q = 1.60–<italic>1.78</italic>–1.94, Qav. = 1.72–1.85 (5 specimens, 160 spores, <xref ref-type="fig" rid="F7">Fig. 7</xref>c), narrowly amygdaloid to fusoid, with blunt apex, finely, densely verrucose, warts not anastomosing, fairly light-coloured, very faintly dextrinoid. <italic>Basidia</italic> 27–39 × 6.5–8.5 μm (40 basidia), clavate, 4-spored, pale brownish, finely granulose at the base, sometimes with a few dark granules. <italic>Lamellar trama hyphae</italic> with small to moderate, dark rice-shaped granules. <italic>Stipe apex hyphae</italic> yellowish brownish, outermost ones with yellow brown to red brown substance and some dark granules. <italic>Pileipellis</italic>: gelatinous layer absent, uppermost hyphae 4–10 μm wide, thin-walled, ochraceous brown, finely, densely, spirally to spot-like incrusted, with many large, dark red brown, angular particles. Lower hyphae with slightly thicker, smooth to incrusted, brown walls and red brown to sepia brown granular contents, granules small to large. The thin-walled upper hyphae lost with age and are mostly not seen in older basidiomata.</p><p>Ecology & Distribution — In mixed forests (<italic>Betula</italic>,<italic> Populus</italic>,<italic> Salix</italic>,<italic> Picea</italic>,<italic> Pinus</italic>) but the tree host is unknown, most of the collections from grassy roadsides. Occurs in the southern boreal to the northern boreal zones, known from South Finland to Lapland. Fruits from mid-August to mid-September.</p><p><italic>Other specimens examined</italic>. F<sc>INLAND</sc>, Etelä-Savo, Mäntyharju, Juolasvesi, mesic spruce heath forest with swampy depressions, <italic>Pinus</italic>,<italic> Betula</italic> and <italic>Populus tremula</italic>, road ditch, 2 Sept. 1996, <italic>I. Kytövuori</italic> 96-782, H6032415 (H); Salmela, half-open forest of <italic>Betula</italic> and <italic>Populus tremula</italic> and some pines and spruces, 22 Aug. 1998, <italic>I. Kytövuori</italic> 98-1337 (H); Koillismaa, Taivalkoski Jurmu-Kurtti, Koivuoja, pine dominated forest with some <italic>Picea</italic> and <italic>Betula</italic>, by a forest track, 31 Aug. 2008, <italic>I. Kytövuori</italic> 08-584, H6033533 (H); Inarin Lappi, Inari, N side of the river Lutto, N side of the road to Rajajooseppi, Keskikompsio (Hätäpuhelin), above the river bank, mesic heath forest with swamp depressions, <italic>Pinus</italic>,<italic> Betula</italic> and <italic>Salix</italic> spp., 14 Aug. 1995, <italic>I. Kytövuori</italic> 95-382, H6032729 (H).</p><p>Notes — <italic>Cortinarius balteaticlavatus</italic> belongs to /Balteatoalbi together with <italic>C. balteatoalbus</italic> Rob. Henry and <italic>C. balteatialutaceus</italic>. In ITS regions it differs by 10 substitutions and indel positions from <italic>C. balteatoalbus</italic> and 7 substitutions and indel positions from <italic>C. balteatialutaceus</italic>. The sequences of <italic>C. balteaticlavatus</italic> are identical. Typical for the species of the clade, compared to many other species of /Phlegmacioides, are basidiomata without bluish tints and narrow, rather small spores. <italic>Cortinarius balteaticlavatus</italic> grows in mixed forests in boreal zones. It is a stout species that resembles <italic>C. crassus </italic>Fr. The latter, however, has narrower spores (7–9 × 3.5–4.5 μm), abundant cylindrical to clavate cystidia especially at the lamellar edge, and no <italic>KOH-reaction</italic> in context. <italic>Cortinarius balteatoalbus</italic> differs from <italic>C. balteaticlavatus</italic> by its lower and wider appearance, larger spores, and more incrusted pileipellis hyphae. <italic>Cortinarius balteatialutaceus</italic> has a relatively shorter stipe, whitish to very pale brown pileus margin, and larger, moderately to fairly strongly verrucose spores.</p><p><bold><italic>Cortinarius brunneiaurantius</italic></bold> Kytöv., Liimat. & Niskanen, <italic>sp. nov</italic>. — MycoBank MB805880; <xref ref-type="fig" rid="F6">Fig. 6</xref>g, <xref ref-type="fig" rid="F7">7</xref>d</p><p><italic>Etymology</italic>. The name refers to the colour of the pileus.</p><p><italic>Type</italic>. F<sc>INLAND</sc>, Varsinais-Suomi, Turku, Ruissalo, Kansanpuisto, <italic>Tilia</italic> alley, also <italic>Quercus robur</italic> nearby, on clayey-mull soil, 22 Sept. 2001, <italic>J. Vauras</italic> 17979 (holotype H6032422; isotype TUR171972). GenBank KF732600.</p><p><italic>Pileus</italic> 4–9 cm broad, hemispherical to plano-convex, at first viscid but soon dry, centre with small, appressed scales, innately fibrillose toward margin, pale ochraceous brown. <italic>Lamellae</italic> emarginate, crowded, bluish to pale grey when young, later pale brown. <italic>Stipe</italic> 4–9 cm long, 0.8–1.5 cm thick at apex, 1.2–2 cm wide at base, clavate, first whitish, later pale brown. <italic>Universal veil</italic> white, soon brownish, rather sparse. <italic>Context</italic> white in the pileus, bluish at least in the upper part of the stipe. <italic>Odour</italic> not recorded. <italic>KOH-reaction</italic> in context yellow. <italic>Exsiccata</italic>: pileus orange brown to brown at the centre, paler yellowish brown at the margin, stipe pale brown.</p><p>In MLZ: <italic>Spores</italic> 8.2–<italic>8.9</italic>–9.7 × 4.8–<italic>5.2</italic>–5.4 μm, av. = 8.9 × 5.2 μm, Q = 1.58–<italic>1.71</italic>–1.83, Qav. = 1.70–1.71 (2 specimens, 100 spores, <xref ref-type="fig" rid="F7">Fig. 7</xref>d), amygdaloid to citriform, with a shallow suprahilar depression, moderately to fairly strongly verrusose, warts dark, moderately dextrinoid, often very dark. <italic>Basidia</italic> 30–38 × 7–8 μm, 4-spored, clavate, sand brown, with very small granules. <italic>Lamellar trama hyphae</italic> with dark small granules and blood red substance or large granules and chips on greenish greyish ground colour. <italic>Stipe apex hyphae</italic> bright yellow, some with granules and chips, outermost ones yellow to reddish brown with blood blackish contents. <italic>Pileipellis </italic>simplex, in overall view orange red, uppermost hyphae 4–8 μm wide, with clearly discernible spirally incrusted wall and some dark granule mounds inside. Lower hyphae wider, more yellow, almost smooth, with abundant dark granular substance. Hypoderm absent.</p><p>Ecology & Distribution — In deciduous forests, presumably with <italic>Quercus</italic>,<italic> Corylus</italic> and <italic>Populus</italic>. To date only known from South Finland. Fruits in autumn. No sequences of the species exist in the public databases.</p><p><italic>Other specimens examined</italic>. F<sc>INLAND</sc>, Uusimaa, Espoo, Nuuksio, Pirttimäki, fairly rich grass-herb forest with <italic>Populus tremula</italic>,<italic> Betula</italic>,<italic> Alnus incana</italic>,<italic> Quercus</italic>,<italic> Corylus</italic>,<italic> Prunus padus</italic>,<italic> Salix</italic> spp., and some old pines and young spruces, 2 Sept. 1993, <italic>I. Kytövuori</italic> 93-644, H6032406 (H).</p><p>Notes — <italic>Cortinarius brunneiaurantius</italic> belongs to /Phlegmacioides and forms a well supported group (0.99 PP) with <italic>C. sobrius</italic> P. Karst.,<italic> C. balteatus</italic> (Fr.) Fr.,<italic> C. brunneoviolaceus</italic> Bidaud,<italic> C. pseudonaeovosus</italic> Rob. Henry and <italic>C. clarobaltoides</italic> var. <italic>longispermus</italic> Reumaux. The ITS sequences of <italic>C. brunneiaurantius</italic> are identical and differ from the ones of the sister species <italic>C. sobrius</italic> by 9 substitutions and indel positions. Morphologically, it does not resemble any of its closest relatives. Typical for the species are pale ochraceous brown pileus, rather small, amygdaloid spores, positive granular MLZ reaction, and habitat with deciduous trees.</p><p><bold><italic>Cortinarius caesiocolor</italic></bold> Kytöv., Liimat. & Niskanen, <italic>sp. nov</italic>. — MycoBank MB805881; <xref ref-type="fig" rid="F6">Fig. 6</xref>h, <xref ref-type="fig" rid="F7">7</xref>e</p><p><italic>Etymology</italic>. The name refers to the colour of the pileus.</p><p><italic>Type</italic>. F<sc>INLAND</sc>, Uusimaa, Lohja, Jalassaari, Tamminiemi, by a track with <italic>Betula</italic>,<italic> Populus tremula</italic>,<italic> Quercus</italic>,<italic> Corylus</italic> and <italic>Salix caprea</italic>, on calcareous soil, 27 Aug. 2000, <italic>I. Kytövuori</italic> 00-029 (holotype H; isotype NY). GenBank KF732603.</p><p><italic>Pileus</italic> 5–9 cm broad, hemispherical to convex, then plano-convex, slightly viscid, innately fibrillose, more so toward margin, bluish violet to violet brown. <italic>Lamellae</italic> emarginate, crowded, first pale greyish brown with a bluish tint, later pale brown to brown. <italic>Stipe</italic> 6–10 cm long, 1–2 cm thick at apex, 2–3.5 cm wide at base, clavate, whitish, with a bluish tint at the apex. <italic>Universal veil</italic> first pale blue, later becoming brown, sparse. <italic>Context </italic>blue in all parts or partially whitish but blue at least close to lamellae. <italic>Odour</italic> indistinct. <italic>KOH-reaction</italic> in context yellow. <italic>Exsiccata</italic>: pileus pale greyish brown to violet brown, stipe pale greyish brown, brown at base.</p><p>In MLZ: <italic>Spores</italic> 9.1–<italic>9.9</italic>–10.9 × 5.4–<italic>5.8</italic>–6.3 μm, av. = 9.8–9.9 × 5.7–5.9 μm, Q = 1.59–<italic>1.70</italic>–1.84, Qav. = 1.67–1.72 (2 specimens, 120 spores, <xref ref-type="fig" rid="F7">Fig. 7</xref>e), weakly citriform, sometimes with a low suprahilar depression, dark-coloured, moderately dextrinoid, fairly strongly verrucose, warts anastomosing, dark. <italic>Basidia</italic> 30–41 × 7.5–9.0 μm (40 basidia), 4-spored, clavate, reddish sand brown, mostly with some blood red particles. <italic>Lamellar trama hyphae</italic> yellow to orange yellowish, with small granules. <italic>Stipe apex hyphae</italic> yellow to orange yellow, with small granules, the outermost ones fairly red, with very small blood red granules.<italic> Velum/cortina hyphae</italic> fairly red, with very small blood red granules. <italic>Pileipellis</italic>: epicutis weakly gelatinous, uppermost hyphae narrow, 2–4 μm wide, ochraceous brown, spirally incrusted, mostly not granulose, mostly without large redbrown particles, lower hyphae up to 8 μm wide, mostly incrusted, somewhat granulose with small, brown granules. Hypoderm absent.</p><p>Ecology & Distribution — Under <italic>Quercus</italic>,<italic> Populus</italic> and <italic>Corylus</italic> on mull soil in parks, roadsides and deciduous forests. Known from southern Finland. No sequences of this species exist in public databases.</p><p><italic>Other specimen examined</italic>. F<sc>INLAND</sc>, Uusimaa, Helsinki, the cemetery of Malmi, <italic>Quercus robur</italic> alley, on cut meadow, 17 Aug. 1997, <italic>I. Kytövuori</italic> 97-207, H6032730 (H).</p><p>Notes — <italic>Cortinarius caesiocolor</italic> is a typical member of /Phlegmacioides. Typical for the species are bluish to violet pileus and upper part of the stipe (appearance somewhat like <italic>C. porphyropus</italic>), small spores, and habitat with deciduous trees. The sister species <italic>C. chromataphilus</italic> Rob. Henry and more distantly related <italic>C. largus</italic> Fr., which also grow in deciduous forests, are at pileus centre greyish brown, towards margin greyish blue, and have larger spores (spores of <italic>C. chromataphilus</italic> 10.5–13 × 6–6.5 μm, of <italic>C. largus</italic> 10.0–11.8 × 6.1–8.8 μm). Another small-spored species with bluish colours is <italic>C. violaceomaculatus</italic> Brandrud, but it has smaller (8.8–10.2 × 5.2–5.7 μm), more weakly verrucose spores, and grows with conifers. <italic>Cortinarius caesiocolor</italic> formed a well-supported clade in our phylogenetic analysis and differs in ITS regions from <italic>C. chromataphilus</italic> by 16 substitutions and indel positions. The sequences of <italic>C. caesiocolor </italic>have three base and one length polymorphisms and the maximum pairwise distance is zero.</p><p><bold><italic>Cortinarius myrtilliphilus</italic></bold> Kytöv., Liimat., Niskanen & Brandrud, <italic>sp. nov</italic>. — MycoBank MB805882; <xref ref-type="fig" rid="F7">Fig. 7</xref>f</p><p><italic>Etymology</italic>. The name refers to the habitat with <italic>Vaccinium myrtillus</italic>.</p><p>= <italic>Cortinarius vacciniophilus</italic> Brandrud, Edinburgh J. Bot. 54, 1: 114. 1997. p.p.</p><p><italic>Type</italic>. F<sc>INLAND</sc>, Kainuu, Suomussalmi, Suolijärvi, Siikajärvi, NW foot of Siikavaara (Rönninvaara), Pöksäkorpi, gently sloping, partly swampy, grass-herb-spruce forest with some <italic>Pinus</italic>,<italic> Betula</italic>,<italic> Populus</italic>,<italic> Alnus incana</italic> and <italic>Salix</italic> spp., 14 Sept. 1997, <italic>I. Kytövuori</italic> 97-1469, H6032751 (holotype H; isotype NY). GenBank KF732605.</p><p><italic>Pileus</italic> 4.5–9(–11) cm broad, hemispherical to plano-convex, with slightly incurved margin, weakly to distinctly viscid when young, soon dry, with appressed veil scales, especially at centre, pale yellowish brown to darker ochraceous brown to leather brown (like <italic>C. balteatus</italic>), young margin paler, almost whitish; sometimes with hygrophanous streaks. <italic>Lamellae</italic> emarginate, crowded, greyish white when young, later pale brown. <italic>Stipe</italic> 5.5–11 cm long, 1–2(–2.5) cm thick at apex, 1.5–3 cm wide at base, clavate, at first white, later becoming pale brown from base. <italic>Universal veil</italic> white, sparse. <italic>Context</italic> white, faintly greyish hygrophanous at stipe apex. <italic>Odour</italic> indistinct to weak, pleasant, resembling corn. <italic>KOH-reaction</italic> in context strongly yellow when young, later weakly yellow or with yellow margin. <italic>Exsiccata</italic>: pileus uniformly pale warm yellowish brown to brown or somewhat darker at the middle, stipe with the same tint as pileus but lighter.</p><p>In MLZ: <italic>Spores</italic> 10.9–<italic>12.1</italic>–13.4 × 6.3–<italic>6.9</italic>–7.5 μm, Q = 1.63–<italic>1.79</italic>–1.91 (2 specimens, 100 spores, <xref ref-type="fig" rid="F7">Fig. 7</xref>f), amygdaloid-fusoid with a distinct suprahilar depression and a long apical part (in form very much like those in <italic>C. collinitus</italic>), (finely to) moderately verrucose, somewhat dextrinoid. <italic>Basidia</italic> 33–45 × 8.5–11 μm (40 basidia), sand brown, dark granules very few. <italic>Lamellar trama hyphae</italic> with small granules. <italic>Stipe apex hyphae</italic>: pale brown, outermost hyphae more reddish, dark granules few. <italic>Pileipellis</italic>: gelatinous layer present, erect-sinuose gelatinous hyphae 4–6(–8) μm wide, repent subsurface hyphae 5–10 μm wide, fairly thin-walled, finely to fairly strongly spirally incrusted, with some dark red brown, angular particles. Lower hyphae with slightly thicker, smooth to strongly and coarsely incrusted hyphae with dark red brown, farinose to granulose contents.</p><p>Ecology & Distribution — In mesic <italic>Picea</italic> dominated forests, from richer to poor <italic>Vaccinium myrtillus</italic> type. So far only known from middle boreal zone from Finland and Norway and might be rare. Produces fruitbodies in autumn.</p><p><italic>Other specimens examined</italic>. N<sc>ORWAY</sc>, Oppland, Ringebu, Venabygd, c. 700 m asl, <italic>Picea abies</italic> forest, ± poor raw humus, mossy <italic>Vaccinium myrtillus</italic> type, 30 Aug. 1994, <italic>E. Bendiksen & T.E. Brandrud</italic>, TEB 4-94, O 125949 (O, sub nom. <italic>C. vacciniophilus</italic>).</p><p>Notes — <italic>Cortinarius myrtilliphilus</italic> belongs to /Phlegmacioides. It has an ochraceous brown pileus and pale greyish white lamellae, both without bluish tints, large spores, and it grows in mesic <italic>Picea</italic> dominated forests. It has previously been confused with <italic>C. pseudonaevosus</italic> Rob. Henry (= <italic>C. acidophilus</italic> Brandrud) which is a common spruce forest species in northern Europe and also occurs in mountain areas of Central Europe. Typically, <italic>C. pseudonaevosus</italic> has bluish pileus margin, bluish tints in context of the pileus and stipe apex, and smaller spores. However, <italic>C. myrtilliphilus</italic> strongly resembles the large-spored and non-bluish specimens of <italic>C. pseudonaevosus</italic>. Previously, these variants were called <italic>C. vacciniophilus</italic> Brandrud (<xref rid="R15" ref-type="bibr">Brandrud 1998</xref>). These large-spored <italic>C. vacciniophilus</italic> specimens are in some regions as frequent as the smaller spored <italic>C. acidophilus</italic> specimens (<xref rid="R41" ref-type="bibr">Gjerde et al. 2012</xref>). Our molecular studies showed that a majority of the large-spored specimens are in fact <italic>C. pseudonaevosus</italic>, leaving <italic>C. pseudonaevosus</italic> as an apparently genetic uniform but morphologically heterogeneous species. On the other hand, analysis of ITS sequences also showed that <italic>C.</italic><italic>myrtilliphilus</italic> represents an autonomous species and <italic>C.</italic><italic>myrtilliphilus</italic> and <italic>C. pseudonaevosus</italic> are not closely related. <italic>Cortinarius pseudonaevosus</italic> belongs to a strongly supported clade (0.99 PP) with <italic>C. balteatus</italic> (Fr.) Fr.,<italic> C. brunneiaurantius</italic> Kytöv., Liimat. & Niskanen,<italic> C. brunneoviolaceus</italic> Bidaud,<italic> C. sobrius</italic> and <italic>C. clarobaltoides </italic>var.<italic> longispermus</italic> Reumaux, while <italic>C. myrtilliphilus</italic> is placed elsewhere in /Phlegmacioides, but further relationships with the other members of the clade were not fully resolved. Based on the pairwise comparison of the ITS sequences the closest species is <italic>C. badiolatus</italic> (M.M. Moser) M.M. Moser (= <italic>C. durus</italic> s.auct.) from which it differs by 11 substitutions and indel positions. <italic>Cortinarius badiolatus</italic> differs from <italic>C. myrtilliphilus</italic> by paler, initially almost whitish colours of the pileus, and habitat mainly in subalpine <italic>Betula</italic> forests. Based on molecular and morphological data we conclude that <italic>C. myrtilliphilus</italic> represent an autonomous species and describe it here as new to science. <italic>Cortinarius myrtilliphilus</italic> must be a very rare species, by us only found twice during many years of study. Due to limited material, we do not fully know the morphological variation of <italic>C. myrtilliphilus</italic>, and the morphological overlap with the similar, but genetically quite distant, large-spored variants of <italic>C. pseudonaevosus</italic> (= <italic>C. vacciniophilus</italic> p.p.) needs further study.</p></sec><sec id="s14"><title>DISCUSSION</title><sec id="s14a"><title>Type studies</title><p>The majority of names published in the genus <italic>Cortinarius</italic> have been by French taxonomists, representing over half of the types we studied, 132 names representing 77 species, but for 17 of these an earlier name by other taxonomists exist, so in reality the number of truly new species they described is 60. They represent species from both broadleaf hardwood (about 60 %) and conifer forests. Thus, the <italic>Cortinarii</italic> of France are among the most extensively studied in the world. Although Friesian names often dominate the nomenclatural discussions in <italic>Cortinarius</italic>, the number of species he described is significantly less than the numbers for French authors. In <xref rid="R27" ref-type="bibr">Fries (1836–1838</xref>) there are about 50 names belonging to the groups we studied and of these 32 are currently included in Funga Nordica (<xref rid="R52" ref-type="bibr">Jeppesen et al. 2012</xref>). This demonstrates how difficult it is to interpret old names with vague descriptions and no type material. The Friesian names currently in use include mainly species for which either published or unpublished illustrations exists making them easier to interpret. Species with Friesian names represent about 20 % of the species recognized in this study. A similar trend occurs in subg. <italic>Telamonia</italic>. For example, in sections <italic>Armillati</italic> and <italic>Brunnei </italic>30 % of the total known species were described by Fries, and in the morphologically challenging sect. <italic>Bovini</italic> only 8 % (<xref rid="R70" ref-type="bibr">Niskanen et al. 2009</xref>, <xref rid="R71" ref-type="bibr">2011a</xref>, <xref rid="R72" ref-type="bibr">2013a</xref>).</p><p>From North America we studied 60 types, representing 59 species. Of these, 52 species (88 %) have apparently not been described from anywhere else in the world, while one of them is the older synonym for a European name (<italic>C. metarius</italic> Kauffman = <italic>C. barbarorum</italic> Bidaud, Moënne-Locc. & Reumaux). Thirty-five of these species were described from the mountains and coastal areas of western North America, 13 species are from the Rocky Mountains and 12 species were collected from eastern North America, mainly Michigan. Almost all the names that are synonyms of European species are associated with conifers, i.e., <italic>C. crassus</italic> Fr. (= <italic>C. subaustralis</italic> A.H. Sm. & Hesler, described from North Carolina), <italic>C. glaucopus </italic>(Schaeff.) Fr. (= <italic>C. glaucopoides</italic> Kauffman, described from Colorado) and <italic>C. metarius</italic>; the only exception is <italic>C. triumphans</italic> Fr. s.auct. (= <italic>C. ophiopus</italic> Peck from Maryland). Species described from North America and reported here for the first time from Europe, include <italic>C. olympianus </italic>(USA, Washington state and Finland), <italic>C. patrickensis </italic>(USA, California and Sweden) and <italic>C. sannio </italic>(USA, Wyoming, Finland and Sweden), all are conifer-associated species. These findings correlate well with previous studies by <xref rid="R35" ref-type="bibr">Garnica et al. (2011)</xref>, <xref rid="R43" ref-type="bibr">Harrower et al. (2011)</xref> and <xref rid="R72" ref-type="bibr">Niskanen et al. (2013a</xref>, <xref rid="R73" ref-type="bibr">b</xref>). Interestingly, many names described from Europe have been used in North America, whereas few names from North America have been applied to taxa in Europe.</p><p>Of the 154 species recognised in this study 114 of them have no synonyms, however, the remaining 40 species each have one or more synonyms. The species with a significant number of synonyms include <italic>C. largus</italic> (14), <italic>C. pseudonaevosus</italic> (6), <italic>C. talus</italic> (5), <italic>C. crassus</italic> (4) and <italic>C. varius</italic> (4), all represent rather common species. The authors whose names we studied have all described synonyms. This is in part due to the difficulty in determining which species already have been described, especially based only on morphology and ecology. The narrow species concept of Bidaud, Henry, Moënne-Loccoz and Remaux was not supported by our study nor were the broad species concepts of Fries, Brandrud and Jeppesen et al. (2012) as supported by the results of <xref rid="R32" ref-type="bibr">Frøslev et al. (2007)</xref> and <xref rid="R70" ref-type="bibr">Niskanen et al. (2009</xref>,
<xref rid="R71" ref-type="bibr">2011a</xref>).</p><p>In this paper, species names have been synonymised when both molecular and morphological data have supported it. Of course there is some risk that morphological differences have been overlooked by studying too few specimens. Based on the overall data we have on <italic>Cortinarius</italic> it would seem logical, however, that in the majority of cases the synonymization is correct. In doubtful cases, when the variation in ITS regions of a group of specimens has been more than 2 substitutions and/or indel positions and we have not had enough specimens to make further comparisons, we have left names unchanged, i.e. <italic>C. albescens/C. gentianeus/C. volvatus</italic>, <italic>C. herpeticus/C. violaceonitens</italic>, and <italic>C. misermontii/C. olidoamarus </italic>var.<italic> valentinus/ C.subaccedens/C. van-campiae.</italic> Also, the following species with higher intraspecific variation may in reality represent two to several species: <italic>C. aureofulvus </italic>s.auct. /<italic>C. orichalceus </italic>var.<italic> xanthocephalus</italic>, <italic>C. pallidirimosus</italic>, <italic>C. porphyropus</italic> and <italic>C. scaurus.</italic></p><p>Our study includes the majority of names described in subg.<italic>Phlegmacium</italic>, excluding sect. <italic>Riederi</italic> and parts of sections<italic> Calochroi </italic>and<italic> Fulvi </italic>already treated or to be treated by other taxonomists (e.g. <xref rid="R32" ref-type="bibr">Frøslev et al. 2007</xref>). In some instances, type material could not be acquired or found in herbaria, i.e. the material of C.H. Peck (NYSM) and some of the material of R. Henry (PC). In other cases they were not possible to study within the time frame of this study, i.e. the early names described by M. Moser (M). Peck published from 1870 to 1912 about 80 names in the genus <italic>Cortinarius</italic> and the names represent at least twenty species of subg. <italic>Phlegmacium</italic> s.lato based on morphological and microscopical features. The material deposited in München includes about 30 <italic>Phlegmacium</italic> species. For about 55 <italic>Phlegmacium</italic> species described by Henry the type material was not found and over 20 additional ones remain to be studied. Therefore, it is possible that some of these names have priority over currently applied <italic>Cortinarius</italic> names, including some of those in this paper. Our plan is to acquire Peck’s, the remaining materials of Moser’s and Henry’s, and perhaps others that we have overlooked in the near future and attempt to sequence them for comparison.</p></sec><sec id="s14b"><title>New species</title><p>The majority of new species described here represent boreal taxa, i.e. <italic>C. pallidirimosus</italic>,<italic> C. balteaticlavatus</italic> and <italic>C. myrtilliphilus</italic>, and/or are species that previously have been included within the morphological limits of other, ‘well-known’ taxa, these include <italic>C. boreicyanites</italic> and <italic>C. subfraudulosus</italic>. The large number of unknown, boreal species is not surprising, since the majority of existing names are described from hemiboreal and temperate zones or from the mountains of Central and southern Europe. Even though the <italic>Cortinarius</italic> species of Europe have been extensively studied, there undoubtedly remain a lot of taxa to be discovered, not to mention in other areas of the world, which are less explored.</p></sec><sec id="s14c"><title>Factors affecting on the success of molecular work</title><p>The success of the molecular studies was significantly influenced by the condition of type specimens, including the age of the specimen, how it was dried and/or preserved, and whether or not it was mouldy. Also, the length polymorphism of the ITS region caused some problems with direct sequencing. In general, <italic>Cortinarius</italic> is one of the easiest genera of <italic>Agaricales</italic> for molecular work. Often DNA can be extracted with regular methods even from old specimens. For example, we obtained a complete ITS region from the type material fo <italic>C. sobrius</italic> P. Karst. (collected in 1890) and <italic>C. virentophyllus </italic>Kauffman (collected in 1912). The success rate usually decreases with older specimens, however, there is some difference between the specimens collected by different authors. For example, we were not able to successfully sequence any of the specimens collected by T. Hongo during 1960–1968 (<xref rid="R71" ref-type="bibr">Niskanen et al. 2011a</xref>, and unpublished data of <italic>C. cinnamomeoides</italic> Hongo, <italic>C. neoarmillatus</italic> Hongo and <italic>C. nigrosquamosus</italic> Hongo) whereas the success rate for the specimens collected by A.H. Smith mainly during 1935–1941 was much better. This could be due to the drying process itself, for example, drying temperature, length of drying time, which is related to air circulation, and the condition of fresh specimens when placed on the dryer. In particular, the type collections of R. Henry proved challenging, since often the material was sparse and poorly labelled, and many were mouldy. In some instances types were missing, could not be located at the time they were requested, or were not available for study.</p><p>Several things can improve the success rate of problematic specimens. For example, methods that provided high quality DNA extraction, and PCR and sequencing machines improved the results. If the DNA is fragmented, amplifying and sequencing ITS1 and ITS2 regions separately was often helpful.</p></sec><sec id="s14d"><title>Recommendations for stabilizing the nomenclature</title><p>For achieving a stable and unambiguous use of names, the following requirements should be fulfilled whenever possible. Types of existing names should be studied. Even easily identifiable species like <italic>C. balteatoalbus</italic> can be misinterpreted, or a species may turn out to be a group of cryptic species, for example, <italic>C. dionysae </italic>and <italic>C. elegantior</italic> (<xref rid="R35" ref-type="bibr">Garnica et al. 2011</xref>). Taxonomists should not use names in barcoding databases unless they have been verified by molecular type studies. For old names that do not have type specimens or where the type specimen is in poor condition or not available because of historical preservation, a neotype, lectotype or epitype should be chosen. Names rejected due to the problems of interpretation should be excluded from consideration. If the name is not verified with a sequence from a type specimen its application will be uncertain and a source of confusion. A significant problem with neo-, lecto- and epitypes is the lack of a database for them. It is difficult to find out if a type already has been designated thus hindering nomenclatorial work.</p><p>New names should not be accepted for publication without the support of molecular data in genera where the amplification of the ITS or other sequences is easily done, thus preventing the description of synonyms and creating a sound reference database for further taxonomical and ecological studies. Finally, when studying types it is essential to mark the basidiomata from which the DNA has been extracted. Some collection might turn out to be mixed, and thus, knowing exactly which basidiocarp has been studied, will be necessary.</p><p>These recommendations may seem self-evident, but have been neglected in many cases. Often the reason is a lack of resources and/or knowledge. At least in <italic>Cortinarius</italic> the amount of already existing names is overwhelming and we need to proceed carefully from this point forward.</p></sec><sec id="s14e"><title>Infrageneric classification</title><p>The classification of <italic>Cortinarius</italic> is not yet stabilised and many traditional infrageneric groups have shown to be at least partly artificial. Therefore, when studying a certain group of species, it might be difficult to find all the relevant species described from literature. The aim of the phylogenetic estimates in this study was not to solve the infrageneric classification of <italic>Cortinarius</italic> but to show the preliminary placement of the studied species, which then could be used for guidance in further taxonomical studies.</p><p>In our phylogenetic analysis ten of the twelve clades representing phlegmacioid species sensu <xref rid="R38" ref-type="bibr">Garnica et al. (2005)</xref> were recovered: /Alluti (= Multiformes), /Amarescentes (= Infracti), /Arguti/Calochroi, /Percomes, /Phlegmacium, /Phlegmacioides, /Praestantes (= Claricolores), and /Scauri. In addition, /Glaucopodes was well supported in our analysis. Clades /Caerulescentes and /Vulpini were split.</p></sec><sec id="s14f"><title>Barcoding Cortinarius</title><p>This study provides ITS barcodes for 175 <italic>Cortinarius</italic> species. This data enables the identification of a majority of boreal and temperate species of <italic>Phlegmacia</italic>, except for sections <italic>Calochroi</italic>, <italic>Fulvi</italic>, for which part of the ITS sequences already are published (e.g. <xref rid="R32" ref-type="bibr">Frøslev et al. 2007</xref>) and <italic>Riederi</italic>, from Europe and parts of North America.</p><p>Of the 236 names we studied, only 61 were currently represented in GenBank. It is noteworthy that almost half of these names are Friesian names and a third of them are names published from North America. Only 4 % of the names described by French authors have made their way into general use in GenBank. This emphasizes the importance of type studies and the role of taxonomists in the creation of an identification database. If we do not produce sound basic data on <italic>Cortinarius </italic>species, it is difficult or impossible in many cases for non-taxonomists to identify specimens or environmental samples.</p><p>A barcoding database based on type studies is essential for ecological, environmental or further taxonomic research. Once completed, it will create a solid base for future studies. Barcoding is a powerful tool, which enables us to identify and compare species from different regions in a completely new way. This is especially true and important for fungi like <italic>Cortinarius</italic>, which are morphologically challenging and difficult to identify. Finally, we are able to reliably compare our knowledge on species from different areas. A base for <italic>Phlegmacia</italic> is created here and provides a beginning and direction for future studies in <italic>Cortinarius</italic>.</p></sec></sec></body><back><ack><p>We are grateful to the curators of C, G, H, IB, MICH, PC and S, and especially to Bart Buyck, Xavier Carteret, Philippe Clerc, Regina Kühner-Winkler, Ursula Peintner and Patricia Rogers. Anna-Lena Anderberg is also warmly thanked for providing photos of unpublished plates of Fries. Tor Erik Brandrud is thanked for valuable comments concerning the neo- and epitypifications and providing the photo of <italic>C. caesiophylloides</italic>. Andy Taylor and Dimitar Bojantchev are thanked for providing material and sequences for the study and Andy Taylor also for providing the photo of <italic>C. talimultiformis</italic>. Ertugrul Sesli is thanked for providing material of <italic>C. talimultiformis</italic>, Tapio Kekki for providing the photo of <italic>C. flavipallens</italic>, Jukka Vauras for providing the photo of <italic>C. brunneiaurantius</italic> and Karen Hughes for providing an additional sequence of <italic>C. acystidiosus</italic>. Bálint Dima and Tobias G. Frøslev thank members of the ‘J.E.C. <italic>Phlegmacium</italic> Research Group’ (Francesco Bellù, Tor Erik Brandrud, Bernhard Oertel, Günter Saar and Geert Schmidt-Stohn) and Thomas Stjernegaard Jeppesen, and László Albert for valuable taxonomic discussions during the past years. László Albert and Zoltán Lukács are thanked for making older literature available for study. 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Short ITS sequences excluded from the analysis marked with *.</p></caption><table frame="hsides" rules="groups"><thead><tr><th align="left" rowspan="1" colspan="1">Species</th><th align="left" rowspan="1" colspan="1">Voucher</th><th align="left" rowspan="1" colspan="1">Herb</th><th align="left" rowspan="1" colspan="1">Locality</th><th align="left" rowspan="1" colspan="1">Ecology</th><th align="left" rowspan="1" colspan="1">Current name</th><th align="left" rowspan="1" colspan="1">Collection date</th><th align="left" rowspan="1" colspan="1">Collector</th><th align="left" rowspan="1" colspan="1">GenBank number</th></tr></thead><tbody><tr><td align="left" rowspan="1" colspan="1"><italic>C. acidophilus</italic> Brandrud 1997 (holotype)</td><td align="left" rowspan="1" colspan="1">TEB61-79</td><td align="left" rowspan="1" colspan="1">O</td><td align="left" rowspan="1" colspan="1">Norway, Oppl, Lunner, Østäsen</td><td align="left" rowspan="1" colspan="1">In forest of <italic>Picea abies</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. pseudonaevosus</italic> Rob. Henry 1957</td><td align="left" rowspan="1" colspan="1">01.08.1979</td><td align="left" rowspan="1" colspan="1">T.E. Brandrud</td><td align="left" rowspan="1" colspan="1">KF732241</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. acystidiosus</italic> Thiers 1960 (holotype)*</td><td align="left" rowspan="1" colspan="1">1902</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Texas, San Jacinto Co., Sam Houston National Forest, Coldspring</td><td align="left" rowspan="1" colspan="1">In pine-hardwood forest</td><td align="left" rowspan="1" colspan="1"><italic>C. acystidiosus</italic> Thiers 1960</td><td align="left" rowspan="1" colspan="1">23.05.1953</td><td align="left" rowspan="1" colspan="1">H.D. Thiers</td><td align="left" rowspan="1" colspan="1">KF732242</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. aggregatus </italic>Kauffman 1918*</td><td align="left" rowspan="1" colspan="1">MICH10311</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Michigan, Jackson, Vandercook Park, Jackson</td><td align="left" rowspan="1" colspan="1">In low woods</td><td align="left" rowspan="1" colspan="1"><italic>C. aggregatus </italic>Kauffman 1918</td><td align="left" rowspan="1" colspan="1">09.09.1907</td><td align="left" rowspan="1" colspan="1">C.H. Kauffman</td><td align="left" rowspan="1" colspan="1">KF732243</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. albescens</italic> A.H. Sm. 1944 (holotype)</td><td align="left" rowspan="1" colspan="1">17522</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Washington, Olympic National Park, Olympic Hot Springs</td><td align="left" rowspan="1" colspan="1">Under conifers</td><td align="left" rowspan="1" colspan="1"><italic>C. albescens</italic> A.H. Sm. 1944</td><td align="left" rowspan="1" colspan="1">02.10.1941</td><td align="left" rowspan="1" colspan="1">A.H. Smith</td><td align="left" rowspan="1" colspan="1">KF732244</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. albofragrans</italic> Ammirati & M.M. Moser 1997 (holotype)</td><td align="left" rowspan="1" colspan="1">95/595</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">USA, California, Del Norte Co., Highway 199, Danger Point</td><td align="left" rowspan="1" colspan="1">Under <italic>Quercus</italic>, also evergreen oaks (<italic>Q. vacciniifolia</italic>, <italic>Q. chrysolepis</italic>, <italic>Lithocarpus densi-</italic><italic>flora</italic>) with some <italic>Pseudotsuga menziesii</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. albofragrans</italic> Ammirati & M.M. Moser 1997</td><td align="left" rowspan="1" colspan="1">20.11.1996</td><td align="left" rowspan="1" colspan="1">M. 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Sm. 1942 (holotype)</td><td align="left" rowspan="1" colspan="1">15381</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Michigan, Ann Arbor</td><td align="left" rowspan="1" colspan="1">Under butternut, walnut, and bassword on low ground along streams low ground along streams</td><td align="left" rowspan="1" colspan="1"><italic>C. amnicola</italic> A.H. Sm. 1942</td><td align="left" rowspan="1" colspan="1">15.09.1940</td><td align="left" rowspan="1" colspan="1">A.H. Smith</td><td align="left" rowspan="1" colspan="1">KF732249</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. anfractoides</italic> Rob. Henry & Trescol 1987 (holotype)*</td><td align="left" rowspan="1" colspan="1">RH1898</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France</td><td align="left" rowspan="1" colspan="1">In frondose forest under <italic>Quercus ilex</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. anfractoides</italic> Rob. Henry & Trescol 1987</td><td align="left" rowspan="1" colspan="1">11.1984</td><td align="left" rowspan="1" colspan="1">G. Chevassut & F. Trescol</td><td align="left" rowspan="1" colspan="1">KF732250</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. anfractoides</italic> var. <italic>cinereoclarus</italic> Bidaud 2009 (holotype)</td><td align="left" rowspan="1" colspan="1">AB08-09-155</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Ain, Cerin</td><td align="left" rowspan="1" colspan="1">In calcareous deciduous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. persoonianus</italic> Bidaud 2009</td><td align="left" rowspan="1" colspan="1">07.09.2008</td><td align="left" rowspan="1" colspan="1">A. Bidaud</td><td align="left" rowspan="1" colspan="1">KF732251</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. arenicola</italic> A.H. Sm. 1942 (holotype)</td><td align="left" rowspan="1" colspan="1">15315</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Michigan, Waterloo, Waterloo Project</td><td align="left" rowspan="1" colspan="1">Under <italic>Sassafras</italic> in dry sandy open woods</td><td align="left" rowspan="1" colspan="1"><italic>C. arenicola</italic> A.H. Sm. 1942</td><td align="left" rowspan="1" colspan="1">12.09.1940</td><td align="left" rowspan="1" colspan="1">A.H. Smith</td><td align="left" rowspan="1" colspan="1">KF732252</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. areni-silvae</italic> (Brandrud) Brandrud 2012 (holotype)</td><td align="left" rowspan="1" colspan="1">CFP461b</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Ång, Graninge, Viksmon</td><td align="left" rowspan="1" colspan="1">In boreal coniferous forest with <italic>Pinus</italic> and <italic>Picea</italic> on sandy soil</td><td align="left" rowspan="1" colspan="1"><italic>C. areni-silvae</italic> (Brandrud) Brandrud 2012</td><td align="left" rowspan="1" colspan="1">25.08.1986</td><td align="left" rowspan="1" colspan="1">T.E. Brandrud et al.</td><td align="left" rowspan="1" colspan="1">KF732253</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. argutipes</italic> Bidaud & Reumaux 1996 (holotype)*</td><td align="left" rowspan="1" colspan="1">713</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Creuse</td><td align="left" rowspan="1" colspan="1">Under <italic>Fagus</italic> on acid soil</td><td align="left" rowspan="1" colspan="1"><italic>C. chromataphilus</italic> Rob. Henry 1989</td><td align="left" rowspan="1" colspan="1">15.10.1987</td><td align="left" rowspan="1" colspan="1">D. Brion</td><td align="left" rowspan="1" colspan="1">KF732254</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. atrochalybaeus</italic> M.M. Moser & Ammirati 2000 (holotype)</td><td align="left" rowspan="1" colspan="1">95/630</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">USA, California, Del Norte Co., Highway 199, Smith River Middle Fork, Danger Point</td><td align="left" rowspan="1" colspan="1">Under <italic>Lithocarpus densiflora</italic>, <italic>Arbutus</italic><italic>menziesii</italic>, <italic>Quercus vacciniifolius</italic> and <italic>Q. chrysolepis</italic> on calcareous soil</td><td align="left" rowspan="1" colspan="1"><italic>C. atrochalybaeus</italic> M.M. Moser & Ammirati 2000</td><td align="left" rowspan="1" colspan="1">29.11.1995</td><td align="left" rowspan="1" colspan="1">J. Ammirati</td><td align="left" rowspan="1" colspan="1">KF732255</td></tr><tr><td align="left" rowspan="1" colspan="1">C. <italic>aurantionapus</italic> Bidaud & Reumaux 2006 (holotype)*</td><td align="left" rowspan="1" colspan="1">PML4893</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Drôme, Romeyer</td><td align="left" rowspan="1" colspan="1">Under <italic>Picea</italic> and <italic>Pinus sylvestris</italic> on calcareous soil</td><td align="left" rowspan="1" colspan="1"><italic>C. talus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">09.10.1992</td><td align="left" rowspan="1" colspan="1">J. Garin</td><td align="left" rowspan="1" colspan="1">KF732256</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. aurantionapus</italic> var. <italic>similis</italic> Moënne-Locc. 2006 (holotype)</td><td align="left" rowspan="1" colspan="1">PML883</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Haute-Savoie, Avernioz</td><td align="left" rowspan="1" colspan="1">Under <italic>Picea</italic> on calcareous soil</td><td align="left" rowspan="1" colspan="1"><italic>C. talimultiformis </italic>Kytöv., Liimat., Nis-kanen, A.F.S. Taylor & Sesli sp. nov.</td><td align="left" rowspan="1" colspan="1">06.06.1988</td><td align="left" rowspan="1" colspan="1">P. Moënne-Loccoz</td><td align="left" rowspan="1" colspan="1">KF732257</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. aurantiopallidus</italic> Bidaud 2006 (holotype)</td><td align="left" rowspan="1" colspan="1">AB05-11-404</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Ardèche, Lagorce</td><td align="left" rowspan="1" colspan="1">Under <italic>Quercus ilex</italic> on calcareous soil</td><td align="left" rowspan="1" colspan="1"><italic>C. aurantiopallidus</italic> Bidaud 2006</td><td align="left" rowspan="1" colspan="1">11.11.2005</td><td align="left" rowspan="1" colspan="1">A. Bidaud</td><td align="left" rowspan="1" colspan="1">KF732258</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. badiolatus</italic> (M.M. Moser) M.M. Moser 1967 (holotype)*</td><td align="left" rowspan="1" colspan="1">53/10</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">Germany, Schwenningen</td><td align="left" rowspan="1" colspan="1">In coniferous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. badiolatus</italic> (M.M. Moser) M.M. Moser 1967</td><td align="left" rowspan="1" colspan="1">28.08.1953</td><td align="left" rowspan="1" colspan="1">H. Haas</td><td align="left" rowspan="1" colspan="1">KF732259</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. balteatialutaceus </italic>Kytöv., Liimat. & Niskanen sp. nov. (holotype)</td><td align="left" rowspan="1" colspan="1">IK09-751</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Sweden, Jmt, Frostviken, Jormlien, Säterklumpen</td><td align="left" rowspan="1" colspan="1">In <italic>Betula</italic> forest with solitary <italic>Picea</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. balteatialutaceus </italic>Kytöv., Liimat. & Niskanen sp. nov.</td><td align="left" rowspan="1" colspan="1">09.07.2009</td><td align="left" rowspan="1" colspan="1">P. & I. Kytövuori</td><td align="left" rowspan="1" colspan="1">KF732586</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. balteatibulbosus</italic> Kytöv., Niskanen, Liimat., Bojantchev & A.F.S. Taylor sp. nov. (holotype)</td><td align="left" rowspan="1" colspan="1">IK98-1624 (H6033539)</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, U, Espoo, Nuuksio, Pirttimäki</td><td align="left" rowspan="1" colspan="1">In half-open cut meadow</td><td align="left" rowspan="1" colspan="1"><italic>C. balteatibulbosus</italic> Kytöv., Niskanen, Liimat., Bojantchev & A.F.S. Taylor sp. nov.</td><td align="left" rowspan="1" colspan="1">04.09.1998</td><td align="left" rowspan="1" colspan="1">I. Kytövuori</td><td align="left" rowspan="1" colspan="1">KF732589</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. balteaticlavatus</italic> Kytöv., Liimat. & Niskanen sp. nov. (holotype)</td><td align="left" rowspan="1" colspan="1">IK96-595 (H6032412)</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, PH, Virrat, Hauhuu, Sikosaari, Salmela</td><td align="left" rowspan="1" colspan="1">On the grassy roadside with young <italic>Picea</italic>, <italic>Betula</italic>, <italic>Populus tremula</italic>, <italic>Alnus incana</italic> and <italic>Salix</italic> spp.</td><td align="left" rowspan="1" colspan="1"><italic>C. balteaticlavatus</italic> Kytöv., Liimat. & Niskanen sp. nov.</td><td align="left" rowspan="1" colspan="1">23.08.1996</td><td align="left" rowspan="1" colspan="1">I. Kytövuori</td><td align="left" rowspan="1" colspan="1">KF732596</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. balteatoalbus </italic>Rob. Henry 1985 (isotype)*</td><td align="left" rowspan="1" colspan="1">RH82.98</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France</td><td align="left" rowspan="1" colspan="1">In submontane <italic>Picea</italic> forest</td><td align="left" rowspan="1" colspan="1"><italic>C. balteatoalbus</italic> Rob. Henry 1985</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">KF732260</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. balteatotomentosus</italic> Rob. Henry ex Rob. Henry 1985 (holotype)</td><td align="left" rowspan="1" colspan="1">RH306</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Doubs, Boujaille</td><td align="left" rowspan="1" colspan="1">In montane forest (mostly) of <italic>Picea abies</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. balteatus</italic> (Fr.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">KF732261</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. balteatus</italic> (Fr.) Fr. 1838 (neotype)</td><td align="left" rowspan="1" colspan="1">CFP940</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Ång, Säbrå, Överdal</td><td align="left" rowspan="1" colspan="1">Under <italic>Picea</italic> in cultivated area</td><td align="left" rowspan="1" colspan="1"><italic>C. balteatus</italic> (Fr.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">03.08.1990</td><td align="left" rowspan="1" colspan="1">T.E. Brandrud et al.</td><td align="left" rowspan="1" colspan="1">KF732262</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. balteatus</italic> var. <italic>praestantoides</italic> Reumaux 1996 (holotype)</td><td align="left" rowspan="1" colspan="1">760</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Ile-de-France, Forêt de Rambouillet, Etang d’Or</td><td align="left" rowspan="1" colspan="1">In shrubs in <italic>Quercus</italic> forest</td><td align="left" rowspan="1" colspan="1"><italic>C. flavescentipes</italic> Reumaux 1996</td><td align="left" rowspan="1" colspan="1">23.10.1982</td><td align="left" rowspan="1" colspan="1">P. Reumaux</td><td align="left" rowspan="1" colspan="1">KF732263</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. barrentium</italic> Poirier & Reumaux 1993 (holotype)*</td><td align="left" rowspan="1" colspan="1">2398</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Loiret, Arboretum des Barres</td><td align="left" rowspan="1" colspan="1">Under conifers</td><td align="left" rowspan="1" colspan="1"><italic>C. barrentium</italic> Poirier & Reumaux 1993</td><td align="left" rowspan="1" colspan="1">13.11.1991</td><td align="left" rowspan="1" colspan="1">J. Poirier</td><td align="left" rowspan="1" colspan="1">KF732264</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. bigelowii</italic> Thiers & A.H. Sm. 1969 (holotype)</td><td align="left" rowspan="1" colspan="1">45385</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Idaho, Seven Devils Mts, Heaven’s Gate Ridge</td><td align="left" rowspan="1" colspan="1">Under conifers</td><td align="left" rowspan="1" colspan="1"><italic>C. bigelowii</italic> Thiers & A.H. Sm. 1969</td><td align="left" rowspan="1" colspan="1">26.07.1954</td><td align="left" rowspan="1" colspan="1">H.E. Bigelow</td><td align="left" rowspan="1" colspan="1">KF732265</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. boreicyanites </italic>Kytöv., Liimat., Niskanen & A.F.S. Taylor sp. nov. (holotype)</td><td align="left" rowspan="1" colspan="1">CFP931</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Jmt, Ragunda, Böle</td><td align="left" rowspan="1" colspan="1">In birch forest on rich ground (<italic>Betula</italic>,<italic> Picea</italic>)</td><td align="left" rowspan="1" colspan="1"><italic>C. boreicyanites </italic>Kytöv., Liimat., Niskanen & A.F.S. Taylor sp. nov.</td><td align="left" rowspan="1" colspan="1">07.24.1990</td><td align="left" rowspan="1" colspan="1">T.E. Brandrud et al.</td><td align="left" rowspan="1" colspan="1">KF732296</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. boreidionysae </italic>Kytöv., Liimat., Niskanen & Dima sp. nov. (holotype)</td><td align="left" rowspan="1" colspan="1">IK97-1220</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, PeP, Tervola, Peura, Raemäki</td><td align="left" rowspan="1" colspan="1">In grass-herb-spruce forest with spring-fed depressions, on calcareous ground</td><td align="left" rowspan="1" colspan="1"><italic>C. boreidionysae </italic>Kytöv., Liimat., Niskanen & Dima sp. nov.</td><td align="left" rowspan="1" colspan="1">11.09.1997</td><td align="left" rowspan="1" colspan="1">I. Kytövuori</td><td align="left" rowspan="1" colspan="1">KF732488</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. borgsjoeensis</italic> Brandrud 1992 (isotype)</td><td align="left" rowspan="1" colspan="1">CFP728</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Jmt, Ragunda, Kullstabodarna</td><td align="left" rowspan="1" colspan="1">In herbaceous spruce forest</td><td align="left" rowspan="1" colspan="1"><italic>C. borgsjoeensis</italic> Brandrud 1992</td><td align="left" rowspan="1" colspan="1">31.08.1988</td><td align="left" rowspan="1" colspan="1">T.E. Brandrud et al.</td><td align="left" rowspan="1" colspan="1">KF732266</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. brunneiaurantius </italic>Kytöv., Liimat. & Niskanen sp. nov. (holotype)</td><td align="left" rowspan="1" colspan="1">JV17979 (H6032422)</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, V, Turku, Ruissalo, Kansanpuisto</td><td align="left" rowspan="1" colspan="1"><italic>Tilia</italic> alley, also <italic>Quercus robur</italic> nearby, on clayey-mull soil</td><td align="left" rowspan="1" colspan="1"><italic>C. brunneiaurantius </italic>Kytöv., Liimat. & Niskanen sp. nov.</td><td align="left" rowspan="1" colspan="1">22.09.2001</td><td align="left" rowspan="1" colspan="1">J. Vauras</td><td align="left" rowspan="1" colspan="1">KF732600</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. brunneolividus</italic> Bidaud 1996 (holotype)</td><td align="left" rowspan="1" colspan="1">3734</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Isère, Optevoz</td><td align="left" rowspan="1" colspan="1">In calcareous deciduous forest of <italic>Quercus</italic> and <italic>Carpinus</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. brunneolividus</italic> Bidaud 1996</td><td align="left" rowspan="1" colspan="1">14.09.1993</td><td align="left" rowspan="1" colspan="1">C. Blanc</td><td align="left" rowspan="1" colspan="1">KF732268</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. brunneoviolaceus</italic> Bidaud 1996 (holotype)</td><td align="left" rowspan="1" colspan="1">2951</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Isère, Arzay</td><td align="left" rowspan="1" colspan="1">Under <italic>Castanea</italic> on acid soil</td><td align="left" rowspan="1" colspan="1"><italic>C. brunneolividus</italic> Bidaud 1996</td><td align="left" rowspan="1" colspan="1">01.10.1992</td><td align="left" rowspan="1" colspan="1">A. Bidaud</td><td align="left" rowspan="1" colspan="1">KF732269</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cacodes</italic> M.M. Moser & Ammirati 2000 (holotype)</td><td align="left" rowspan="1" colspan="1">91/618</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">USA, California, Mendocino, Russian Gulch State Park</td><td align="left" rowspan="1" colspan="1">In mixed coniferous forest with <italic>Tsuga</italic>, <italic>Pseudotsuga</italic>, <italic>Abies</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. cacodes</italic> M.M. Moser & Ammirati 2000</td><td align="left" rowspan="1" colspan="1">30.11.1991</td><td align="left" rowspan="1" colspan="1">M. Moser</td><td align="left" rowspan="1" colspan="1">KF732270</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. caerulescens</italic> (Schaeff.) Fr. 1838 (epitype)</td><td align="left" rowspan="1" colspan="1">CFP853</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Belgium, Brabant, Tervuren</td><td align="left" rowspan="1" colspan="1">In beech forest on calcareous ground</td><td align="left" rowspan="1" colspan="1"><italic>C. caerulescens</italic> (Schaeff.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">23.09.1989</td><td align="left" rowspan="1" colspan="1">T.E. Brandrud et al.</td><td align="left" rowspan="1" colspan="1">KF732271</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. caesiocinctus </italic>Kühner 1989 (holotype)</td><td align="left" rowspan="1" colspan="1">57-13</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Haute-Savoie, Le Môle</td><td align="left" rowspan="1" colspan="1">In mixed forest with <italic>Fagus</italic> and <italic>Picea</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. caesiocinctus</italic> Kühner 1989</td><td align="left" rowspan="1" colspan="1">30.08.1957</td><td align="left" rowspan="1" colspan="1">R. Kühner</td><td align="left" rowspan="1" colspan="1">DQ663239</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. caesiocolor </italic>Kytöv., Liimat. & Niskanen sp. nov. (holotype)</td><td align="left" rowspan="1" colspan="1">IK00-029</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, U, Lohja, Jalassaari, Tamminiemi, by a track</td><td align="left" rowspan="1" colspan="1">With <italic>Betula</italic>, <italic>Populus tremula</italic>, <italic>Quercus</italic>, <italic>Corylus</italic> and <italic>Salix caprea</italic>, on calcareous ground</td><td align="left" rowspan="1" colspan="1"><italic>C. caesiocolor</italic> Kytöv., Liimat. & Niskanen sp. nov.</td><td align="left" rowspan="1" colspan="1">27.08.2000</td><td align="left" rowspan="1" colspan="1">I. Kytövuori</td><td align="left" rowspan="1" colspan="1">KF732603</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. caesiophylloides</italic> Kytöv., Liimat., Niskanen, Brandrud & Frøslev sp. nov. (holotype)</td><td align="left" rowspan="1" colspan="1">TN05-016 (H6029792)</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, ES, Joutsa, Koivuranta</td><td align="left" rowspan="1" colspan="1">In fairly young, mesic to damp, <italic>Picea abies</italic> dominated forest with some <italic>Betula</italic> and</td><td align="left" rowspan="1" colspan="1"><italic>C. caesiophylloides</italic> Kytöv., Liimat., Niskanen, Brandrud & Frøslev sp. nov.</td><td align="left" rowspan="1" colspan="1">30.08.2005</td><td align="left" rowspan="1" colspan="1">K.Liimatainen & T. Niskanen</td><td align="left" rowspan="1" colspan="1">KF732572</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. calojanthinus</italic> M.M. Moser & Ammirati 1999 (holotype)</td><td align="left" rowspan="1" colspan="1">97/220</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">USA, Wyoming, Teton National Forest, Calypso Creek, Flagstaff Road</td><td align="left" rowspan="1" colspan="1">In subalpine forest under <italic>Picea engelmannii</italic>, <italic>Abies lasiocarpa</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. calojanthinus</italic> M.M. Moser & Ammirati 1999</td><td align="left" rowspan="1" colspan="1">21.08.1997</td><td align="left" rowspan="1" colspan="1">M. Moser & J. Ammirati</td><td align="left" rowspan="1" colspan="1">KF732272</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. calyptratus</italic> A.H. Sm. 1939 (holotype)</td><td align="left" rowspan="1" colspan="1">8352</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, California, Crescent City</td><td align="left" rowspan="1" colspan="1">Under mixed spruce and redwood, pre-sumably also <italic>Lithocarpus</italic> and<italic> Quercus</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. calyptratus</italic> A.H. Sm. 1939</td><td align="left" rowspan="1" colspan="1">03.11.1937</td><td align="left" rowspan="1" colspan="1">A.H. Smith</td><td align="left" rowspan="1" colspan="1">KF732273</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. calyptrodermus</italic> A.H. Sm. 1942 (holotype)</td><td align="left" rowspan="1" colspan="1">15356</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Michigan, Sharron Hollow</td><td align="left" rowspan="1" colspan="1">In low woods of second growth oak and basswood (<italic>Tilia</italic>) and various shrubs</td><td align="left" rowspan="1" colspan="1"><italic>C. calyptrodermus</italic> A.H. Sm. 1942</td><td align="left" rowspan="1" colspan="1">14.09.1940</td><td align="left" rowspan="1" colspan="1">A.H. Smith</td><td align="left" rowspan="1" colspan="1">KF732274</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. castaneicolor</italic> A.H. Sm. 1944 (holotype)</td><td align="left" rowspan="1" colspan="1">17926</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Washington, Olympic National Park, Olympic Hot Springs</td><td align="left" rowspan="1" colspan="1">Under conifers</td><td align="left" rowspan="1" colspan="1"><italic>C. castaneicolor</italic> A.H. Sm. 1944</td><td align="left" rowspan="1" colspan="1">15.10.1941</td><td align="left" rowspan="1" colspan="1">A.H. Smith</td><td align="left" rowspan="1" colspan="1">KF732275</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cephalixoides</italic> M.M. Moser & Thiers 1995 (holotype)</td><td align="left" rowspan="1" colspan="1">87/188</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">USA, Wyoming, Teton National Forest, Flagstaff Road</td><td align="left" rowspan="1" colspan="1">In subalpin forest under <italic>Picea engelmannii</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. cephalixoides</italic> M.M. Moser & Thiers 1995</td><td align="left" rowspan="1" colspan="1">09.08.1987</td><td align="left" rowspan="1" colspan="1">H.D. Thiers</td><td align="left" rowspan="1" colspan="1">KF732276</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cephalixolargus</italic> Rob. Henry 1977 (holotype)</td><td align="left" rowspan="1" colspan="1">6048</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Bois de Dampierre</td><td align="left" rowspan="1" colspan="1">In mixed frondose forest under<italic> Fagus</italic>, <italic>Quercus</italic>, <italic>Carpinus </italic>and <italic>Betula</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. largus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">R. Henry</td><td align="left" rowspan="1" colspan="1">KF732277</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. chromataphilus</italic> Rob. Henry 1989 (holotype)</td><td align="left" rowspan="1" colspan="1">86.90</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Luxeuil</td><td align="left" rowspan="1" colspan="1">In mixed forest</td><td align="left" rowspan="1" colspan="1"><italic>C. chromataphilus</italic> Rob. Henry 1989</td><td align="left" rowspan="1" colspan="1">1986</td><td align="left" rowspan="1" colspan="1">Exposition</td><td align="left" rowspan="1" colspan="1">KF732278</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cinctipes</italic> Bidaud, Eyssart. & Hermitte 2004 (holotype)</td><td align="left" rowspan="1" colspan="1">GE 02-100</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Var, Les Mayons</td><td align="left" rowspan="1" colspan="1">Under <italic>Quercus suber </italic>and <italic>Erica arborea</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. pseudocephalixus</italic> Bidaud & Moënne-Locc. 2000</td><td align="left" rowspan="1" colspan="1">01.11.2002</td><td align="left" rowspan="1" colspan="1">J.-C. Hermitte</td><td align="left" rowspan="1" colspan="1">KF732279</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. citrinifolius</italic> A.H. Sm. 1939 (holotype)</td><td align="left" rowspan="1" colspan="1">3158</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Washington, Olympic National Park, Boulder Creek</td><td align="left" rowspan="1" colspan="1">Under fir</td><td align="left" rowspan="1" colspan="1"><italic>C. citrinifolius</italic> A.H. Sm. 1939</td><td align="left" rowspan="1" colspan="1">15.10.1935</td><td align="left" rowspan="1" colspan="1">A.H. Smith</td><td align="left" rowspan="1" colspan="1">KF732280</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. citrinipedes</italic> A.H. Sm. 1942 (holotype)</td><td align="left" rowspan="1" colspan="1">15305</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Michigan, Ann Arbor</td><td align="left" rowspan="1" colspan="1">On humus in oak woods</td><td align="left" rowspan="1" colspan="1"><italic>C. citrinipedes</italic> A.H. Sm. 1942</td><td align="left" rowspan="1" colspan="1">11.09.1940</td><td align="left" rowspan="1" colspan="1">A.H. Smith</td><td align="left" rowspan="1" colspan="1">KF732281</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. citriolens</italic> Ammirati & M.M. Moser 1999 (holotype)</td><td align="left" rowspan="1" colspan="1">97/122</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">USA, Wyoming, Teton National Forest Flagstaff Creek</td><td align="left" rowspan="1" colspan="1">In subalpine forest under <italic>Picea engelmannii</italic>,<italic>Abies lasiocarpa</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. citriolens</italic> Ammirati & M.M. Moser 1999</td><td align="left" rowspan="1" colspan="1">06.08.1997</td><td align="left" rowspan="1" colspan="1">M. Moser & J. Ammirati</td><td align="left" rowspan="1" colspan="1">KF732282</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. claricolor</italic> (Fr.) Fr. 1838 (neotype)</td><td align="left" rowspan="1" colspan="1">CFP691</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Ång, Stigsjö, Uland, Langmyrberget</td><td align="left" rowspan="1" colspan="1">In spruce forest with blueberry</td><td align="left" rowspan="1" colspan="1"><italic>C. claricolor</italic> (Fr.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">09.08.1988</td><td align="left" rowspan="1" colspan="1">T.E. Brandrud et al.</td><td align="left" rowspan="1" colspan="1">KF732283</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. clarobaltoides</italic> var. <italic>longispermus</italic> Reumaux 1996 (holotype)</td><td align="left" rowspan="1" colspan="1">833</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Ile-de-France</td><td align="left" rowspan="1" colspan="1">Under conifers</td><td align="left" rowspan="1" colspan="1"><italic>C. clarobaltoides</italic> var. <italic>longispermus</italic> Reumaux 1996</td><td align="left" rowspan="1" colspan="1">15.09.1987</td><td align="left" rowspan="1" colspan="1">M. Pelerin</td><td align="left" rowspan="1" colspan="1">KF732284</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. clarus</italic> Reumaux 1996 (holotype)</td><td align="left" rowspan="1" colspan="1">4038</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Yonne, Forêt de Hervaux</td><td align="left" rowspan="1" colspan="1">In frondose forest</td><td align="left" rowspan="1" colspan="1"><italic>C. largus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">04.10.1994</td><td align="left" rowspan="1" colspan="1">Mlle Maité</td><td align="left" rowspan="1" colspan="1">KF732285</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. claviceps</italic> Reumaux 1996 (holotype)</td><td align="left" rowspan="1" colspan="1">2932</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Ardennes, Forêt de Belval</td><td align="left" rowspan="1" colspan="1">Under <italic>Quercus</italic> and <italic>Fagus</italic> on clayey-calcaerous soil</td><td align="left" rowspan="1" colspan="1"><italic>C. largus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">19.09.1992</td><td align="left" rowspan="1" colspan="1">P. Reumaux &F. Reumaux</td><td align="left" rowspan="1" colspan="1">KF732286</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cobaltinus</italic> Kytöv., Liimat. & Niskanen 2013 (holotype)</td><td align="left" rowspan="1" colspan="1">TN02-1012 (H6032404)</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, Ks, Kuusamo, Oulanka National Park</td><td align="left" rowspan="1" colspan="1">In herb-rich <italic>Picea abies</italic> forest with some <italic>Pinus</italic>, <italic>Betula</italic> and <italic>Populus</italic> on calcareous ground</td><td align="left" rowspan="1" colspan="1"><italic>C. cobaltinus</italic> Kytöv., Liimat. & Niskanen 2013</td><td align="left" rowspan="1" colspan="1">22.09.2002</td><td align="left" rowspan="1" colspan="1">T. Niskanen, I. Kytövuori & K.Liimatainen</td><td align="left" rowspan="1" colspan="1">KF673470</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. collocandoides</italic> Reumaux 2009 (holotype)</td><td align="left" rowspan="1" colspan="1">PML5087</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Yvelines, étang d’Or, Rambouillet</td><td align="left" rowspan="1" colspan="1">In mixed forest</td><td align="left" rowspan="1" colspan="1"><italic>C. collocandoides</italic> Reumaux 2009</td><td align="left" rowspan="1" colspan="1">27.10.1996</td><td align="left" rowspan="1" colspan="1">G. Redeuilh & P. Reumaux</td><td align="left" rowspan="1" colspan="1">KF732287</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. concrescens</italic> Secr. ex Bidaud, Moënne-Locc. & Reumaux 1996 (holotype)</td><td align="left" rowspan="1" colspan="1">3578</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Haute-Savoie, Le Danay, St-Jean-de-Sixt</td><td align="left" rowspan="1" colspan="1">Under <italic>Picea abies</italic> and <italic>Alnus viridis</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. balteatoalbus</italic> Rob. Henry 1985</td><td align="left" rowspan="1" colspan="1">10.10.1993</td><td align="left" rowspan="1" colspan="1">M. Mugnier</td><td align="left" rowspan="1" colspan="1">KF732288</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. congeminus</italic> Moënne-Locc. & Reumaux 1995 (holotype) 1995 (holotype)</td><td align="left" rowspan="1" colspan="1">3422</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Ardennes, Forêt de Belval</td><td align="left" rowspan="1" colspan="1">In deciduous forest on clayey-calcareous soil</td><td align="left" rowspan="1" colspan="1"><italic>C. largus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">03.10.1992</td><td align="left" rowspan="1" colspan="1">P. Reumaux &F. Reumaux</td><td align="left" rowspan="1" colspan="1">KF732289</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. corrugis</italic> A.H. Sm. 1944 (holotype)</td><td align="left" rowspan="1" colspan="1">16842</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Washington, Baker National Forest, Anderson Lookout, Ermine Creek Trail</td><td align="left" rowspan="1" colspan="1">Under conifers</td><td align="left" rowspan="1" colspan="1"><italic>C. turmalis</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">11.09.1941</td><td align="left" rowspan="1" colspan="1">A.H. Smith</td><td align="left" rowspan="1" colspan="1">KF732290</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. crassus</italic> Fr. 1838 (neotype)</td><td align="left" rowspan="1" colspan="1">CFP938</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Ång, Säbrä, Hårsta</td><td align="left" rowspan="1" colspan="1">In wet mixed forest with <italic>Sphagnum </italic>and <italic>Pinus</italic>, <italic>Picea</italic>, <italic>Betula</italic></td><td align="left" rowspan="1" colspan="1">C. <italic>crassus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">28.07.1990</td><td align="left" rowspan="1" colspan="1">T.E. Brandrud et al.</td><td align="left" rowspan="1" colspan="1">KF732291</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cremeiamarescens </italic>Kytöv., Liimat. & Niskanen sp. nov. (holotype)</td><td align="left" rowspan="1" colspan="1">IK11-014</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Sweden, Gtl, Alskog and När parish</td><td align="left" rowspan="1" colspan="1">Mesic to damp spruce forest with some <italic>Pinus</italic>, <italic>Quercus</italic> and <italic>Corylus</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. cremeiamarescens </italic>Kytöv., Liimat. & Niskanen sp. nov.</td><td align="left" rowspan="1" colspan="1">27.09.2011</td><td align="left" rowspan="1" colspan="1">I. Kytövuori</td><td align="left" rowspan="1" colspan="1">KF732493</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. crenulatus</italic> Rob. Henry ex Bidaud & Reumaux 2006 (holotype)</td><td align="left" rowspan="1" colspan="1">PML4866</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Allier, Forêt de Tronçais</td><td align="left" rowspan="1" colspan="1">In deciduous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. talus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">17.10.1992</td><td align="left" rowspan="1" colspan="1">F. Lopez</td><td align="left" rowspan="1" colspan="1">KF732292</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cruentipellis </italic>Kytöv., Liimat., Niskanen & Dima sp. nov. (holotype)</td><td align="left" rowspan="1" colspan="1">TN03-1451</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Sweden, Öl, Långlöt, Åstad, Nitares hägn</td><td align="left" rowspan="1" colspan="1">Grassy pasture with <italic>Corylus</italic> and <italic>Juniperus</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. cruentipellis </italic>Kytöv., Liimat., Niskanen & Dima sp. nov.</td><td align="left" rowspan="1" colspan="1">13.09.2003</td><td align="left" rowspan="1" colspan="1">I. Kytövuori, K.Liimatainen &T. Niskanen</td><td align="left" rowspan="1" colspan="1">KF732539</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cumatilis</italic> Fr. 1838 (neotype)</td><td align="left" rowspan="1" colspan="1">IK98-2164</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Sweden, Nrk, Hidinge, Garphyttan</td><td align="left" rowspan="1" colspan="1">Spruce forest</td><td align="left" rowspan="1" colspan="1"><italic>C. cumatilis</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">20.09.1998</td><td align="left" rowspan="1" colspan="1">I. Kytövuori</td><td align="left" rowspan="1" colspan="1">KF732293</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cupreorufus</italic> Brandrud 1994 (isotype)</td><td align="left" rowspan="1" colspan="1">CFP1038</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Upl, Älvkarleby, Billudden</td><td align="left" rowspan="1" colspan="1">In dry spruce forest on rich ground</td><td align="left" rowspan="1" colspan="1"><italic>C. cupreorufus</italic> Brandrud 1994</td><td align="left" rowspan="1" colspan="1">03.10.1990</td><td align="left" rowspan="1" colspan="1">T.E. Brandrud et al.</td><td align="left" rowspan="1" colspan="1">KF732294</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cupreoviolaceus</italic> Bidaud & Reumaux 1996 (holotype)</td><td align="left" rowspan="1" colspan="1">3426</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Ardennes, Forêt de Belval</td><td align="left" rowspan="1" colspan="1">In deciduous forest on clayey-calcareous soil</td><td align="left" rowspan="1" colspan="1"><italic>C. largus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">27.09.1992</td><td align="left" rowspan="1" colspan="1">P. Reumaux &F. Reumaux</td><td align="left" rowspan="1" colspan="1">KF732295</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cyanites</italic> Fr. 1838 (neotype)</td><td align="left" rowspan="1" colspan="1">AT2005069</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Sweden, Upl, Uppsala, Stadsskogen</td><td align="left" rowspan="1" colspan="1">In mixed forest</td><td align="left" rowspan="1" colspan="1"><italic>C. cyanites</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">26.08.2005</td><td align="left" rowspan="1" colspan="1">A. Taylor</td><td align="left" rowspan="1" colspan="1">KF732355</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. delaportei</italic> Rob. Henry 1988 (holotype)</td><td align="left" rowspan="1" colspan="1">RH8673</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Paris region</td><td align="left" rowspan="1" colspan="1">In frondose forest</td><td align="left" rowspan="1" colspan="1"><italic>C. delaportei</italic> Rob. Henry 1988</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">A. Delaporte</td><td align="left" rowspan="1" colspan="1">KF732297</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. dionysae</italic> var. <italic>avellaneus</italic> Rob. Henry ex Bidaud & Carteret 2008 (holotype)</td><td align="left" rowspan="1" colspan="1">AB97-10-361</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Ain, Champdor</td><td align="left" rowspan="1" colspan="1">In mixed calcareous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. dionysae</italic> var. <italic>avellaneus</italic> Rob. Henry ex Bidaud & Carteret 2008</td><td align="left" rowspan="1" colspan="1">21.10.1997</td><td align="left" rowspan="1" colspan="1">G. Chamonaz</td><td align="left" rowspan="1" colspan="1">KF732298</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. eliae</italic> Bidaud, Moënne-Locc. & Reumaux 1996 (holotype)</td><td align="left" rowspan="1" colspan="1">1032</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Haute-Savoie, Vieugy</td><td align="left" rowspan="1" colspan="1">In herbaceous <italic>Quercus</italic> forest</td><td align="left" rowspan="1" colspan="1"><italic>C. eliae</italic> Bidaud, Moënne-Locc. & Reumaux 1996</td><td align="left" rowspan="1" colspan="1">27.10.1988</td><td align="left" rowspan="1" colspan="1">P. Moënne-Loccoz</td><td align="left" rowspan="1" colspan="1">KF732299</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. elotoides</italic> M.M. Moser & McKnight 1995 (holotype)</td><td align="left" rowspan="1" colspan="1">87/60</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">USA, Wyoming, Teton National Forest, Turpin Meadow</td><td align="left" rowspan="1" colspan="1">Under <italic>Picea pungens</italic>, <italic>P. engelmannii</italic>, <italic>Pseudotsuga menziesii</italic> in subalpine forest</td><td align="left" rowspan="1" colspan="1"><italic>C. elotoides</italic> M.M. Moser & McKnight 1995</td><td align="left" rowspan="1" colspan="1">23.07.1987</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">KF732300</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. eumarginatus</italic> Rob. Henry ex Bidaud, Carteret & Reumaux 2009 (holotype)</td><td align="left" rowspan="1" colspan="1">AB07-10-175</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Ain, Chanay, col de Richemont</td><td align="left" rowspan="1" colspan="1">In coniferous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. purpurascens</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">16.10.2007</td><td align="left" rowspan="1" colspan="1">A. Bidaud</td><td align="left" rowspan="1" colspan="1">KF732301</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. evosmus</italic> Joachim ex Bidaud & Reumaux 2006 (holotype)*</td><td align="left" rowspan="1" colspan="1">PML4837</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Oise, Coye-la-Forêt</td><td align="left" rowspan="1" colspan="1">In deciduous forest on calcareous soil</td><td align="left" rowspan="1" colspan="1"><italic>C. evosmus</italic> Joachim ex Bidaud & Reumaux 2006</td><td align="left" rowspan="1" colspan="1">20.10.1995</td><td align="left" rowspan="1" colspan="1">M. Diamond</td><td align="left" rowspan="1" colspan="1">KF732302</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. flavaurora</italic> M.M. Moser & McKnight 1995 (holotype)*</td><td align="left" rowspan="1" colspan="1">89/187</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">USA, Wyoming, Teton National Forest, Fourmile meadow</td><td align="left" rowspan="1" colspan="1">Under <italic>Picea engelmannii</italic> in subalpine forest</td><td align="left" rowspan="1" colspan="1"><italic>C. elotoides</italic> M.M. Moser & McKnight 1995</td><td align="left" rowspan="1" colspan="1">07.08.1989</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">KF732303</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. flavescentipes</italic> Reumaux 1996 (holotype)</td><td align="left" rowspan="1" colspan="1">3606</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Loiret, Forêt de Montargis</td><td align="left" rowspan="1" colspan="1">Under <italic>Quercus</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. flavescentipes</italic> Reumaux 1996</td><td align="left" rowspan="1" colspan="1">27.10.1993</td><td align="left" rowspan="1" colspan="1">Participants of mycological field trip in Sully-sur-Loire</td><td align="left" rowspan="1" colspan="1">KF732304</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. flavipallens </italic>Kytöv., Liimat. & Niskanen sp. nov. (holotype)</td><td align="left" rowspan="1" colspan="1">IK08-1729 (H6032745)</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, Kn, Kajaani, Hietalahti</td><td align="left" rowspan="1" colspan="1">In fairly damp grass-herb-spruce forest with <italic>Pinus</italic>, <italic>Betula</italic> and <italic>Populus</italic><italic>tremula</italic>,<italic></italic>on calcareous ground</td><td align="left" rowspan="1" colspan="1"><italic>C. flavipallens </italic>Kytöv., Liimat. & Niskanen sp. nov.</td><td align="left" rowspan="1" colspan="1">13.09.2008</td><td align="left" rowspan="1" colspan="1">I. Kytövuori</td><td align="left" rowspan="1" colspan="1">KF732554</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. flavivelatus </italic>Kytöv., Liimat. & Niskanen sp. nov. (holotype)</td><td align="left" rowspan="1" colspan="1">IK98-885</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Sweden, Nb, Pajala, Junosuando</td><td align="left" rowspan="1" colspan="1">In dryish <italic>Picea abies</italic> heath forest with <italic>Pinus</italic>, <italic>Betula</italic>, and open meadows</td><td align="left" rowspan="1" colspan="1"><italic>C. flavivelatus </italic>Kytöv., Liimat. & Niskanen sp. nov. </td><td align="left" rowspan="1" colspan="1">15.08.1998</td><td align="left" rowspan="1" colspan="1">I. Kytövuori</td><td align="left" rowspan="1" colspan="1">KF732528</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. flavobulbus</italic> Ammirati & M.M. Moser 1997 (holotype)</td><td align="left" rowspan="1" colspan="1">95/629</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">USA, California, Del Norte Co., Highway 199, Danger Point</td><td align="left" rowspan="1" colspan="1">Under <italic>Quercus vacciniifolia</italic> and <italic>Q. garrayana</italic> in rather dry habitats on basic soils</td><td align="left" rowspan="1" colspan="1"><italic>C. flavobulbus</italic> Ammirati & M.M. Moser 1997</td><td align="left" rowspan="1" colspan="1">29.11.1995</td><td align="left" rowspan="1" colspan="1">M. Moser</td><td align="left" rowspan="1" colspan="1">KF732305</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. foetens</italic> (M.M. Moser) M.M. Moser 1967 (holotype)</td><td align="left" rowspan="1" colspan="1">51/128</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">Austria, Tyrol, Hötting-Innsbruck, Buchtal</td><td align="left" rowspan="1" colspan="1">In mixed deciduous forest under <italic>Fagus</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. foetens</italic> (M.M. Moser) M.M. Moser 1967</td><td align="left" rowspan="1" colspan="1">13.09.1951</td><td align="left" rowspan="1" colspan="1">M. Moser</td><td align="left" rowspan="1" colspan="1">KF732306</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. fraudulosus </italic>Britzelm. 1885 (neotype)</td><td align="left" rowspan="1" colspan="1">IK95-1852</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Germany, Baden-Württemberg, Freudenstadt, Heiligenbronn</td><td align="left" rowspan="1" colspan="1">In gently sloping grass-herb coniferous forest (<italic>Abies alba</italic>, <italic>Picea abies</italic>, <italic>Fagus sylvatica</italic>) on calcareous ground</td><td align="left" rowspan="1" colspan="1"><italic>C. fraudulosus </italic>Britzelm. 1885</td><td align="left" rowspan="1" colspan="1">05.10.1995</td><td align="left" rowspan="1" colspan="1">I. Kytövuori</td><td align="left" rowspan="1" colspan="1">KF732518</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. fraudulosus</italic> var. <italic>patrickensis</italic> M.M. Moser 2000 (holotype)</td><td align="left" rowspan="1" colspan="1">95/617</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">USA, California, Humboldt Co., Trinidad, Patrick’s Point State Park</td><td align="left" rowspan="1" colspan="1">Under <italic>Picea sitkensis</italic> and <italic>Pseudotsuga</italic><italic>menziesii</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. patrickensis </italic>(M.M. Moser) Niskanen, Liimat., Kytöv., Bojantchev & Ammirati comb. nov.</td><td align="left" rowspan="1" colspan="1">25.11.1995</td><td align="left" rowspan="1" colspan="1">M. Moser</td><td align="left" rowspan="1" colspan="1">KF732307</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. frondosophilus </italic>Bidaud 2001 (holotype)</td><td align="left" rowspan="1" colspan="1">PML4817</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Ain, Cerin</td><td align="left" rowspan="1" colspan="1">In deciduous forest on calcareous soil</td><td align="left" rowspan="1" colspan="1"><italic>C. platypus </italic>(M.M. Moser) M.M. Moser 1967</td><td align="left" rowspan="1" colspan="1">16.11.1997</td><td align="left" rowspan="1" colspan="1">A. Bidaud</td><td align="left" rowspan="1" colspan="1">KF732562</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. fuligineofolius</italic> (M.M. Moser) M.M. Moser & Peintner 2002 (holotype)</td><td align="left" rowspan="1" colspan="1">91/682</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">USA, California, Mendocino, Caspar Little Lake Road</td><td align="left" rowspan="1" colspan="1">In coniferous forest with <italic>Pseudotsuga, i>Tsuga</italic> and <italic>Sequoia</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. fuligineofolius</italic> (M.M. Moser) M.M. Moser & Peintner 2002</td><td align="left" rowspan="1" colspan="1">06.12.1991</td><td align="left" rowspan="1" colspan="1">M. Moser</td><td align="left" rowspan="1" colspan="1">KF732308</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. fulmineus</italic> var. <italic>sulphureus</italic> Kauffman 1918 (lectotype)*</td><td align="left" rowspan="1" colspan="1">MICH10351</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Michigan, Washtenaw, Ann Arbor</td><td align="left" rowspan="1" colspan="1">On the ground among humus, in frondose or mixed wood</td><td align="left" rowspan="1" colspan="1"><italic>C. fulmineus </italic>var. <italic>sulphureus </italic>Kauffman 1918</td><td align="left" rowspan="1" colspan="1">03.10.1910</td><td align="left" rowspan="1" colspan="1">C.H. Kauffman</td><td align="left" rowspan="1" colspan="1">KF732309</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. gentianeus</italic> Bidaud 1993 (holotype)</td><td align="left" rowspan="1" colspan="1">2575</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Ain, Lampnas</td><td align="left" rowspan="1" colspan="1">In calcareous <italic>Fagus</italic> forest with <italic>Betula</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. gentianeus</italic> Bidaud 1993</td><td align="left" rowspan="1" colspan="1">08.10.1991</td><td align="left" rowspan="1" colspan="1">M. Russi</td><td align="left" rowspan="1" colspan="1">KF732310</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. genuinus</italic> Rob. Henry ex Bidaud & Carteret 2009 (holotype)</td><td align="left" rowspan="1" colspan="1">XC2005-132</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Seine-Maritime, Les Petites Dalles, Valmont</td><td align="left" rowspan="1" colspan="1">In deciduous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. collocandoides</italic> Reumaux 2009</td><td align="left" rowspan="1" colspan="1">02.09.2005</td><td align="left" rowspan="1" colspan="1">X. Carteret</td><td align="left" rowspan="1" colspan="1">KF732311</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. georgiolens</italic> Rob. Henry 1986 (holotype)*</td><td align="left" rowspan="1" colspan="1">RH3153</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Languedoc-Roussillon</td><td align="left" rowspan="1" colspan="1">In forest of <italic>Quercus ilex</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. georgiolens</italic> Rob. Henry 1986</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">KF732312</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. glaucocephalus</italic> M.M. Moser, Ammirati & Halling 2000 (holotype)</td><td align="left" rowspan="1" colspan="1">95/679</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">USA, California, Mendocino Co., Caspar Little Lake Rd.</td><td align="left" rowspan="1" colspan="1">In mixed forests with <italic>Tsuga</italic>, <italic>Pseudotsuga</italic>, <italic>Pinus ponderosa</italic>, <italic>Abies</italic> and <italic>Arctostaphylos</italic><italic>manzanita</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. glaucocephalus</italic> M.M. Moser, Ammirati & Halling 2000</td><td align="left" rowspan="1" colspan="1">07.12.1995</td><td align="left" rowspan="1" colspan="1">J. Ammirati & M. Moser</td><td align="left" rowspan="1" colspan="1">KF732313</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. glaucopoides</italic> Kauffman 1923 (holotype)</td><td align="left" rowspan="1" colspan="1">MICH10358</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Colorado, Grand, Leal</td><td align="left" rowspan="1" colspan="1">Under spruce and fir</td><td align="left" rowspan="1" colspan="1"><italic>C. glaucopus</italic> (Schaeff.: Fr.) Gray 1821</td><td align="left" rowspan="1" colspan="1">16.08.1917</td><td align="left" rowspan="1" colspan="1">C.H. Kauffman</td><td align="left" rowspan="1" colspan="1">KF732314</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. glaucopus</italic> (Schaeff.: Fr.) Gray 1821 (neotype)</td><td align="left" rowspan="1" colspan="1">CFP786</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Mpd, Alnö, Ås brygga</td><td align="left" rowspan="1" colspan="1">In dry spruce forest on calcareous ground</td><td align="left" rowspan="1" colspan="1"><italic>C. glaucopus</italic> (Schaeff.: Fr.) Gray 1821</td><td align="left" rowspan="1" colspan="1">21.09.1988</td><td align="left" rowspan="1" colspan="1">T.E. Brandrud et al.</td><td align="left" rowspan="1" colspan="1">KF732315</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. glaucopus</italic> var. <italic>olivaceus</italic> f. <italic>ingratus</italic> Moënne-Locc. 2008 (holotype)</td><td align="left" rowspan="1" colspan="1">PML762</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Haute-Savoie, forêt de la Semine, Clarafond</td><td align="left" rowspan="1" colspan="1">In calcareous deciduous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. scaurocaninus </italic>Chevassut & Rob. Henry 1982</td><td align="left" rowspan="1" colspan="1">05.11.1987</td><td align="left" rowspan="1" colspan="1">P. Moënne-Loccoz</td><td align="left" rowspan="1" colspan="1">KF732316</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. glaucopus</italic> var. <italic>subrubrovelatus</italic> Bidaud 2008 (holotype)</td><td align="left" rowspan="1" colspan="1">AB97-11-431</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Ain, Farges, in mixed forest with <italic>Populus tremula</italic>, <italic>Corylus sativa</italic>, <italic>Fagus</italic><italic>sylvatica</italic>, <italic>Abies alba</italic></td><td align="left" rowspan="1" colspan="1">In deciduous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. subrubrovelatus </italic>(Bidaud) Kytöv., Liimat., Niskanen & Dima comb. nov.</td><td align="left" rowspan="1" colspan="1">01.11.1997</td><td align="left" rowspan="1" colspan="1">A. Bidaud</td><td align="left" rowspan="1" colspan="1">KF732317</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. gracilior </italic>(Jul. Schäff. ex M.M. Moser) M.M. Moser 1967 (holotype)</td><td align="left" rowspan="1" colspan="1">58/73</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">Germany, Baden-Württemberg, Randengebiet near Zollhaus</td><td align="left" rowspan="1" colspan="1">In <italic>Fagus</italic> forest on calcareous soil</td><td align="left" rowspan="1" colspan="1"><italic>C. gracilior </italic>(Jul. Schäff. ex M.M. Moser) M.M. Moser 1967</td><td align="left" rowspan="1" colspan="1">09.10.1958</td><td align="left" rowspan="1" colspan="1">M. Moser</td><td align="left" rowspan="1" colspan="1">UDB001082</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. gratus </italic>Reumaux 2008 (holotype)</td><td align="left" rowspan="1" colspan="1">PML85</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Haute-Savoie, forêt de la Mandallaz, Choisy</td><td align="left" rowspan="1" colspan="1">In deciduous forest under <italic>Fagus</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. leonicolor</italic> Reumaux 2001</td><td align="left" rowspan="1" colspan="1">27.09.1986</td><td align="left" rowspan="1" colspan="1">P. Moënne-Loccoz</td><td align="left" rowspan="1" colspan="1">KF732318</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. griseocoeruleus </italic>Ammirati & M.M. Moser 1997 (holotype)</td><td align="left" rowspan="1" colspan="1">95/685</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">USA, California, Mendocino Co., about 8 km east of Mendocino on road 408</td><td align="left" rowspan="1" colspan="1">Under <italic>Lithocarpus densiflora </italic>Moser 1997</td><td align="left" rowspan="1" colspan="1"><italic>C. griseocoeruleus </italic>Ammirati & M.M.</td><td align="left" rowspan="1" colspan="1">08.12.1995</td><td align="left" rowspan="1" colspan="1">M. Moser</td><td align="left" rowspan="1" colspan="1">KF732319</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. herculeolens</italic> Bidaud 1996 (holotype)*</td><td align="left" rowspan="1" colspan="1">3700</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Montbrison, Loire</td><td align="left" rowspan="1" colspan="1">Under <italic>Quercus petraea</italic> on basaltic soil</td><td align="left" rowspan="1" colspan="1"><italic>C. chromataphilus</italic> Rob. Henry 1989</td><td align="left" rowspan="1" colspan="1">11.11.1992</td><td align="left" rowspan="1" colspan="1">A. Bidaud</td><td align="left" rowspan="1" colspan="1">KF732320</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. herpeticus</italic> Fr. 1838 (neotype)</td><td align="left" rowspan="1" colspan="1">CFP936</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Ång, Säbrå, Hällenyland</td><td align="left" rowspan="1" colspan="1">Under <italic>Picea</italic> on cultivated area</td><td align="left" rowspan="1" colspan="1"><italic>C. herpeticus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">20.07.1990</td><td align="left" rowspan="1" colspan="1">T.E. Brandrud et al.</td><td align="left" rowspan="1" colspan="1">KF732321</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. hysginicolor</italic> Bidaud 1995 (holotype)*</td><td align="left" rowspan="1" colspan="1">2955</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Ain, Innimont</td><td align="left" rowspan="1" colspan="1">In young herbaceus, calcareous <italic>Picea</italic> forest</td><td align="left" rowspan="1" colspan="1"><italic>C. badiolatus</italic> (M.M. Moser) M.M. Moser 1967</td><td align="left" rowspan="1" colspan="1">03.10.1992</td><td align="left" rowspan="1" colspan="1">A. Bidaud</td><td align="left" rowspan="1" colspan="1">KF732322</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. immixtus</italic> Kauffman 1932 (holotype)*</td><td align="left" rowspan="1" colspan="1">MICH10365</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Washington, Mason, Lake Cushman, Olympic Mountains</td><td align="left" rowspan="1" colspan="1">Under fir and hemlock</td><td align="left" rowspan="1" colspan="1"><italic>C. immixtus</italic> Kauffman 1932</td><td align="left" rowspan="1" colspan="1">20.10.1915</td><td align="left" rowspan="1" colspan="1">C.H. Kauffman</td><td align="left" rowspan="1" colspan="1">KF732323</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. inamoenus</italic> (J. Favre) Quadr. 1985 (holotype)*</td><td align="left" rowspan="1" colspan="1">13522</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">Switzerland</td><td align="left" rowspan="1" colspan="1">In subalpine coniferous forest on calcareous ground</td><td align="left" rowspan="1" colspan="1"><italic>C. rosargutus</italic> Chevassut & Rob. Henry 1978</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">J. Favre</td><td align="left" rowspan="1" colspan="1">KF732324</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. infractus</italic> (Pers.: Fr.) Fr. 1838 (neotype)</td><td align="left" rowspan="1" colspan="1">CFP495</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Boh, Tossene, Anneröd</td><td align="left" rowspan="1" colspan="1">In beech forest, on medium rich ground</td><td align="left" rowspan="1" colspan="1"><italic>C. infractus</italic> (Pers.: Fr.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">15.09.1986</td><td align="left" rowspan="1" colspan="1">T.E. Brandrud et al.</td><td align="left" rowspan="1" colspan="1">KF732325</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. infractus</italic> var. <italic>aeruginosus</italic> Rob. Henry ex Reumaux 2009</td><td align="left" rowspan="1" colspan="1">XC2006-66</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Seine-et-Marne, forêt de Villefermoy</td><td align="left" rowspan="1" colspan="1">In deciduous forest on clayey-calcareous soil</td><td align="left" rowspan="1" colspan="1"><italic>C. infractus</italic> (Pers.: Fr.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">11.10.2006</td><td align="left" rowspan="1" colspan="1">R. Chalange</td><td align="left" rowspan="1" colspan="1">KF732326</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. infractus</italic> var. <italic>flavus</italic> M.M. Moser 1999 (holotype)*</td><td align="left" rowspan="1" colspan="1">97/169</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">USA, Wyoming, Shoshone National Forest, Brooks Lodge</td><td align="left" rowspan="1" colspan="1">In coniferous forest under <italic>Picea</italic><italic>engelmannii</italic> and <italic>Abies lasiocarpa</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. infractiflavus </italic>(M.M. Moser) Kytöv., Niskanen, Liimat., Bojantchev & Ammirati stat. nov. & nom. nov.</td><td align="left" rowspan="1" colspan="1">12.08.1997</td><td align="left" rowspan="1" colspan="1">M. Moser</td><td align="left" rowspan="1" colspan="1">KF732327</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. josephii</italic> Reumaux 2006 (holotype)</td><td align="left" rowspan="1" colspan="1">PML5193</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Ardennes, forêt d’Elan</td><td align="left" rowspan="1" colspan="1">In calcareous deciduous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. gracilior</italic> (Jul. Schäff. ex M.M. Moser) M.M. Moser 1967</td><td align="left" rowspan="1" colspan="1">08.10.1999</td><td align="left" rowspan="1" colspan="1">P. Reumaux</td><td align="left" rowspan="1" colspan="1">KF732328</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. juxtadibaphus</italic> Rob. Henry 1983 (holotype)</td><td align="left" rowspan="1" colspan="1">RH3880</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Haut-Doubs, L’Hôpital du Grosbois</td><td align="left" rowspan="1" colspan="1">In submontane, calcareous <italic>Carpinus</italic> forest</td><td align="left" rowspan="1" colspan="1"><italic>C. juxtadibaphus</italic> Rob. Henry 1983</td><td align="left" rowspan="1" colspan="1">1950</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">KF732329</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. kuehneri</italic> M.M. Moser 1974 (holotype)</td><td align="left" rowspan="1" colspan="1">1965/0042</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">Austria, Tyrol, Oetztal, Untergurgl</td><td align="left" rowspan="1" colspan="1">Under <italic>Alnus viridis</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. kuehneri</italic> M.M. Moser 1974</td><td align="left" rowspan="1" colspan="1">03.09.1965</td><td align="left" rowspan="1" colspan="1">M. Moser</td><td align="left" rowspan="1" colspan="1">KF732330</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. kytoevuorii </italic>Niskanen & Liimat. sp. nov. (holotype)</td><td align="left" rowspan="1" colspan="1">TN05-158, H6029355</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, Ks, Kuusamo, Oulanka, Ampumavaara</td><td align="left" rowspan="1" colspan="1">In old, grass-herb <italic>Picea abies</italic> forest with some <italic>Betula</italic>, <italic>Pinus</italic><italic>sylvestris</italic> and <italic>Populus tremula</italic>,</td><td align="left" rowspan="1" colspan="1"><italic>C. kytoevuorii </italic>Niskanen & Liimat. sp. nov.</td><td align="left" rowspan="1" colspan="1">17.09.2005</td><td align="left" rowspan="1" colspan="1">K. Liimatainen & T. Niskanen</td><td align="left" rowspan="1" colspan="1">KF732529</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. laetargutus</italic> Chevassut & Rob. Henry 1978 (holotype)</td><td align="left" rowspan="1" colspan="1">RH70405</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Mont Aigoual</td><td align="left" rowspan="1" colspan="1">In mixed forest of <italic>Picea</italic> and <italic>Fagus</italic></td><td align="left" rowspan="1" colspan="1">C. <italic>crassus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">29.09.1975</td><td align="left" rowspan="1" colspan="1">Mme Moutet</td><td align="left" rowspan="1" colspan="1">KF732331</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. largoides</italic> Rob. Henry ex Bidaud, Carteret & Reumaux 2009 (holotype)</td><td align="left" rowspan="1" colspan="1">PML2336</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Ain, bois de la Morgne, Ordonnaz</td><td align="left" rowspan="1" colspan="1">In deciduous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. subpurpurascens</italic> (Batsch) Fr. 1838</td><td align="left" rowspan="1" colspan="1">06.10.1991</td><td align="left" rowspan="1" colspan="1">A. Bidaud</td><td align="left" rowspan="1" colspan="1">KF732332</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. largus</italic> Fr. 1838 (neotype)</td><td align="left" rowspan="1" colspan="1">TN08-060 (H6001957)</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, V, Turku, Ruissalo</td><td align="left" rowspan="1" colspan="1">In deciduous forest of <italic>Quercus robur</italic>, <italic>Corylus</italic><italic>avellana</italic> and some <italic>Betula</italic> on mull soil</td><td align="left" rowspan="1" colspan="1"><italic>C. largus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">03.09.2008</td><td align="left" rowspan="1" colspan="1">K. Liimatainen & T. Niskanen</td><td align="left" rowspan="1" colspan="1">AB859985</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. largusiellus</italic> Reumaux 1996 (holotype)</td><td align="left" rowspan="1" colspan="1">3411</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Ardennes, Forêt de Boult</td><td align="left" rowspan="1" colspan="1">Under deciduous trees (<italic>Betula</italic>, <italic>Carpinus</italic>) on clayey-calcareous soil</td><td align="left" rowspan="1" colspan="1"><italic>C. largus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">09.10.1992</td><td align="left" rowspan="1" colspan="1">P. Reumaux</td><td align="left" rowspan="1" colspan="1">KF732334</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. latoclaricolor </italic>Rob. Henry 1989 (holotype)</td><td align="left" rowspan="1" colspan="1">RH1199</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France</td><td align="left" rowspan="1" colspan="1">In <italic>Picea</italic> forest</td><td align="left" rowspan="1" colspan="1"><italic>C. badiolatus</italic> (M.M. Moser) M.M. Moser 1967</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">KF732335</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. lemanicus</italic> A. Favre & Vialard 2000 (holotype)</td><td align="left" rowspan="1" colspan="1">452177</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Savoy, Thonon-les-Bains, Margencel</td><td align="left" rowspan="1" colspan="1">Under <italic>Pinus</italic> and <italic>Picea</italic> on calcareous soil</td><td align="left" rowspan="1" colspan="1"><italic>C. delaportei</italic> Rob. Henry 1988</td><td align="left" rowspan="1" colspan="1">06.11.1999</td><td align="left" rowspan="1" colspan="1">A. Favre, J. Vialard</td><td align="left" rowspan="1" colspan="1">KF732336</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. leonicolor</italic> Reumaux 2001 (holotype)</td><td align="left" rowspan="1" colspan="1">4739</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Ardennes, bois du Vivier</td><td align="left" rowspan="1" colspan="1">In clayey-calcareous deciduous forest under <italic>Quercus</italic> and <italic>Carpinus</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. leonicolor</italic> Reumaux 2001</td><td align="left" rowspan="1" colspan="1">04.10.1992</td><td align="left" rowspan="1" colspan="1">P. Reumaux</td><td align="left" rowspan="1" colspan="1">KF732337</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. lilacinicolor</italic> Reumaux 1996 (holotype)</td><td align="left" rowspan="1" colspan="1">3512</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Ardennes, La Croix-aux-Bois</td><td align="left" rowspan="1" colspan="1">In deciduous forest on clayey-calcareous soil</td><td align="left" rowspan="1" colspan="1"><italic>C. largus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">16.09.1979</td><td align="left" rowspan="1" colspan="1">P. Reumaux</td><td align="left" rowspan="1" colspan="1">KF732338</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. lintrisporus</italic> Reumaux 1997 (holotype)</td><td align="left" rowspan="1" colspan="1">2935</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Ardennes, Mont Dieu</td><td align="left" rowspan="1" colspan="1">In clayey-calcareous deciduous forest (<italic>Carpinus</italic>, <italic>Betula</italic>)</td><td align="left" rowspan="1" colspan="1"><italic>C. largus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">23.09.1992</td><td align="left" rowspan="1" colspan="1">P. Reumaux</td><td align="left" rowspan="1" colspan="1">KF732339</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. lividoviolaceus</italic> Rob. Henry 1987 (paratype)*</td><td align="left" rowspan="1" colspan="1">RH2932</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France</td><td align="left" rowspan="1" colspan="1">In frondose woodland</td><td align="left" rowspan="1" colspan="1"><italic>C. largus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">1969</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">KF732340</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. luteiaureus </italic> Kytöv., Liimat. & Niskanen sp. nov. (holotype)</td><td align="left" rowspan="1" colspan="1">IK07-247b (H6033617)</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, OP, Kiiminki, Juuvansydänmaa</td><td align="left" rowspan="1" colspan="1">In grass-herb <italic>Picea abies</italic> forest with some <italic>Betula</italic>, <italic>Populus</italic><italic>tremula</italic> and <italic>Pinus</italic>, on calcareous ground</td><td align="left" rowspan="1" colspan="1"><italic>C. luteiaureus </italic>Kytöv., Liimat. & Niskanen sp. nov.</td><td align="left" rowspan="1" colspan="1">17.08.2007</td><td align="left" rowspan="1" colspan="1">I. Kytövuori</td><td align="left" rowspan="1" colspan="1">KF732568</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. luteoarmillatus</italic> A.H. Sm. 1942 (holotype)*</td><td align="left" rowspan="1" colspan="1">15360</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Michigan, Sharron Hollow</td><td align="left" rowspan="1" colspan="1">On muck soil in low woods</td><td align="left" rowspan="1" colspan="1"><italic>C. luteoarmillatus</italic> A.H. Sm. 1942</td><td align="left" rowspan="1" colspan="1">14.09.1940</td><td align="left" rowspan="1" colspan="1">A.H. Smith</td><td align="left" rowspan="1" colspan="1">KF732341</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. luteobrunnescens</italic> A.H. Sm. 1944 (holotype)</td><td align="left" rowspan="1" colspan="1">17785</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Washington, Olympic National Park, Olympic Hot Springs</td><td align="left" rowspan="1" colspan="1">Under conifers</td><td align="left" rowspan="1" colspan="1"><italic>C. luteobrunnescens</italic> A.H. Sm. 1944</td><td align="left" rowspan="1" colspan="1">11.10.1941</td><td align="left" rowspan="1" colspan="1">D.E. Stunz & A.H. Smith</td><td align="left" rowspan="1" colspan="1">KF732342</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. luteocingulatus </italic>Bidaud & Fillion 1992 (holotype)*</td><td align="left" rowspan="1" colspan="1">AB91-10-260</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Haute-Savoie, Forêt de la Semine</td><td align="left" rowspan="1" colspan="1">Under <italic>Quercus</italic> and <italic>Carpinus</italic> on clayey, slightly acid soil</td><td align="left" rowspan="1" colspan="1"><italic>C. luteocingulatus </italic>Bidaud & Fillion 1992</td><td align="left" rowspan="1" colspan="1">10.1991</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">KF732343</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. luteovaginans</italic> Bidaud & Faurite-Gendron 2006 (holotype)</td><td align="left" rowspan="1" colspan="1">AB03-11-73</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Ardèche, Saint-Alban-sur- Sampzon</td><td align="left" rowspan="1" colspan="1">In deciduous forest under <italic>Quercus ilex</italic> and <italic>Q. humilis</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. aurantiopallidus</italic> Bidaud 2006</td><td align="left" rowspan="1" colspan="1">05.11.2003</td><td align="left" rowspan="1" colspan="1">A. Faurite-Gendron</td><td align="left" rowspan="1" colspan="1">KF732344</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. maculatipes</italic> Bidaud 1996 (holotype)</td><td align="left" rowspan="1" colspan="1">2845</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Savoie, Les Arcs</td><td align="left" rowspan="1" colspan="1">In subalpine zone under <italic>Picea abies</italic> and <italic>Alnus viridis</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. pseudonaevosus</italic> Rob. Henry 1957</td><td align="left" rowspan="1" colspan="1">13.08.1992</td><td align="left" rowspan="1" colspan="1">P.A. Moreau</td><td align="left" rowspan="1" colspan="1">KF732345</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. maculatocaespitosus</italic> Bidaud 2009 (holotype)</td><td align="left" rowspan="1" colspan="1">AB08-10-302</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Ain, Cerin</td><td align="left" rowspan="1" colspan="1">In calcareous deciduous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. maculatocaespitosus</italic> Bidaud 2009</td><td align="left" rowspan="1" colspan="1">11.10.2008</td><td align="left" rowspan="1" colspan="1">A. Bidaud</td><td align="left" rowspan="1" colspan="1">KF732346</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. mahiquesii </italic>Vila, A. Ortega & Suár.-Sant. 2008 (holotype)</td><td align="left" rowspan="1" colspan="1">GDA54298</td><td align="left" rowspan="1" colspan="1">GDA</td><td align="left" rowspan="1" colspan="1">Spain, Catalonia, Perafita</td><td align="left" rowspan="1" colspan="1">Under <italic>Cistus monspeliensis</italic>, on acid soil</td><td align="left" rowspan="1" colspan="1"><italic>C. mahiquesii</italic> Vila, A. Ortega & Suár.-Sant. 2008</td><td align="left" rowspan="1" colspan="1">18.01.2008</td><td align="left" rowspan="1" colspan="1">J.Vila, X.Llimona</td><td align="left" rowspan="1" colspan="1">FM202139</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. melleicarneus </italic>Kytöv., Liimat., Niskanen & Brandrud sp. nov. (holotype)</td><td align="left" rowspan="1" colspan="1">IK01-053</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Estonia, Hiiumaa, Sarve, Soonlepa</td><td align="left" rowspan="1" colspan="1">In deciduous forest <italic>(Corylus</italic>, <italic>Quercus</italic>) with some <italic>Pinus</italic> on calcareous ground</td><td align="left" rowspan="1" colspan="1"><italic>C.melleicarneus</italic>Kytöv., Liimat., Niskanen & Brandrud sp. nov.</td><td align="left" rowspan="1" colspan="1">16.09.2001</td><td align="left" rowspan="1" colspan="1">I. Kytövuori</td><td align="left" rowspan="1" colspan="1">KF732577</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. mendax</italic> Bidaud, Mahiques & Reumaux 2011 (holotype)</td><td align="left" rowspan="1" colspan="1">AB07-10-162</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Ain, col de Richemont, Chanay</td><td align="left" rowspan="1" colspan="1">In mixed forest under <italic>Betula</italic> and <italic>Picea</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. mendax</italic> Bidaud, Mahiques & Reumaux 2011</td><td align="left" rowspan="1" colspan="1">12.10.2007</td><td align="left" rowspan="1" colspan="1">E. Bidaud & R. Fillion</td><td align="left" rowspan="1" colspan="1">KF732401</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. metarius</italic> Kauffman 1921 (holotype)</td><td align="left" rowspan="1" colspan="1">MICH10374</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Colorado, Grand, Leal</td><td align="left" rowspan="1" colspan="1">Under spruce and fir</td><td align="left" rowspan="1" colspan="1"><italic>C. metarius</italic> Kauffman 1921</td><td align="left" rowspan="1" colspan="1">29.08.1917</td><td align="left" rowspan="1" colspan="1">C.H. Kauffman</td><td align="left" rowspan="1" colspan="1">KF732347</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. misermontii</italic> Chevassut & Rob. Henry 1986 (holotype)</td><td align="left" rowspan="1" colspan="1">RH84.134</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Montpellier</td><td align="left" rowspan="1" colspan="1">Under <italic>Quercus ilex</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. misermontii</italic> Chevassut & Rob. Henry 1986</td><td align="left" rowspan="1" colspan="1">12.1984</td><td align="left" rowspan="1" colspan="1">G. Chevassut</td><td align="left" rowspan="1" colspan="1">KF732348</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. montanus</italic> Kauffman 1932 (holotype)</td><td align="left" rowspan="1" colspan="1">MICH10377</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Oregon, Clackamas, Mt Hood, near Welches Post Office</td><td align="left" rowspan="1" colspan="1">In hemlock and cedar forest</td><td align="left" rowspan="1" colspan="1"><italic>C. montanus</italic> Kauffman 1932</td><td align="left" rowspan="1" colspan="1">03.10.1922</td><td align="left" rowspan="1" colspan="1">C.H. Kauffman</td><td align="left" rowspan="1" colspan="1">KF732349</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. multiformis</italic> Fr. 1838 (neotype)</td><td align="left" rowspan="1" colspan="1">CFP445</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Ång, Häggdånger, Sjö</td><td align="left" rowspan="1" colspan="1">In spruce forest with blueberry</td><td align="left" rowspan="1" colspan="1"><italic>C. multiformis</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">21.08.1986</td><td align="left" rowspan="1" colspan="1">T.E. Brandrud et al.</td><td align="left" rowspan="1" colspan="1">KF732350</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C.multiformis</italic> var. <italic>caesiophyllus</italic> Moënne-Locc. 2006 (holotype)</td><td align="left" rowspan="1" colspan="1">PML882</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Savoie, Arith</td><td align="left" rowspan="1" colspan="1">Under <italic>Picea</italic> on calcareous soil</td><td align="left" rowspan="1" colspan="1"><italic>C.caesiolamellatus</italic>(Bidaud) Kytöv., Liimat., Niskanen, Brandrud, Frøslev & A.F.S. Taylor comb. nov.</td><td align="left" rowspan="1" colspan="1">03.06.1988</td><td align="left" rowspan="1" colspan="1">P. Moënne-Loccoz</td><td align="left" rowspan="1" colspan="1">KF732351</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. muricinicolor</italic> Moënne-Locc. 1996 (holotype)</td><td align="left" rowspan="1" colspan="1">3582</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Haute-Savoie, Quintal</td><td align="left" rowspan="1" colspan="1">Under <italic>Picea</italic> on calcareous soil</td><td align="left" rowspan="1" colspan="1"><italic>C. variecolor</italic> (Pers.: Fr.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">11.10.1993</td><td align="left" rowspan="1" colspan="1">P. Moënne-Loccoz</td><td align="left" rowspan="1" colspan="1">KF732352</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. mutabilis</italic> A.H. Sm. 1944 (holotype)</td><td align="left" rowspan="1" colspan="1">17451</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Washington, Olympic National Park, Olympic Hot Springs</td><td align="left" rowspan="1" colspan="1">Under conifers, pine, Douglas fir and mountain hemlock</td><td align="left" rowspan="1" colspan="1"><italic>C. occidentalis</italic> A.H. Sm. 1939</td><td align="left" rowspan="1" colspan="1">30.09.1941</td><td align="left" rowspan="1" colspan="1">A.H. Smith</td><td align="left" rowspan="1" colspan="1">KF732353</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. myrtilliphilus</italic> Kytöv., Liimat., Niskanen & Brandrud sp. nov. (holotype)</td><td align="left" rowspan="1" colspan="1">IK97-1469 (H6032751)</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, Kn, Suomussalmi, Suolijärvi</td><td align="left" rowspan="1" colspan="1">In gently sloping, partly swampy, grass- herb spruce forest with some <italic>Pinus</italic>,<italic>Betula</italic>,<italic>Populus</italic>, <italic>Alnus incana</italic> and <italic>Salix</italic> spp.</td><td align="left" rowspan="1" colspan="1"><italic>C. myrtilliphilus</italic> Kytöv., Liimat., Niskanen & Brandrud sp. nov.</td><td align="left" rowspan="1" colspan="1">14.09.1997</td><td align="left" rowspan="1" colspan="1">I. Kytövuori</td><td align="left" rowspan="1" colspan="1">KF732605</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. neotriumphans</italic> Bidaud, Moënne-Locc. & Reumaux 2000 (holotype)</td><td align="left" rowspan="1" colspan="1">PML209</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Haute-Savoie, Semnoz</td><td align="left" rowspan="1" colspan="1">Under <italic>Picea</italic> on calcareous soil</td><td align="left" rowspan="1" colspan="1"><italic>C. neotriumphans</italic> Bidaud, Moënne- Locc. & Reumaux 2000</td><td align="left" rowspan="1" colspan="1">18.09.1985</td><td align="left" rowspan="1" colspan="1">P. Moënne-Loccoz</td><td align="left" rowspan="1" colspan="1">KF732354</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. norrlandicus</italic> Brandrud 1989 (isotype)</td><td align="left" rowspan="1" colspan="1">CFP526</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Ång, Häggdånger, Torrom</td><td align="left" rowspan="1" colspan="1">High herbaceous spruce forest on rich ground</td><td align="left" rowspan="1" colspan="1"><italic>C. norrlandicus</italic> Brandrud</td><td align="left" rowspan="1" colspan="1">22.09.1986</td><td align="left" rowspan="1" colspan="1">T.E. Brandrud</td><td align="left" rowspan="1" colspan="1">DQ117928</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. obsoletus</italic> Kühner 1955 (syntype)</td><td align="left" rowspan="1" colspan="1">Sa-52-66 (G00262069)</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Haute-Savoie, Samoëns</td><td align="left" rowspan="1" colspan="1">Under <italic>Fagus</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. obsoletus</italic> Kühner 1955</td><td align="left" rowspan="1" colspan="1">22.09.1952</td><td align="left" rowspan="1" colspan="1">R. Kühner</td><td align="left" rowspan="1" colspan="1">KF732628</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. obsoletus</italic> Kühner 1955 (syntype)</td><td align="left" rowspan="1" colspan="1">Sa-52-91 (G00262070)</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Haute-Savoie, Samoëns</td><td align="left" rowspan="1" colspan="1">Under <italic>Fagus</italic> and <italic>Quercus</italic> among moss and grass</td><td align="left" rowspan="1" colspan="1"><italic>C. obsoletus</italic> Kühner 1955</td><td align="left" rowspan="1" colspan="1">27.09.1952</td><td align="left" rowspan="1" colspan="1">R. Kühner</td><td align="left" rowspan="1" colspan="1">KF732356</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. occidentalis</italic> A.H. Sm. 1939 (holotype)</td><td align="left" rowspan="1" colspan="1">8654</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, California, Trinidad</td><td align="left" rowspan="1" colspan="1">Under redwood</td><td align="left" rowspan="1" colspan="1"><italic>C. occidentalis</italic> A.H. Sm. 1939</td><td align="left" rowspan="1" colspan="1">12.11.1937</td><td align="left" rowspan="1" colspan="1">A.H. Smith</td><td align="left" rowspan="1" colspan="1">KF732357</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. occultus</italic> Moënne-Locc. & Reumaux 1996 (holotype)</td><td align="left" rowspan="1" colspan="1">3591</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Ile-de-France, région de Versailles</td><td align="left" rowspan="1" colspan="1">In calcareous deciduous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. largus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">14.10.1993</td><td align="left" rowspan="1" colspan="1">M. Cerutti</td><td align="left" rowspan="1" colspan="1">KF732358</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. ochraceobrunneus</italic> Rob. Henry ex Bidaud, Moënne-Locc. & Reumaux 2000 (holotype)</td><td align="left" rowspan="1" colspan="1">4027</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Haute-Savoie, Bois de Vorcier</td><td align="left" rowspan="1" colspan="1">In mixed forest dominated by <italic>Picea</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. ochraceobrunneus</italic> Rob. Henry ex Bidaud, Moënne-Locc. & Reumaux 2000</td><td align="left" rowspan="1" colspan="1">05.11.1995</td><td align="left" rowspan="1" colspan="1">H. Debauvais</td><td align="left" rowspan="1" colspan="1">KF732359</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. ochribubalinus </italic>Kytöv., Liimat. & Niskanen sp. nov. (holotype)</td><td align="left" rowspan="1" colspan="1">IK93-641, H6032734</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, U, Espoo, Nuuksio</td><td align="left" rowspan="1" colspan="1">In fairly rich grass-herb forest with <italic>Populus</italic><italic>tremula</italic>, <italic>Betula</italic>, <italic>Alnus incana</italic>, <italic>Quercus</italic>, <italic>Corylus</italic>, <italic>Prunus padus</italic>, <italic>Salix</italic> spp., and some old pines and young spruces</td><td align="left" rowspan="1" colspan="1"><italic>C. ochribubalinus </italic>Kytöv., Liimat. & Niskanen sp. nov.</td><td align="left" rowspan="1" colspan="1">02.09.1993</td><td align="left" rowspan="1" colspan="1">I. Kytövuori</td><td align="left" rowspan="1" colspan="1">KF732530</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. ochroclarus</italic> Rob. Henry ex Rob. Henry 1996 (holotype)*</td><td align="left" rowspan="1" colspan="1">2792</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Haute-Savoie, Forêt de la Semine</td><td align="left" rowspan="1" colspan="1">In deciduous forest mixed with some <italic>Picea</italic> on clayey-calcareous soil</td><td align="left" rowspan="1" colspan="1"><italic>C. ochroclarus</italic> Rob. Henry ex Rob. Henry 1996</td><td align="left" rowspan="1" colspan="1">11.07.1992</td><td align="left" rowspan="1" colspan="1">R. Fillion</td><td align="left" rowspan="1" colspan="1">KF732360</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. ochropudorinus</italic> Rob. Henry ex Bidaud & Reumaux 2006 (holotype)</td><td align="left" rowspan="1" colspan="1">PML2339</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Ain, Meyriat</td><td align="left" rowspan="1" colspan="1">In deciduous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. talus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">09.10.1991</td><td align="left" rowspan="1" colspan="1">A. Bidaud</td><td align="left" rowspan="1" colspan="1">KF732361</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. olidoamarus</italic> var. <italic>valentinus</italic> Mahiques & A. Favre 1999 (holotype)</td><td align="left" rowspan="1" colspan="1">452173</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">Spain, Valencia</td><td align="left" rowspan="1" colspan="1">Under <italic>Quercus suber</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. olidoamarus</italic> var. <italic>valentinus</italic> Mahiques & A. Favre 1999</td><td align="left" rowspan="1" colspan="1">26.10.1996</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">KF732362</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. oliveopetasatus</italic> M.M. Moser 2000 (holotype)</td><td align="left" rowspan="1" colspan="1">95/360</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">USA, Oregon, Wasco Co., Mt Hood, Clear Creek Camp Ground</td><td align="left" rowspan="1" colspan="1">In mixed coniferous forest (<italic>Abies</italic>,<italic>Picea</italic>, <italic>Pinus</italic>,<italic>Tsuga</italic>, <italic>Pseudotsuga</italic>,<italic> Larix</italic>)</td><td align="left" rowspan="1" colspan="1"><italic>C. oliveopetasatus</italic> M.M. Moser 2000</td><td align="left" rowspan="1" colspan="1">25.10.1995</td><td align="left" rowspan="1" colspan="1">M. Moser</td><td align="left" rowspan="1" colspan="1">KF732363</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. olympianus</italic> A.H. Sm. 1939 (holotype)</td><td align="left" rowspan="1" colspan="1">3242</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Washington, Olympic Mountains, Olympic Hot Springs, Boulder Creek</td><td align="left" rowspan="1" colspan="1">Under fir</td><td align="left" rowspan="1" colspan="1"><italic>C. olympianus</italic> A.H. Sm. 1939</td><td align="left" rowspan="1" colspan="1">19.10.1935</td><td align="left" rowspan="1" colspan="1">A.H. Smith</td><td align="left" rowspan="1" colspan="1">KF732364</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. ophiopus</italic> Peck 1878 (part of type)*</td><td align="left" rowspan="1" colspan="1">s.n.</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">USA, Maryland</td><td align="left" rowspan="1" colspan="1">Among fallen leaves in woods</td><td align="left" rowspan="1" colspan="1"><italic>C. ophiopus</italic> Peck 1878</td><td align="left" rowspan="1" colspan="1">September</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">KF732365</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. oregonensis</italic> A.H. Sm. 1939 (holotype)</td><td align="left" rowspan="1" colspan="1">3557</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Oregon, Florence, Lake Tahkenitch</td><td align="left" rowspan="1" colspan="1">Under spruce</td><td align="left" rowspan="1" colspan="1"><italic>C. oregonensis</italic> A.H. Sm. 1939</td><td align="left" rowspan="1" colspan="1">19.11.1935</td><td align="left" rowspan="1" colspan="1">A.H. Smith</td><td align="left" rowspan="1" colspan="1">KF732366</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C.orichalceus</italic> var. <italic>olympianus</italic> A.H. Sm. 1944 (holotype)</td><td align="left" rowspan="1" colspan="1">17513</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Washington, Olympic National Park, Olympic Hot Springs</td><td align="left" rowspan="1" colspan="1">Under conifers</td><td align="left" rowspan="1" colspan="1"><italic>C. pseudocupreorufus </italic>(A.H. Sm.) Nis- kanen, Liimat. & Ammirati stat. nov. & nom. nov.</td><td align="left" rowspan="1" colspan="1">02.10.1941</td><td align="left" rowspan="1" colspan="1">A.H. Smith</td><td align="left" rowspan="1" colspan="1">KF732367</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. orichalceus</italic> var. <italic>olympianus </italic>f. <italic>luteifolius</italic> A.H. Sm. 1944 (holotype)</td><td align="left" rowspan="1" colspan="1">16970</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Washington, Olympic Mts, Lake Angeles</td><td align="left" rowspan="1" colspan="1">Under conifers</td><td align="left" rowspan="1" colspan="1"><italic>C. luteicolor </italic>(A.H. Sm.) Ammirati, Bojant- chev, Niskanen & Liimat. stat. nov. & nom. nov</td><td align="left" rowspan="1" colspan="1">19.09.1941</td><td align="left" rowspan="1" colspan="1">A.H. Smith</td><td align="left" rowspan="1" colspan="1">KF732368</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. orichalceus</italic> var. <italic>xanthocephalus</italic> A.H. Sm. 1944 (holotype)*</td><td align="left" rowspan="1" colspan="1">17514</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Washington, Olympic National Park, Olympic Hot Springs</td><td align="left" rowspan="1" colspan="1">Under conifers</td><td align="left" rowspan="1" colspan="1"><italic>C. orichalceus</italic> var. <italic>xanthocephalus</italic> A.H. Sm. 1944</td><td align="left" rowspan="1" colspan="1">02.10.1941</td><td align="left" rowspan="1" colspan="1">A.H. Smith</td><td align="left" rowspan="1" colspan="1">KF732369</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pallidifolius</italic> A.H. Sm. 1939 (holotype)</td><td align="left" rowspan="1" colspan="1">3244</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Washington, Olympic Mountains, Olympic Hot Springs</td><td align="left" rowspan="1" colspan="1">Under fir</td><td align="left" rowspan="1" colspan="1"><italic>C. pallidifolius</italic> A.H. Sm. 1939</td><td align="left" rowspan="1" colspan="1">19.10.1935</td><td align="left" rowspan="1" colspan="1">A.H. Smith</td><td align="left" rowspan="1" colspan="1">KF732370</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pallidirimosus </italic>Kytöv., Liimat. & Niskanen sp. nov. (holotype)</td><td align="left" rowspan="1" colspan="1">IK95-585, H6035694</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, InL, Utsjoki, Kevo, Tsieskuljohka</td><td align="left" rowspan="1" colspan="1">In mesic heath forest with <italic>Betula</italic> and <italic>Pinus</italic>, some moist depressions</td><td align="left" rowspan="1" colspan="1"><italic>C. pallidirimosus </italic>Kytöv., Liimat. & Niskanen sp. nov.</td><td align="left" rowspan="1" colspan="1">17.08.1995</td><td align="left" rowspan="1" colspan="1">I. Kytövuori</td><td align="left" rowspan="1" colspan="1">KF732578</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pansa</italic> (Fr.) Sacc. 1887 (neotype)</td><td align="left" rowspan="1" colspan="1">IK90-1826</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, V, Kemiö, Pederså</td><td align="left" rowspan="1" colspan="1">At small, abandoned limestone quarries, spruce heath forest, roadside</td><td align="left" rowspan="1" colspan="1"><italic>C. pansa</italic> (Fr.) Sacc. 1887</td><td align="left" rowspan="1" colspan="1">21.09.1990</td><td align="left" rowspan="1" colspan="1">I. Kytövuori</td><td align="left" rowspan="1" colspan="1">KF732522</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. papulosus</italic> Fr. 1838 (neotype)*</td><td align="left" rowspan="1" colspan="1">CFP344</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Mpd, Alnö, Ås</td><td align="left" rowspan="1" colspan="1">In spruce forest on calcareous ground</td><td align="left" rowspan="1" colspan="1"><italic>C. papulosus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">07.09.1985</td><td align="left" rowspan="1" colspan="1">T.E. Brandrud et al.</td><td align="left" rowspan="1" colspan="1">KF732371</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. paracrassus</italic> Reumaux 1995*</td><td align="left" rowspan="1" colspan="1">3522</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Ile-de-France, Fontainebleau</td><td align="left" rowspan="1" colspan="1">In calcareous deciduous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. largus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">07.10.1979</td><td align="left" rowspan="1" colspan="1">N. Martelli</td><td align="left" rowspan="1" colspan="1">KF732372</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. paracyanopus</italic> Moënne-Locc. & Reumaux 1996 (holotype)</td><td align="left" rowspan="1" colspan="1">3510</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Ardennes, Bois du Mont Dieu</td><td align="left" rowspan="1" colspan="1">Under <italic>Carpinus</italic> and <italic>Betula</italic>, on clayey- calcareous soil</td><td align="left" rowspan="1" colspan="1"><italic>C. largus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">25.09.1980</td><td align="left" rowspan="1" colspan="1">P. Reumaux</td><td align="left" rowspan="1" colspan="1">KF732373</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. parafulmineus </italic>Rob. Henry 1993</td><td align="left" rowspan="1" colspan="1">2384</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Drôme, Romeyer</td><td align="left" rowspan="1" colspan="1">In montane forest of <italic>Fagus</italic> and <italic>Abies</italic>, on calcareous ground</td><td align="left" rowspan="1" colspan="1"><italic>C. parafulmineus</italic> Rob. Henry 1993</td><td align="left" rowspan="1" colspan="1">02.11.1991</td><td align="left" rowspan="1" colspan="1">A. Bidaud & P. Reumaux</td><td align="left" rowspan="1" colspan="1">KF732552</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. parargutus</italic> Bidaud, Moënne-Locc. & Reumaux 1999 (holotype)</td><td align="left" rowspan="1" colspan="1">1144</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Ile-de-France</td><td align="left" rowspan="1" colspan="1">In deciduous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. pardinus</italic> Reumaux 1995</td><td align="left" rowspan="1" colspan="1">24.08.1987</td><td align="left" rowspan="1" colspan="1">J. Poirier</td><td align="left" rowspan="1" colspan="1">KF732374</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pardinus</italic> Reumaux 1995 (holotype)</td><td align="left" rowspan="1" colspan="1">3432</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Ardennes, Forêt de Belval</td><td align="left" rowspan="1" colspan="1">In deciduous forest on clayey-calcareous soil</td><td align="left" rowspan="1" colspan="1"><italic>C. pardinus</italic> Reumaux 1995</td><td align="left" rowspan="1" colspan="1">27.09.1992</td><td align="left" rowspan="1" colspan="1">P. Reumaux &F. Reumaux</td><td align="left" rowspan="1" colspan="1">KF732375</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. parolivascens</italic> Moënne-Locc. & Reumaux 2009 (holotype)</td><td align="left" rowspan="1" colspan="1">PML29</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Haute-Savoie, plateau des Glières</td><td align="left" rowspan="1" colspan="1">Under <italic>Pinus</italic> among <italic>Sphagnum</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. scaurus</italic> (Fr.: Fr.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">08.09.1984</td><td align="left" rowspan="1" colspan="1">R. Baubet</td><td align="left" rowspan="1" colspan="1">KF732377</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. patibilis</italic> Brandrud & Melot 1983 (holotype)*</td><td align="left" rowspan="1" colspan="1">213-78</td><td align="left" rowspan="1" colspan="1">O</td><td align="left" rowspan="1" colspan="1">Norway, Akh, Nannestad, Hornsjøen</td><td align="left" rowspan="1" colspan="1">In oligotrophic <italic>Picea abies</italic> forest</td><td align="left" rowspan="1" colspan="1"><italic>C. patibilis</italic> Brandrud & Melot 1983</td><td align="left" rowspan="1" colspan="1">10.08.1978</td><td align="left" rowspan="1" colspan="1">T.E. Brandrud</td><td align="left" rowspan="1" colspan="1">KF732378</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. patibilis</italic> var. <italic>scoticus</italic> Brandrud 1997 (holotype)*</td><td align="left" rowspan="1" colspan="1">TEB161-83</td><td align="left" rowspan="1" colspan="1">E</td><td align="left" rowspan="1" colspan="1">Scotland, Perthshire, Calvine, Struan Wood</td><td align="left" rowspan="1" colspan="1">In subalpine zone under <italic>Betula pubescens</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. largus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">22.09.1983</td><td align="left" rowspan="1" colspan="1">T.E. Brandrud</td><td align="left" rowspan="1" colspan="1">KF732379</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. percomis</italic> Fr. 1838 (neotype)</td><td align="left" rowspan="1" colspan="1">TN08-041</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, V, Karjaa, Kohagen</td><td align="left" rowspan="1" colspan="1">In herb-rich <italic>Picea abies</italic> forest with some <italic>Corylus avellana</italic>, <italic>Quercus robur</italic>, <italic>Betula</italic> and <italic>Populus tremula</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. percomis</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">02.09.2008</td><td align="left" rowspan="1" colspan="1">K.Liimatainen & T.Niskanen</td><td align="left" rowspan="1" colspan="1">KF732380</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. perpallens</italic> Chevassut & Rob. Henry 1978 (holotype)</td><td align="left" rowspan="1" colspan="1">RH3928</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Mont Aigoual</td><td align="left" rowspan="1" colspan="1">Under <italic>Picea</italic> and <italic>Fagus</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. perpallens</italic> Chevassut & Rob. Henry 1978</td><td align="left" rowspan="1" colspan="1">28.10.1973</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">KF732381</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. persoonianus</italic> Bidaud 2009 (holotype)*</td><td align="left" rowspan="1" colspan="1">AB97-11-496</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Ain, Cerin</td><td align="left" rowspan="1" colspan="1">In grassy meadow</td><td align="left" rowspan="1" colspan="1"><italic>C. persoonianus</italic> Bidaud 2009</td><td align="left" rowspan="1" colspan="1">16.11.1997</td><td align="left" rowspan="1" colspan="1">A. Bidaud</td><td align="left" rowspan="1" colspan="1">KF732382</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. phylladus</italic> Rob. Henry 1983 (holotype)*</td><td align="left" rowspan="1" colspan="1">RH2046</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Jura, Forêt de Chaux</td><td align="left" rowspan="1" colspan="1">In frondose forest with <italic>Quercus </italic>and <italic>Fagus</italic></td><td align="left" rowspan="1" colspan="1">C. <italic>crassus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">07.1965</td><td align="left" rowspan="1" colspan="1">R. Henry</td><td align="left" rowspan="1" colspan="1">KF732383</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pini</italic> Brandrud 1996 (isotype)</td><td align="left" rowspan="1" colspan="1">CFP394</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Norway, Oppl, Østre Toten, Balke</td><td align="left" rowspan="1" colspan="1">In coniferous forest on calcareous ground</td><td align="left" rowspan="1" colspan="1"><italic>C. pini</italic> Brandrud 1996</td><td align="left" rowspan="1" colspan="1">25.09.1985</td><td align="left" rowspan="1" colspan="1">T.E. Brandrud et al.</td><td align="left" rowspan="1" colspan="1">KF732384</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. piriodolens</italic> Moënne-Locc. 1996 (holotype)</td><td align="left" rowspan="1" colspan="1">642</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Haute-Savoie, Plateau de Solaison</td><td align="left" rowspan="1" colspan="1">In young calcareous <italic>Picea</italic> forest</td><td align="left" rowspan="1" colspan="1"><italic>C. variecolor</italic> (Pers.: Fr.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">01.09.1987</td><td align="left" rowspan="1" colspan="1">R. Baubet</td><td align="left" rowspan="1" colspan="1">KF732385</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. platypus </italic>(M.M. Moser) M.M. Moser 1967 (holotype)</td><td align="left" rowspan="1" colspan="1">58/64 (19580064) </td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">Germany, Baden-Württemberg, Inzigkofen</td><td align="left" rowspan="1" colspan="1">In <italic>Fagus</italic> forest on calcareous soil</td><td align="left" rowspan="1" colspan="1"><italic>C. platypus </italic>(M.M. Moser) M.M. Moser 1967</td><td align="left" rowspan="1" colspan="1">09.10.1948</td><td align="left" rowspan="1" colspan="1">M.M. Moser</td><td align="left" rowspan="1" colspan="1">KF732563</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. ponderosus</italic> A.H. Sm. 1939 (holotype)</td><td align="left" rowspan="1" colspan="1">9273</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Oregon, Cave City</td><td align="left" rowspan="1" colspan="1">Under <italic>Pinus ponderosa</italic> and <italic>Quercus</italic> spp.</td><td align="left" rowspan="1" colspan="1"><italic>C. ponderosus</italic> A.H. Sm. 1939</td><td align="left" rowspan="1" colspan="1">01.12.1937</td><td align="left" rowspan="1" colspan="1">A.H. Smith</td><td align="left" rowspan="1" colspan="1">KF732386</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. porphyropus</italic> (Alb. & Schwein.) Fr. 1838 (neotype)</td><td align="left" rowspan="1" colspan="1">CFP717</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Jmt, Ragunda, Ragunda</td><td align="left" rowspan="1" colspan="1">In birch forest on rich ground</td><td align="left" rowspan="1" colspan="1"><italic>C. porphyropus</italic> (Alb. & Schwein.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">20.08.1988</td><td align="left" rowspan="1" colspan="1">T.E. Brandrud et al.</td><td align="left" rowspan="1" colspan="1">KF732387</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. porphyropus</italic> var. <italic>porphyrophorus</italic> Reumaux 2009 (holotype)</td><td align="left" rowspan="1" colspan="1">PML5086</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Ardennes, La Croix-aux-Bois</td><td align="left" rowspan="1" colspan="1">In deciduous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. porphyropus</italic> (Alb. & Schwein.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">06.10.1995</td><td align="left" rowspan="1" colspan="1">P. Reumaux</td><td align="left" rowspan="1" colspan="1">KF732388</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. praestans</italic> (Cordier) Gillet 1874 (epitype)</td><td align="left" rowspan="1" colspan="1">IK94-1861</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">France, Ain, communen d’Echallan</td><td align="left" rowspan="1" colspan="1">In <italic>Fagus</italic> forest with <italic>Picea</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. praestans</italic> (Cordier) Gillet 1874</td><td align="left" rowspan="1" colspan="1">27.10.1994</td><td align="left" rowspan="1" colspan="1">I. Kytövuori</td><td align="left" rowspan="1" colspan="1">KF732389</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. psalliotoides</italic> Chevassut & Rob. Henry 1978 (holotype)*</td><td align="left" rowspan="1" colspan="1">RH3154</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, St. Mitre-les-Remparts</td><td align="left" rowspan="1" colspan="1">Under <italic>Quercus ilex</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. psalliotoides</italic> Chevassut & Rob. Henry 1978</td><td align="left" rowspan="1" colspan="1">22.11.1970</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">KF732390</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pseudocephalixus</italic> Bidaud & Moënne-Locc. 2000 (holotype)*</td><td align="left" rowspan="1" colspan="1">4446</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Drôme, Forêt de Romeyer</td><td align="left" rowspan="1" colspan="1">In mixed calcareous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. pseudocephalixus</italic> Bidaud & Moënne-Locc. 2000</td><td align="left" rowspan="1" colspan="1">02.11.1991</td><td align="left" rowspan="1" colspan="1">A. Bidaud</td><td align="left" rowspan="1" colspan="1">KF732392</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pseudocyanites</italic> var. <italic>paucus</italic> Reumaux 2005 (holotype)</td><td align="left" rowspan="1" colspan="1">PML5261</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Val-de-Marne, bois d’Ivry</td><td align="left" rowspan="1" colspan="1">In deciduous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. cyanites</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">17.10.1998</td><td align="left" rowspan="1" colspan="1">A. Bardet & R. Bardet</td><td align="left" rowspan="1" colspan="1">KF732393</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pseudogracilior</italic> Reumaux 2006 (holotype)</td><td align="left" rowspan="1" colspan="1">PML4858</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Dordogne, Tursac</td><td align="left" rowspan="1" colspan="1">In calcareous deciduous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. pseudogracilior</italic> Reumaux 2006</td><td align="left" rowspan="1" colspan="1">26.10.1997</td><td align="left" rowspan="1" colspan="1">P. Reumaux</td><td align="left" rowspan="1" colspan="1">KF732394</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pseudolargus</italic> Rob. Henry 1987 (holotype)</td><td align="left" rowspan="1" colspan="1">RH70218</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Doubs</td><td align="left" rowspan="1" colspan="1">In frondose and coniferous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. pseudonaevosus</italic> Rob. Henry 1957</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">KF732395</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pseudominor</italic> Rob. Henry ex Reumaux 2006 (holotype)</td><td align="left" rowspan="1" colspan="1">PML4750</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Ardennes, Bois des Alleux</td><td align="left" rowspan="1" colspan="1">In deciduous forest on clayey-calcareous soil</td><td align="left" rowspan="1" colspan="1"><italic>C. talus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">27.09.1997</td><td align="left" rowspan="1" colspan="1">P. Reumaux</td><td align="left" rowspan="1" colspan="1">KF732396</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pseudonaevosus</italic> Rob. Henry 1957 (type)</td><td align="left" rowspan="1" colspan="1">RH162</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France</td><td align="left" rowspan="1" colspan="1">In montane <italic>Picea</italic> forest</td><td align="left" rowspan="1" colspan="1"><italic>C. pseudonaevosus</italic> Rob. Henry 1957</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">KF732397</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pseudonebularis</italic> Moënne-Locc. 1996 (holotype)</td><td align="left" rowspan="1" colspan="1">42</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Haute-Savoie, Massif du Semnoz</td><td align="left" rowspan="1" colspan="1">In subalpine <italic>Picea</italic> forest</td><td align="left" rowspan="1" colspan="1"><italic>C. pseudonebularis</italic> Moënne-Locc. 1996</td><td align="left" rowspan="1" colspan="1">22.09.1985</td><td align="left" rowspan="1" colspan="1">P. Moënne-Loccoz</td><td align="left" rowspan="1" colspan="1">KF732398</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pseudopansa</italic> Bidaud 2000 (holotype)</td><td align="left" rowspan="1" colspan="1">4401</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Ain, Le Poizat</td><td align="left" rowspan="1" colspan="1">In coniferous forest on calcareous ground</td><td align="left" rowspan="1" colspan="1"><italic>C. varius</italic> (Schaeff.: Fr.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">14.10.1995</td><td align="left" rowspan="1" colspan="1">P.A. Moreau</td><td align="left" rowspan="1" colspan="1">KF732399</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pseudopimus </italic>Rob. Henry ex Rob. Henry 2000 (holotype)</td><td align="left" rowspan="1" colspan="1">4516</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Ain, Innimont</td><td align="left" rowspan="1" colspan="1">In mixed forest</td><td align="left" rowspan="1" colspan="1"><italic>C. varius</italic> (Schaeff.: Fr.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">08.10.1990</td><td align="left" rowspan="1" colspan="1">D. Mazuir</td><td align="left" rowspan="1" colspan="1">KF732400</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pseudotalus</italic> Rob. Henry ex Bidaud & Reumaux 2006 (holotype)</td><td align="left" rowspan="1" colspan="1">PML4859</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Seine-Maritime, Forêt de la Londe-Rouvray</td><td align="left" rowspan="1" colspan="1">Under deciduous trees on calcareous soil</td><td align="left" rowspan="1" colspan="1"><italic>C. talus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">24.10.1997</td><td align="left" rowspan="1" colspan="1">J.-C. Malaval</td><td align="left" rowspan="1" colspan="1">KF732402</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pseudoturmalis</italic> Bidaud & Moënne-Locc. 2010 (holotype)</td><td align="left" rowspan="1" colspan="1">PML3465</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Haute-Savoie, Thorens-Glières</td><td align="left" rowspan="1" colspan="1">In coniferous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. claricolor</italic> (Fr.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">29.07.1993</td><td align="left" rowspan="1" colspan="1">A. Faurite-Gendron</td><td align="left" rowspan="1" colspan="1">KF732403</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pseudovariegatus</italic> M.M Moser 1999 (holotype)</td><td align="left" rowspan="1" colspan="1">97/296</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">USA, Wyoming, Shoshone National Forest, Lake east of Two Ocean Mt</td><td align="left" rowspan="1" colspan="1">In subalpine forest under <italic>Picea</italic><italic>engelmannii</italic>,<italic>Pinus albicaulius</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. pseudovariegatus</italic> M.M Moser 1999</td><td align="left" rowspan="1" colspan="1">31.08.1997</td><td align="left" rowspan="1" colspan="1">M. Moser</td><td align="left" rowspan="1" colspan="1">KF732404</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pseudovarius</italic> Moënne-Locc. & Reumaux 2000 (holotype)*</td><td align="left" rowspan="1" colspan="1">4391</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Ile de France, Bois de Noisiel</td><td align="left" rowspan="1" colspan="1">In calcareous deciduous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. luteocingulatus </italic>Bidaud & Fillion 1992</td><td align="left" rowspan="1" colspan="1">20.10.1996</td><td align="left" rowspan="1" colspan="1">G. Flantzer</td><td align="left" rowspan="1" colspan="1">KF732405</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. purpurascens</italic> Fr. 1838 (neotype)</td><td align="left" rowspan="1" colspan="1">IK98-2121</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Sweden, Nrk, Hidinge, Garphyttans NP</td><td align="left" rowspan="1" colspan="1">In fairly rich spruce grass-herb forest with <italic>Corylus</italic>, <italic>Populus</italic><italic>tremula</italic>, <italic>Betula</italic> and <italic>Quercus</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. purpurascens</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">20.09.1998</td><td align="left" rowspan="1" colspan="1">I. Kytövuori</td><td align="left" rowspan="1" colspan="1">KF732406</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. rapaceoides</italic> Bidaud, G. Riousset & Riousset 2000 (holotype)</td><td align="left" rowspan="1" colspan="1">AB97-12-527</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, St. Rémy de Provence, Bouches du Rhône</td><td align="left" rowspan="1" colspan="1">In deciduous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. rapaceoides</italic> Bidaud, G. Riousset & Riousset 2000</td><td align="left" rowspan="1" colspan="1">03.11.1997</td><td align="left" rowspan="1" colspan="1">G. Riousset & L. Riousset</td><td align="left" rowspan="1" colspan="1">KF732407</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. reverendissimus</italic> Bidaud, Moënne-Locc. & Reumaux 2000 (holotype)</td><td align="left" rowspan="1" colspan="1">4667</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Haute-Loire, Forêt de Miaune</td><td align="left" rowspan="1" colspan="1">In mixed calcareous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. reverendissimus</italic> Bidaud, Moënne- Locc. & Reumaux 2000</td><td align="left" rowspan="1" colspan="1">16.10.1997</td><td align="left" rowspan="1" colspan="1">P. Chapon</td><td align="left" rowspan="1" colspan="1">KF732408</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. rex-claricolorum</italic> Bidaud, Carteret & Reumaux 2010 (holotype)</td><td align="left" rowspan="1" colspan="1">AB04-09-163</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Ain, la Vèche, Chanay</td><td align="left" rowspan="1" colspan="1">In coniferous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. rex-claricolorum</italic> Bidaud, Carteret & Reumaux 2010</td><td align="left" rowspan="1" colspan="1">15.09.2004</td><td align="left" rowspan="1" colspan="1">A. Bidaud & R. Fillion</td><td align="left" rowspan="1" colspan="1">KF732409</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. rioussetiae</italic> Chevassut & Rob. Henry 1986 (holotype)</td><td align="left" rowspan="1" colspan="1">RH84.70-78</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Gravesen</td><td align="left" rowspan="1" colspan="1">Under <italic>Populus alba</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. rioussetiae</italic> Chevassut & Rob. Henry 1986</td><td align="left" rowspan="1" colspan="1">10.1984</td><td align="left" rowspan="1" colspan="1">Mme. Riousset</td><td align="left" rowspan="1" colspan="1">KF732410</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. rosargutus</italic> Chevassut & Rob. Henry 1978 (holotype)</td><td align="left" rowspan="1" colspan="1">RH70477</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Haut-Doubs</td><td align="left" rowspan="1" colspan="1">Under conifers</td><td align="left" rowspan="1" colspan="1"><italic>C. rosargutus</italic> Chevassut & Rob. Henry 1978</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">KF732411</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. rufior </italic>Reumaux 2000 (holotype)</td><td align="left" rowspan="1" colspan="1">1118</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Haute Ardenne</td><td align="left" rowspan="1" colspan="1">In coniferous forest, under <italic>Picea</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. varius</italic> (Schaeff.: Fr.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">08.11.1988</td><td align="left" rowspan="1" colspan="1">G. Flantzer</td><td align="left" rowspan="1" colspan="1">KF732412</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. rufoallutus</italic> Rob. Henry ex Bidaud & Reumaux 2006 (holotype)</td><td align="left" rowspan="1" colspan="1">PML635</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Haute-Savoie, Plateau de Glières</td><td align="left" rowspan="1" colspan="1">Under <italic>Picea</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. rufoallutus</italic> Rob. Henry ex Bidaud & Reumaux 2006</td><td align="left" rowspan="1" colspan="1">26.08.1987</td><td align="left" rowspan="1" colspan="1">P. Moënne-Loccoz & J. Melot</td><td align="left" rowspan="1" colspan="1">KF732413</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. rufoallutus</italic> var. <italic>caesiolamellatus</italic> Bidaud 2006 (holotype)</td><td align="left" rowspan="1" colspan="1">PML4905</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Ain, col des Bérentin</td><td align="left" rowspan="1" colspan="1">Under <italic>Picea</italic> on calcareous soil</td><td align="left" rowspan="1" colspan="1"><italic>C. caesiolamellatus </italic>(Bidaud) Kytöv., Liimat., Niskanen, Brandrud, Frøslev & A.F.S. Taylor comb. nov.</td><td align="left" rowspan="1" colspan="1">03.10.1993</td><td align="left" rowspan="1" colspan="1">A. Bidaud</td><td align="left" rowspan="1" colspan="1">KF732414</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. rufolatus </italic>Moënne-Locc. 1996 (holotype)</td><td align="left" rowspan="1" colspan="1">664</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Haute-Savoie, Plateau de Glières</td><td align="left" rowspan="1" colspan="1">Under <italic>Picea abies</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. rufolatus </italic>Moënne-Locc. 1996</td><td align="left" rowspan="1" colspan="1">26.09.1987</td><td align="left" rowspan="1" colspan="1">P. Moënne-Loccoz</td><td align="left" rowspan="1" colspan="1">KF732415</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. russus</italic> Fr. 1838 (neotype)</td><td align="left" rowspan="1" colspan="1">CFP941</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Ång, Säbrä, Överdal</td><td align="left" rowspan="1" colspan="1">In dry spruce forest on rich ground</td><td align="left" rowspan="1" colspan="1"><italic>C. russus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">03.08.1990</td><td align="left" rowspan="1" colspan="1">T.E. Brandrud et al.</td><td align="left" rowspan="1" colspan="1">KF732416</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. sabuletorum</italic> Redeuilh & Reumaux 1995 (holotype)*</td><td align="left" rowspan="1" colspan="1">2954</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Ile-de-France, Les Mureaux</td><td align="left" rowspan="1" colspan="1">On river sand-banks under deciduous trees</td><td align="left" rowspan="1" colspan="1"><italic>C. chromataphilus</italic> Rob. Henry 1989</td><td align="left" rowspan="1" colspan="1">01.10.1992</td><td align="left" rowspan="1" colspan="1">G. Redeuilh</td><td align="left" rowspan="1" colspan="1">KF732417</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. saginoides</italic> Bidaud & Reumaux 2000 (holotype)</td><td align="left" rowspan="1" colspan="1">1264</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Ain, Forêt de Meyriat</td><td align="left" rowspan="1" colspan="1">In mixed forest</td><td align="left" rowspan="1" colspan="1"><italic>C. varius</italic> (Schaeff.: Fr.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">22.10.1989</td><td align="left" rowspan="1" colspan="1">A. Bidaud & P. Reumaux</td><td align="left" rowspan="1" colspan="1">KF732418</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. sannio</italic> M.M. Moser 1999 (holotype)</td><td align="left" rowspan="1" colspan="1">97/352</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">USA, Wyoming, Teton National Forest, Lost Lake</td><td align="left" rowspan="1" colspan="1">In subalpine coniferous forest under <italic>Picea</italic><italic>engelmannii</italic>, <italic>Abies lasiocarpa</italic> and <italic>Pinus albicaulis</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. sannio</italic> M.M. Moser 1999</td><td align="left" rowspan="1" colspan="1">10.09.1997</td><td align="left" rowspan="1" colspan="1">M. Moser</td><td align="left" rowspan="1" colspan="1">KF732420</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. saxamontanus</italic> Fogel 1995 (holotype)</td><td align="left" rowspan="1" colspan="1">F2535</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Nevada, White Pine Co., Whealer Peak Campground</td><td align="left" rowspan="1" colspan="1">Under <italic>Pinus</italic> spp. and <italic>Abies</italic> spp.</td><td align="left" rowspan="1" colspan="1"><italic>C. saxamontanus</italic> Fogel 1995</td><td align="left" rowspan="1" colspan="1">30.06.1981</td><td align="left" rowspan="1" colspan="1">R. Fogel</td><td align="left" rowspan="1" colspan="1">KF732421</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. scaurocaninus </italic>Chevassut & Rob. Henry 1982 (holotype)*</td><td align="left" rowspan="1" colspan="1">RH71678</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Montpellier</td><td align="left" rowspan="1" colspan="1">Under <italic>Quercus ilex</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. scaurocaninus</italic> Chevassut & Rob. Henry 1982</td><td align="left" rowspan="1" colspan="1">04.11.1979</td><td align="left" rowspan="1" colspan="1">G. Chevassut</td><td align="left" rowspan="1" colspan="1">KF732422</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. scaurus</italic> (Fr.: Fr.) Fr. 1838 (neotype)</td><td align="left" rowspan="1" colspan="1">CFP1074</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Switzerland, Bern, Fribourg, Rechthalten</td><td align="left" rowspan="1" colspan="1">In the border of a peatbog with <italic>Pinus strobus</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. scaurus</italic> (Fr.: Fr.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">29.09.1991</td><td align="left" rowspan="1" colspan="1">T.E. Brandrud et al.</td><td align="left" rowspan="1" colspan="1">KF732423</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. scaurus</italic> f. <italic>phaeophyllus</italic> M.M. Moser 2001 (holotype)</td><td align="left" rowspan="1" colspan="1">94/243</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">Sweden, Sm, Femsjö, Stora Mosse, Källanäset</td><td align="left" rowspan="1" colspan="1">On bare peat soil in a mire with <italic>Pinus sylvestris</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. scaurus</italic> (Fr.: Fr.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">13.09.1994</td><td align="left" rowspan="1" colspan="1">M. Moser</td><td align="left" rowspan="1" colspan="1">KF732424</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. scaurus</italic> subsp. <italic>violaceonitens </italic>Rob. Henry 1976 (holotype)</td><td align="left" rowspan="1" colspan="1">RH2190</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France</td><td align="left" rowspan="1" colspan="1">In forest with <italic>Fagus</italic>, <italic>Quercus </italic>and <italic>Carpinus</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. violaceonitens</italic> (Rob. Henry) Moënne-Locc. 2009</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">KF732425</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. scaurus</italic> var. <italic>notandus</italic> Bidaud 2009 (holotype)</td><td align="left" rowspan="1" colspan="1">AB94-08-32</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Loire, Jeansagnière</td><td align="left" rowspan="1" colspan="1">In montane coniferous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. scaurus</italic> (Fr.: Fr.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">21.08.1994</td><td align="left" rowspan="1" colspan="1">A. Bidaud</td><td align="left" rowspan="1" colspan="1">KF732426</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. serariicolor </italic>Rob. Henry 1985 (holotype)*</td><td align="left" rowspan="1" colspan="1">RH1731</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France</td><td align="left" rowspan="1" colspan="1">In montane coniferous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. papulosus </italic>Fr. 1838</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">KF732427</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. serarius</italic> Fr. 1838 (neotype)</td><td align="left" rowspan="1" colspan="1">CFP959</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Ång, Häggdånger, Torrom</td><td align="left" rowspan="1" colspan="1">In dry spruce forest on rich ground</td><td align="left" rowspan="1" colspan="1"><italic>C. serarius</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">11.08.1990</td><td align="left" rowspan="1" colspan="1">Brandrud et al.</td><td align="left" rowspan="1" colspan="1">KF732428</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. sobrius</italic> P. Karst. 1890 (type)</td><td align="left" rowspan="1" colspan="1">PAK3235</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, EH, Mustiala, Tammela</td><td align="left" rowspan="1" colspan="1">In frondose forest</td><td align="left" rowspan="1" colspan="1"><italic>C. sobrius</italic> P. Karst. 1890</td><td align="left" rowspan="1" colspan="1">23.09.1890</td><td align="left" rowspan="1" colspan="1">P.A. Karsten</td><td align="left" rowspan="1" colspan="1">KF732429</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. sphagnetorum</italic> Bidaud 1996 (holotype)</td><td align="left" rowspan="1" colspan="1">3746</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Savoie, Lac du Clou - Beaufortin</td><td align="left" rowspan="1" colspan="1">In subalpine zone under <italic>Picea</italic>, among<italic>Sphagnum</italic>and<italic>Vaccinium</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. pseudonaevosus</italic> Rob. Henry 1957</td><td align="left" rowspan="1" colspan="1">31.08.1991</td><td align="left" rowspan="1" colspan="1">A. Bidaud</td><td align="left" rowspan="1" colspan="1">KF732430</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. splendens</italic> var. <italic>papillatosporus</italic> Bidaud & Moënne-Locc. 2003 (holotype)</td><td align="left" rowspan="1" colspan="1">960</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Haute-Savoie, Les Puisots</td><td align="left" rowspan="1" colspan="1">In mixed forest</td><td align="left" rowspan="1" colspan="1"><italic>C. splendens</italic> var. <italic>papillatosporus</italic> Bidaud & Moënne-Locc. 2003</td><td align="left" rowspan="1" colspan="1">25.09.1988</td><td align="left" rowspan="1" colspan="1">Exposotion of Annecy</td><td align="left" rowspan="1" colspan="1">KF732431</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. squamosocephalus</italic> Bidaud, Moënne-Locc. & Reumaux 1999 (holotype)</td><td align="left" rowspan="1" colspan="1">99670</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Ardennes, Bois de la Brouille</td><td align="left" rowspan="1" colspan="1">Under <italic>Quercus</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. squamosocephalus</italic> Bidaud, Moënne-Locc. & Reumaux 1999</td><td align="left" rowspan="1" colspan="1">02.10.1998</td><td align="left" rowspan="1" colspan="1">P. Reumaux</td><td align="left" rowspan="1" colspan="1">KF732432</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subaccedens</italic> Rob. Henry 1989 (holotype)*</td><td align="left" rowspan="1" colspan="1">RH3170</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Languedoc-Roussillon- Cévennes region</td><td align="left" rowspan="1" colspan="1">Under <italic>Quercus ilex</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. subaccedens</italic> Rob. Henry 1989</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">KF732433</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subamaricatus</italic> Bidaud 2008 (holotype)</td><td align="left" rowspan="1" colspan="1">AB94-10-313</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Ain, Meyriat</td><td align="left" rowspan="1" colspan="1">In calcareous mixed forest of <italic>Fagus </italic>and <italic>Abies</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. tirolianus</italic> Bidaud, Moënne-Locc. & Reumaux 2005</td><td align="left" rowspan="1" colspan="1">16.10.1994</td><td align="left" rowspan="1" colspan="1">A. Bidaud</td><td align="left" rowspan="1" colspan="1">KF732434</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subaustralis</italic> A.H. Sm. & Hesler 1944 (holotype)*</td><td align="left" rowspan="1" colspan="1">14336</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, N.C., Great Smoky Mts National Park, Indian Gap</td><td align="left" rowspan="1" colspan="1">In coniferous forest</td><td align="left" rowspan="1" colspan="1"><italic>C.</italic><italic>crassus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">14.07.1942</td><td align="left" rowspan="1" colspan="1">L.R. Hesler</td><td align="left" rowspan="1" colspan="1">KF732435</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subbalteatus</italic> Kühner 1955 (syntype)*</td><td align="left" rowspan="1" colspan="1">8-12 (G00262072)</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Savoie, St-Bon, Praz</td><td align="left" rowspan="1" colspan="1">In <italic>Picea</italic> forest</td><td align="left" rowspan="1" colspan="1"><italic>C. balteatus</italic> (Fr.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">10.09.1927</td><td align="left" rowspan="1" colspan="1">R. Kühner</td><td align="left" rowspan="1" colspan="1">KF732437</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subcrassoides</italic> Moënne-Locc. & Remaux 1995 (holotype)</td><td align="left" rowspan="1" colspan="1">363</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Savoie, Col des Saisies</td><td align="left" rowspan="1" colspan="1">In herbaceous <italic>Picea abies </italic>forest</td><td align="left" rowspan="1" colspan="1"><italic>C. pseudonaevosus</italic> Rob. Henry 1957</td><td align="left" rowspan="1" colspan="1">29.09.1986</td><td align="left" rowspan="1" colspan="1">R. Campia</td><td align="left" rowspan="1" colspan="1">KF732438</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subcrassus</italic> Rob. Henry 1983 (holotype)*</td><td align="left" rowspan="1" colspan="1">RH71520</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Doubs, Seloncourt</td><td align="left" rowspan="1" colspan="1">Under conifers</td><td align="left" rowspan="1" colspan="1">C. <italic>crassus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">30.09.1979</td><td align="left" rowspan="1" colspan="1">From exhibition of Seloncourt</td><td align="left" rowspan="1" colspan="1">KF732439</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subcyanites</italic> Bidaud 2005 (holotype)</td><td align="left" rowspan="1" colspan="1">PML5304</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Ain, Le Poizat</td><td align="left" rowspan="1" colspan="1">Under <italic>Picea</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. cyanites</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">05.09.1999</td><td align="left" rowspan="1" colspan="1">A. Bidaud</td><td align="left" rowspan="1" colspan="1">KF732440</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subdecolorans</italic> M. Langl. & Reumaux 2000 (holotype)</td><td align="left" rowspan="1" colspan="1">4403</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Marne, Bazancourt</td><td align="left" rowspan="1" colspan="1">In deciduous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. subdecolorans</italic> M. Langl. & Reumaux 2000</td><td align="left" rowspan="1" colspan="1">20.10.1991</td><td align="left" rowspan="1" colspan="1">M. Langlois</td><td align="left" rowspan="1" colspan="1">KF732441</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subdecoloratus</italic> Reumaux 2000 (holotype)</td><td align="left" rowspan="1" colspan="1">3951</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Ardennes, Bois des Alleux</td><td align="left" rowspan="1" colspan="1">Under <italic>Betula</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. ochraceobrunneus</italic> Rob. Henry ex Bidaud, Moënne-Locc. & Reumaux 2000</td><td align="left" rowspan="1" colspan="1">30.10.1994</td><td align="left" rowspan="1" colspan="1">G. Laffond & P. Reumaux</td><td align="left" rowspan="1" colspan="1">KF732442</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subfoetens</italic> M.M. Moser & McKnight 1995 (holotype)</td><td align="left" rowspan="1" colspan="1">89/307</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">USA, Wyoming, Teton National Forest, Fourmile meadow</td><td align="left" rowspan="1" colspan="1">Under <italic>Picea engelmannii</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. subfoetens</italic> M.M. Moser & McKnight 1995</td><td align="left" rowspan="1" colspan="1">21.08.1989</td><td align="left" rowspan="1" colspan="1">K.H. McKnight</td><td align="left" rowspan="1" colspan="1">KF732443</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subfoetidus</italic> A.H. Sm. 1944 (holotype)</td><td align="left" rowspan="1" colspan="1">17778</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Washington, Olympic National Park, Olympic Hot Springs</td><td align="left" rowspan="1" colspan="1">Under conifers</td><td align="left" rowspan="1" colspan="1"><italic>C. subfoetidus</italic> A.H. Sm. 1944</td><td align="left" rowspan="1" colspan="1">11.10.1941</td><td align="left" rowspan="1" colspan="1">A.H. Smith</td><td align="left" rowspan="1" colspan="1">KF732444</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subfraudulosus </italic>Kytöv., Liimat. & Niskanen sp. nov. (holotype)</td><td align="left" rowspan="1" colspan="1">IK11-006</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Norway, Oppl, Lunner, Skøyenåsen</td><td align="left" rowspan="1" colspan="1">In <italic>Picea abies</italic> forest with <italic>Corylus</italic> on calcareous ground</td><td align="left" rowspan="1" colspan="1"><italic>C. subfraudulosus </italic>Kytöv., Liimat. & Niskanen sp. nov.</td><td align="left" rowspan="1" colspan="1">03.09.2011</td><td align="left" rowspan="1" colspan="1">I. Kytövuori</td><td align="left" rowspan="1" colspan="1">KF732564</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subfuligineus</italic> Bidaud 2008 (holotype)</td><td align="left" rowspan="1" colspan="1">AB97-10-339</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Ain, Petaray, Aranc</td><td align="left" rowspan="1" colspan="1">In calcareous coniferous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. subrugulosus</italic> Bidaud & Armada 2006</td><td align="left" rowspan="1" colspan="1">21.10.1997</td><td align="left" rowspan="1" colspan="1">A. Bidaud</td><td align="left" rowspan="1" colspan="1">KF732445</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subinops</italic> Reumaux 2009 (holotype)</td><td align="left" rowspan="1" colspan="1">PML5119</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Ardennes, bois de Toges</td><td align="left" rowspan="1" colspan="1">In deciduous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. subpurpurascens</italic> (Batsch) Fr. 1838</td><td align="left" rowspan="1" colspan="1">06.10.1996</td><td align="left" rowspan="1" colspan="1">P. Reumaux</td><td align="left" rowspan="1" colspan="1">KF732446</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. sublilacinopes </italic>Bidaud, Moënne-Locc. & Reumaux 2001 (holotype)</td><td align="left" rowspan="1" colspan="1">PML4819</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Seine-Maritime, La Londe-Rouvray</td><td align="left" rowspan="1" colspan="1">In calcareous deciduous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. sublilacinopes </italic>Bidaud, Moënne-Locc. & Reumaux 2001</td><td align="left" rowspan="1" colspan="1">24.10.1997</td><td align="left" rowspan="1" colspan="1">L.C. Malaval</td><td align="left" rowspan="1" colspan="1">KF732561</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subolivascens</italic> A.H. Sm. 1944 (holotype)</td><td align="left" rowspan="1" colspan="1">14311</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Washington, Olympic National Park, Deer Lake</td><td align="left" rowspan="1" colspan="1">Under conifers</td><td align="left" rowspan="1" colspan="1"><italic>C. subolivascens</italic> A.H. Sm. 1944</td><td align="left" rowspan="1" colspan="1">13.06.1939</td><td align="left" rowspan="1" colspan="1">A.H. Smith</td><td align="left" rowspan="1" colspan="1">KF732447</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subopimus</italic> Bidaud 1995 (holotype)</td><td align="left" rowspan="1" colspan="1">3477</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Haute-Savoie, Plateau de Dran, Les Glières</td><td align="left" rowspan="1" colspan="1">In herbaceous, calcareous <italic>Picea abies</italic> forest near <italic>Alnus viridis</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. balteatus</italic> (Fr.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">05.08.1993</td><td align="left" rowspan="1" colspan="1">A. Bidaud</td><td align="left" rowspan="1" colspan="1">KF732448</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subpurpurascens</italic> (Batsch) Fr. 1838 (epitype)</td><td align="left" rowspan="1" colspan="1">TN08-059</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, V, Turku</td><td align="left" rowspan="1" colspan="1">In deciduous forest of <italic>Quercus robur</italic>, <italic>Corylus i>avellana</italic> and some <italic>Betula</italic> on mull soil</td><td align="left" rowspan="1" colspan="1"><italic>C. subpurpurascens</italic> (Batsch) Fr. 1838</td><td align="left" rowspan="1" colspan="1">03.09.2008</td><td align="left" rowspan="1" colspan="1">K. Liimatainen & T. Niskanen</td><td align="left" rowspan="1" colspan="1">KF732449</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subpurpureophyllus</italic> A.H. Sm. 1939 (holotype)</td><td align="left" rowspan="1" colspan="1">8164</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, California, Crescent City</td><td align="left" rowspan="1" colspan="1">Under spruce</td><td align="left" rowspan="1" colspan="1"><italic>C. subpurpureophyllus</italic> A.H. Sm. 1939</td><td align="left" rowspan="1" colspan="1">28.10.1937</td><td align="left" rowspan="1" colspan="1">A.H. Smith</td><td align="left" rowspan="1" colspan="1">KF732450</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subrugulosus</italic> Bidaud & Armada 2006 (holotype)</td><td align="left" rowspan="1" colspan="1">AB05-10-263</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Isère, Treminis</td><td align="left" rowspan="1" colspan="1">In coniferous forest on calcareous soil</td><td align="left" rowspan="1" colspan="1"><italic>C. subrugulosus</italic> Bidaud & Armada 2006</td><td align="left" rowspan="1" colspan="1">06.10.2005</td><td align="left" rowspan="1" colspan="1">A. Bidaud & F. Armada</td><td align="left" rowspan="1" colspan="1">KF732451</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subsolitarius</italic> A.H. Sm. 1942 (holotype)</td><td align="left" rowspan="1" colspan="1">15377</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Michigan, Ann Arbor</td><td align="left" rowspan="1" colspan="1">On humus in oak-hickory woods</td><td align="left" rowspan="1" colspan="1"><italic>C. subsolitarius</italic> A.H. Sm. 1942</td><td align="left" rowspan="1" colspan="1">11.09.1940</td><td align="left" rowspan="1" colspan="1">A.H. Smith</td><td align="left" rowspan="1" colspan="1">KF732452</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subspadiceus</italic> Reumaux 1996 (holotype)</td><td align="left" rowspan="1" colspan="1">2780</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Haute-Vienne, Limousin</td><td align="left" rowspan="1" colspan="1">Under <italic>Picea abies</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. largus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">15.10.1990</td><td align="left" rowspan="1" colspan="1">Taupenot</td><td align="left" rowspan="1" colspan="1">KF732453</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subtortus</italic> (Pers.: Fr.) Fr. 1838 (neotype)</td><td align="left" rowspan="1" colspan="1">IK87-1510</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Sm, Hylte commune, Femsjö parish</td><td align="left" rowspan="1" colspan="1">In partly paludified spruce forest with some other tree species</td><td align="left" rowspan="1" colspan="1"><italic>C. subtortus</italic> (Pers.: Fr.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">18.09.1987</td><td align="left" rowspan="1" colspan="1">I. Kytövuori</td><td align="left" rowspan="1" colspan="1">KF732454</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subvariiformis</italic> Bidaud 2000 (holotype)</td><td align="left" rowspan="1" colspan="1">4663</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Bouches-du-Rhône, St Rémy de Provence</td><td align="left" rowspan="1" colspan="1">Under <italic>Quercus ilex</italic> on calcareous soil</td><td align="left" rowspan="1" colspan="1"><italic>C. luteocingulatus </italic>Bidaud & Fillion 1992</td><td align="left" rowspan="1" colspan="1">03.12.1997</td><td align="left" rowspan="1" colspan="1">G Riousset & L. Riousset</td><td align="left" rowspan="1" colspan="1">KF732455</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. superbus</italic> A.H. Sm. 1944 (holotype)</td><td align="left" rowspan="1" colspan="1">17680</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Washington, Olympic National Park, Olympic Hot Springs</td><td align="left" rowspan="1" colspan="1">On steep mountain slopes</td><td align="left" rowspan="1" colspan="1"><italic>C. superbus</italic> A.H. Sm. 1944</td><td align="left" rowspan="1" colspan="1">08.10.1941</td><td align="left" rowspan="1" colspan="1">A.H. Smith</td><td align="left" rowspan="1" colspan="1">KF732456</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. talimultiformis </italic>Kytöv., Liimat., Niskanen, A.F.S. Taylor & Sesli sp. nov. (holotype)</td><td align="left" rowspan="1" colspan="1">AT2004096</td><td align="left" rowspan="1" colspan="1">UPS</td><td align="left" rowspan="1" colspan="1">Sweden, Upl, Hässelby Park</td><td align="left" rowspan="1" colspan="1">In mixed forest</td><td align="left" rowspan="1" colspan="1"><italic>C. talimultiformis </italic>Kytöv., Liimat., Niskanen, A.F.S. Taylor & Sesli sp. nov.</td><td align="left" rowspan="1" colspan="1">11.07.2004</td><td align="left" rowspan="1" colspan="1">A. Taylor</td><td align="left" rowspan="1" colspan="1">KF732583</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. talus</italic> Fr. 1838 (neotype)</td><td align="left" rowspan="1" colspan="1">CFP832</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Jmt, Ragunda, Ragunda</td><td align="left" rowspan="1" colspan="1">In birch forest on rich ground (<italic>Betula</italic>,<italic>Populus</italic>, <italic>Pinus</italic>)</td><td align="left" rowspan="1" colspan="1"><italic>C. talus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">26.08.1989</td><td align="left" rowspan="1" colspan="1">T.E. Brandrud et al.</td><td align="left" rowspan="1" colspan="1">KF732457</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. thalliopurpurascens</italic> Rob. Henry 1995 (isotype)</td><td align="left" rowspan="1" colspan="1">RH458677</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Jura, Bois Boucot</td><td align="left" rowspan="1" colspan="1">In deciduous forest mainly of <italic>Carpinus</italic> and <italic>Fagus</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. herpeticus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">10.1986</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">KF732458</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. tiliae</italic> Brandrud 1996 (holotype)</td><td align="left" rowspan="1" colspan="1">TEB141-85</td><td align="left" rowspan="1" colspan="1">O</td><td align="left" rowspan="1" colspan="1">Norway, Akh, Bærum, Løkkeåsen</td><td align="left" rowspan="1" colspan="1">In dry calcareous soil, under <italic>Tilia cordata</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. tiliae</italic> Brandrud 1996</td><td align="left" rowspan="1" colspan="1">28.08.1985</td><td align="left" rowspan="1" colspan="1">T.E. Brandrud</td><td align="left" rowspan="1" colspan="1">KF732459</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. tirolianus</italic> Bidaud, Moënne-Locc. & Reumaux 2005 (holotype)</td><td align="left" rowspan="1" colspan="1">PML4341</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Jura, Moirans-en-Montagne</td><td align="left" rowspan="1" colspan="1">Under <italic>Picea</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. tirolianus</italic> Bidaud, Moënne-Locc. & Reumaux 2005</td><td align="left" rowspan="1" colspan="1">15.09.1996</td><td align="left" rowspan="1" colspan="1">A. Dégrange</td><td align="left" rowspan="1" colspan="1">KF732460</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. triumphalis</italic> Bidaud, Moënne-Locc. & Reumaux 2000 (holotype)</td><td align="left" rowspan="1" colspan="1">3950</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Allier, Forêt des Colettes</td><td align="left" rowspan="1" colspan="1">In coniferous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. triumphalis</italic> Bidaud, Moënne-Locc. & Reumaux 2000</td><td align="left" rowspan="1" colspan="1">23.10.1992</td><td align="left" rowspan="1" colspan="1">R. Chalange</td><td align="left" rowspan="1" colspan="1">KF732462</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. turmalis</italic> Fr. 1838 (neotype)</td><td align="left" rowspan="1" colspan="1">CFP716</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Mpd, Borgsjö, Julåsen</td><td align="left" rowspan="1" colspan="1">In herbaceous spruce forest</td><td align="left" rowspan="1" colspan="1"><italic>C. turmalis</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">20.08.1988</td><td align="left" rowspan="1" colspan="1">T.E. Brandrud et al.</td><td align="left" rowspan="1" colspan="1">KF732464</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. vacciniophilus</italic> Brandrud 1997 (holotype)</td><td align="left" rowspan="1" colspan="1">TEB17-88</td><td align="left" rowspan="1" colspan="1">O</td><td align="left" rowspan="1" colspan="1">Norway, Oppl, Lunner, Søndre Oppdalen</td><td align="left" rowspan="1" colspan="1">In forest of <italic>Picea abies</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. pseudonaevosus</italic> Rob. Henry 1957</td><td align="left" rowspan="1" colspan="1">09.08.1988</td><td align="left" rowspan="1" colspan="1">T.E. Brandrud</td><td align="left" rowspan="1" colspan="1">KF732465</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. van-campiae</italic> Cons. 2000*</td><td align="left" rowspan="1" colspan="1">871</td><td align="left" rowspan="1" colspan="1">MCVE</td><td align="left" rowspan="1" colspan="1">Italy</td><td align="left" rowspan="1" colspan="1">In mixed woodland with <italic>Abies alba </italic>and <italic>Fagus sylvatica</italic></td><td align="left" rowspan="1" colspan="1">C. <italic>van-campiae</italic> Cons. 2000</td><td align="left" rowspan="1" colspan="1">04.10.1995</td><td align="left" rowspan="1" colspan="1">G. Consiglio</td><td align="left" rowspan="1" colspan="1">JF907867</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. variecolor</italic> (Pers.: Fr.) Fr. 1838 (neotype)</td><td align="left" rowspan="1" colspan="1">CFP1021</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Gtl, Viklau, Tjakule</td><td align="left" rowspan="1" colspan="1">In spruce forest on calcareous ground</td><td align="left" rowspan="1" colspan="1"><italic>C. variecolor</italic> (Pers.: Fr.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">29.09.1990</td><td align="left" rowspan="1" colspan="1">T.E. Brandrud et al.</td><td align="left" rowspan="1" colspan="1">KF732466</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. variipes</italic> Rob. Henry 1977 (holotype)</td><td align="left" rowspan="1" colspan="1">RH5026</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Ardennes</td><td align="left" rowspan="1" colspan="1">In montane coniferous forest</td><td align="left" rowspan="1" colspan="1"><italic>C. variipes</italic> Rob. Henry 1977</td><td align="left" rowspan="1" colspan="1">Unknown</td><td align="left" rowspan="1" colspan="1">P. Reumaux</td><td align="left" rowspan="1" colspan="1">KF732467</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. variosimilis</italic> M.M. Moser & Ammirati 1999 (holotype)*</td><td align="left" rowspan="1" colspan="1">89/493</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">USA, Washington, Skagit Co., Trail to Easy Pass</td><td align="left" rowspan="1" colspan="1">In subalpine coniferous forest under <italic>Picea engelmannii</italic>, <italic>Abies lasiocarpa</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. variosimilis</italic> M.M. Moser & Ammirati 1999</td><td align="left" rowspan="1" colspan="1">12.09.1989</td><td align="left" rowspan="1" colspan="1">M. Moser</td><td align="left" rowspan="1" colspan="1">KF732468</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. varius</italic> (Schaeff.: Fr.) Fr. 1838 (neotype)</td><td align="left" rowspan="1" colspan="1">CFP801</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Ång, Häggdånger, Torrom</td><td align="left" rowspan="1" colspan="1">In spruce forest on rich ground</td><td align="left" rowspan="1" colspan="1"><italic>C. varius</italic> (Schaeff.: Fr.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">23.09.1988</td><td align="left" rowspan="1" colspan="1">T.E. Brandrud et al.</td><td align="left" rowspan="1" colspan="1">KF732469</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. velicopius</italic> Kauffman 1918 (lectotype)*</td><td align="left" rowspan="1" colspan="1">MICH10435</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Michigan, Washtenaw, Cascade Glen, Ann Arbor</td><td align="left" rowspan="1" colspan="1">Among fallen leaves in mixed or frondose woods</td><td align="left" rowspan="1" colspan="1"><italic>C. velicopius</italic> Kauffman 1918</td><td align="left" rowspan="1" colspan="1">29.09.1907</td><td align="left" rowspan="1" colspan="1">C.H. Kauffman</td><td align="left" rowspan="1" colspan="1">KF732470</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. veneris</italic> Bidaud, Moënne-Locc. & Reumaux 1996 (holotype)</td><td align="left" rowspan="1" colspan="1">2952</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Haute-Loire, Dunières</td><td align="left" rowspan="1" colspan="1">Under <italic>Abies alba</italic> on granitic soil</td><td align="left" rowspan="1" colspan="1"><italic>C. flavescentipes</italic> Reumaux 1996</td><td align="left" rowspan="1" colspan="1">02.10.1992</td><td align="left" rowspan="1" colspan="1">B. Renon</td><td align="left" rowspan="1" colspan="1">KF732471</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. violaceomaculatus</italic> Brandrud 1997 (holotype)</td><td align="left" rowspan="1" colspan="1">CFP1449</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Gtl, Bäl</td><td align="left" rowspan="1" colspan="1">In forest of <italic>Picea abies</italic> and <italic>Pinus sylvestris</italic> on calcareous soil</td><td align="left" rowspan="1" colspan="1"><italic>C. violaceomaculatus</italic> Brandrud 1997</td><td align="left" rowspan="1" colspan="1">28.09.1993</td><td align="left" rowspan="1" colspan="1">T.E. Brandrud et al.</td><td align="left" rowspan="1" colspan="1">KF732473</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. violaceorubens</italic> Moënne-Locc. & Reumaux 1990 (holotype)</td><td align="left" rowspan="1" colspan="1">PML005</td><td align="left" rowspan="1" colspan="1">G</td><td align="left" rowspan="1" colspan="1">France, Haute-Savoie, Pessière plantée d’Avernioz</td><td align="left" rowspan="1" colspan="1">Under <italic>Picea</italic> in needle litter</td><td align="left" rowspan="1" colspan="1"><italic>C. violaceorubens</italic> Moënne-Locc. & Reumaux 1990</td><td align="left" rowspan="1" colspan="1">21.06.1985</td><td align="left" rowspan="1" colspan="1">P. Moënne-Loccoz</td><td align="left" rowspan="1" colspan="1">KF732474</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. virentophyllus</italic> Kauffman 1918 (holotype)</td><td align="left" rowspan="1" colspan="1">MICH10439</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Michigan, Washtenaw, German Park Woods, SW of Ann Arbor</td><td align="left" rowspan="1" colspan="1">On the ground, among grasses in frondose woods of oak, maple, etc.</td><td align="left" rowspan="1" colspan="1"><italic>C. virentophyllus</italic> Kauffman 1918</td><td align="left" rowspan="1" colspan="1">11.10.1912</td><td align="left" rowspan="1" colspan="1">C.H. Kauffman</td><td align="left" rowspan="1" colspan="1">KF732475</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. viridirubescens</italic> M.M. Moser & Ammirati 1997 (holotype)</td><td align="left" rowspan="1" colspan="1">95/688</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">USA, California, Mendocino Co., on Forest Road 408 about 8 miles from village</td><td align="left" rowspan="1" colspan="1">Under <italic>Lithocarpus densiflora</italic> and <italic>Quercus garrayana</italic></td><td align="left" rowspan="1" colspan="1"><italic>C. viridirubescens</italic> M.M. Moser & Ammirati 1997</td><td align="left" rowspan="1" colspan="1">08.12.1995</td><td align="left" rowspan="1" colspan="1">M. Moser</td><td align="left" rowspan="1" colspan="1">KF732476</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. vixolivascens</italic> Rob. Henry 1992 (holotype)</td><td align="left" rowspan="1" colspan="1">RH89.123</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Habsheim, Forêt de la Hardt</td><td align="left" rowspan="1" colspan="1">In mixed deciduous forest with <italic>Carpinus</italic>, <italic>Acer</italic>, <italic>Crataegus</italic> and <italic>Quercus</italic> on subcalcareous soil</td><td align="left" rowspan="1" colspan="1"><italic>C. vixolivascens</italic> Rob. Henry 1992</td><td align="left" rowspan="1" colspan="1">1989</td><td align="left" rowspan="1" colspan="1">M.V. Rastetter</td><td align="left" rowspan="1" colspan="1">KF732477</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. volvatus</italic> A.H. Sm. 1939 (holotype)</td><td align="left" rowspan="1" colspan="1">8857</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, California, Crescent City</td><td align="left" rowspan="1" colspan="1">Under spruce</td><td align="left" rowspan="1" colspan="1"><italic>C. volvatus</italic> A.H. Sm. 1939</td><td align="left" rowspan="1" colspan="1">18.11.1937</td><td align="left" rowspan="1" colspan="1">A.H. Smith</td><td align="left" rowspan="1" colspan="1">KF732478</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. wiebeae</italic> Thiers & A.H. Sm. 1969 (holotype)</td><td align="left" rowspan="1" colspan="1">8051</td><td align="left" rowspan="1" colspan="1">MICH</td><td align="left" rowspan="1" colspan="1">USA, Oregon, Mt Hood, Camas Corral</td><td align="left" rowspan="1" colspan="1">Under fir</td><td align="left" rowspan="1" colspan="1"><italic>C. wiebeae</italic> Thiers & A.H. Sm. 1969</td><td align="left" rowspan="1" colspan="1">08.06.1958</td><td align="left" rowspan="1" colspan="1">E. Wiebe</td><td align="left" rowspan="1" colspan="1">KF732479</td></tr></tbody></table></table-wrap><table-wrap id="T2" orientation="portrait" position="float"><label>Table 2</label><caption><p><italic>Cortinarius</italic> specimens other than types included in the phylogenetic analysis. Short sequences excluded from the analysis marked with *.</p></caption><table frame="hsides" rules="groups"><thead><tr><th align="left" rowspan="1" colspan="1">Species</th><th align="left" rowspan="1" colspan="1">Voucher</th><th align="left" rowspan="1" colspan="1">Herb.</th><th align="left" rowspan="1" colspan="1">Locality</th><th align="left" rowspan="1" colspan="1">GenBank number</th></tr></thead><tbody><tr><td align="left" rowspan="1" colspan="1"><italic>C. acystidiosus</italic> Thiers 1960</td><td align="left" rowspan="1" colspan="1">CLO4681</td><td align="left" rowspan="1" colspan="1">TENN</td><td align="left" rowspan="1" colspan="1">USA</td><td align="left" rowspan="1" colspan="1">KF732419</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. aggregatus </italic>Kauffman 1918*</td><td align="left" rowspan="1" colspan="1">TN10-179</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Canada, QC, Papineauville</td><td align="left" rowspan="1" colspan="1">KF732512</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. alnobetulae</italic> Kühner 1989</td><td align="left" rowspan="1" colspan="1">JFA12247</td><td align="left" rowspan="1" colspan="1">WTU</td><td align="left" rowspan="1" colspan="1">Italy, Passo del Rolle</td><td align="left" rowspan="1" colspan="1">EU655672</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. alticaudus</italic> Reumaux 2008</td><td align="left" rowspan="1" colspan="1">IK09-1402b</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, PK, Kesälahti</td><td align="left" rowspan="1" colspan="1">KF732610</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. areni-silvae</italic> (Brandrud) Brandrud 2012</td><td align="left" rowspan="1" colspan="1">IK95-341</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, InL, Inari</td><td align="left" rowspan="1" colspan="1">KF732611</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. argutus</italic> Fr. 1838 s. auct</td><td align="left" rowspan="1" colspan="1">O-60164</td><td align="left" rowspan="1" colspan="1">O</td><td align="left" rowspan="1" colspan="1">Norway</td><td align="left" rowspan="1" colspan="1">AY669535</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. argutus</italic> s. auct.</td><td align="left" rowspan="1" colspan="1">T44</td><td align="left" rowspan="1" colspan="1">O</td><td align="left" rowspan="1" colspan="1">Norway</td><td align="left" rowspan="1" colspan="1">UDB000138</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. badiolatus</italic> (M.M. Moser) M.M. Moser 1967</td><td align="left" rowspan="1" colspan="1">IK98-1029</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Sweden, Nb, Pajala</td><td align="left" rowspan="1" colspan="1">KF732612</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. balteatialutaceus </italic>Kytöv., Liimat. & Niskanen sp. nov.</td><td align="left" rowspan="1" colspan="1">JV10452, TUR5282</td><td align="left" rowspan="1" colspan="1">TUR</td><td align="left" rowspan="1" colspan="1">Finland, InL, Utsjoki</td><td align="left" rowspan="1" colspan="1">KF732587</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. balteatialutaceus*</italic></td><td align="left" rowspan="1" colspan="1">JR100900</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Norway, S&F, Sogndal</td><td align="left" rowspan="1" colspan="1">KF732588</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. balteatibulbosus</italic> Kytöv., Niskanen, Liimat., Bojantchev & A.F.S. Taylor sp. nov.</td><td align="left" rowspan="1" colspan="1">DBB33060</td><td align="left" rowspan="1" colspan="1">UC</td><td align="left" rowspan="1" colspan="1">Bulgaria, Saranzi</td><td align="left" rowspan="1" colspan="1">KF732590</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. balteatibulbosus*</italic></td><td align="left" rowspan="1" colspan="1">DBB36892</td><td align="left" rowspan="1" colspan="1">UC</td><td align="left" rowspan="1" colspan="1">Sweden, Järfälla</td><td align="left" rowspan="1" colspan="1">KF732591</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. balteatibulbosus</italic></td><td align="left" rowspan="1" colspan="1">H6027358</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, V, Vihti</td><td align="left" rowspan="1" colspan="1">KF732592</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. balteatibulbosus</italic></td><td align="left" rowspan="1" colspan="1">IK93-639, H6032755</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, U, Espoo</td><td align="left" rowspan="1" colspan="1">KF732593</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. balteatibulbosus</italic></td><td align="left" rowspan="1" colspan="1">IK04-044</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, U, Vantaa</td><td align="left" rowspan="1" colspan="1">KF732594</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. balteatibulbosus*</italic></td><td align="left" rowspan="1" colspan="1">H6032413</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, U, Helsinki</td><td align="left" rowspan="1" colspan="1">KF732595</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. balteatibulbosus</italic></td><td align="left" rowspan="1" colspan="1">AT2004091</td><td align="left" rowspan="1" colspan="1">UPS</td><td align="left" rowspan="1" colspan="1">Sweden, Upl, Berthåga graveyard</td><td align="left" rowspan="1" colspan="1">UDB001132</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. balteatibulbosus</italic></td><td align="left" rowspan="1" colspan="1">AT2004045</td><td align="left" rowspan="1" colspan="1">UPS</td><td align="left" rowspan="1" colspan="1">Sweden, Upl, Slottsbacken</td><td align="left" rowspan="1" colspan="1">UDB000711</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. balteatibulbosus</italic></td><td align="left" rowspan="1" colspan="1">AT2004088</td><td align="left" rowspan="1" colspan="1">UPS</td><td align="left" rowspan="1" colspan="1">Sweden, Upl, Stenhagen</td><td align="left" rowspan="1" colspan="1">UDB000715</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. balteatibulbosus*</italic></td><td align="left" rowspan="1" colspan="1">AT2004127</td><td align="left" rowspan="1" colspan="1">UPS</td><td align="left" rowspan="1" colspan="1">Sweden, Upl, Berthåga graveyard</td><td align="left" rowspan="1" colspan="1">UDB000722</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. balteaticlavatus</italic> Kytöv., Liimat. & Niskanen sp. nov.</td><td align="left" rowspan="1" colspan="1">IK95-382, H6032729</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, InL, Inari</td><td align="left" rowspan="1" colspan="1">KF732597</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. balteaticlavatus</italic></td><td align="left" rowspan="1" colspan="1">IK96-782, H6032415</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, ES, Mäntyharju</td><td align="left" rowspan="1" colspan="1">KF732598</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. balteaticlavatus</italic></td><td align="left" rowspan="1" colspan="1">IK08-584, H6033533</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, Ks, Taivalkoski Jurmu-Kurtti</td><td align="left" rowspan="1" colspan="1">KF732599</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. balteatoalbus</italic> Rob. Henry 1985*</td><td align="left" rowspan="1" colspan="1">CFP1083</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">France, Ain, Ordonnaz</td><td align="left" rowspan="1" colspan="1">KF732613</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. balteatoalbus</italic></td><td align="left" rowspan="1" colspan="1">IK97-761b</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Sweden, Jmt, Revsund</td><td align="left" rowspan="1" colspan="1">KF732614</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. bigelowii</italic> Thiers & A.H. Sm. 1969</td><td align="left" rowspan="1" colspan="1">OSC-81327</td><td align="left" rowspan="1" colspan="1">OSC</td><td align="left" rowspan="1" colspan="1">USA, OR, Klamath</td><td align="left" rowspan="1" colspan="1">EU056976</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. boreicyanites </italic>Kytöv., Liimat., Niskanen & A.F.S. Taylor sp. nov.</td><td align="left" rowspan="1" colspan="1">TN03-112</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, PS, Kuopio</td><td align="left" rowspan="1" colspan="1">KF732500</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. boreicyanites </italic></td><td align="left" rowspan="1" colspan="1">AT2010203</td><td align="left" rowspan="1" colspan="1">UPS</td><td align="left" rowspan="1" colspan="1">Great Britain, Scotland, Grampian</td><td align="left" rowspan="1" colspan="1">KF732501</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. boreidionysae </italic>Kytöv., Liimat., Niskanen & Dima sp. nov.</td><td align="left" rowspan="1" colspan="1">IK07-245</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, OP, Kiiminki</td><td align="left" rowspan="1" colspan="1">KF732489</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. borgsjoeensis</italic> Brandrud 1992</td><td align="left" rowspan="1" colspan="1">IK07-1029</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, Ks, Salla</td><td align="left" rowspan="1" colspan="1">KF732615</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. brunneiaurantius </italic>Kytöv., Liimat. & Niskanen sp. nov.</td><td align="left" rowspan="1" colspan="1">IK93-644, H6032406</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, U, Espoo</td><td align="left" rowspan="1" colspan="1">KF732601</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. brunneiaurantius </italic></td><td align="left" rowspan="1" colspan="1">JV17979b, H6032422</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, V, Turku</td><td align="left" rowspan="1" colspan="1">KF732602</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. caerulescens</italic> (Schaeff.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">UL98-88, TUB012146</td><td align="left" rowspan="1" colspan="1">TUB</td><td align="left" rowspan="1" colspan="1">Germany</td><td align="left" rowspan="1" colspan="1">AY174863</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. caesiocinctus </italic></td><td align="left" rowspan="1" colspan="1">IK97-1573</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland</td><td align="left" rowspan="1" colspan="1">DQ663241</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. caesiocolor</italic> Kytöv., Liimat. & Niskanen sp. nov.</td><td align="left" rowspan="1" colspan="1">IK97-207, H6032730</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, U, Helsinki</td><td align="left" rowspan="1" colspan="1">KF732604</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. caesiolamellatus</italic> (Bidaud) Kytöv., Liimat., Niskanen, Brandrud, Frøslev & A.F.S. Taylor comb. nov.</td><td align="left" rowspan="1" colspan="1">TN09-201</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">USA, WA, Olympic peninsula</td><td align="left" rowspan="1" colspan="1">KF732571</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. caesiolamellatus </italic></td><td align="left" rowspan="1" colspan="1">AT2005085</td><td align="left" rowspan="1" colspan="1">UPS</td><td align="left" rowspan="1" colspan="1">Sweden, Upl</td><td align="left" rowspan="1" colspan="1">UDB002202</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. caesiolamellatus </italic></td><td align="left" rowspan="1" colspan="1">11841</td><td align="left" rowspan="1" colspan="1">TUB</td><td align="left" rowspan="1" colspan="1">Germany</td><td align="left" rowspan="1" colspan="1">AY669531</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. caesiophylloides</italic> Kytöv., Liimat., Niskanen, Brandrud & Frøslev sp. nov.</td><td align="left" rowspan="1" colspan="1">IK92-3044</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, SoL, Pelkosenniemi</td><td align="left" rowspan="1" colspan="1">KF732573</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. caesiophylloides</italic></td><td align="left" rowspan="1" colspan="1">IK08-1554</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, Kn, Paltamo</td><td align="left" rowspan="1" colspan="1">KF732574</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. caesiophylloides</italic></td><td align="left" rowspan="1" colspan="1">TF28</td><td rowspan="1" colspan="1"></td><td rowspan="1" colspan="1"></td><td align="left" rowspan="1" colspan="1">KF732576</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. caesiophylloides</italic></td><td align="left" rowspan="1" colspan="1">TF40</td><td rowspan="1" colspan="1"></td><td rowspan="1" colspan="1"></td><td align="left" rowspan="1" colspan="1">KF732575</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. calojanthinus</italic> M.M. Moser & Ammirati 1999</td><td align="left" rowspan="1" colspan="1">TN08-129</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, V, Västanfjärd</td><td align="left" rowspan="1" colspan="1">KF732538</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. calojanthinus</italic></td><td align="left" rowspan="1" colspan="1">TSJ2003-005</td><td align="left" rowspan="1" colspan="1">C</td><td align="left" rowspan="1" colspan="1">Sweden, Vg</td><td align="left" rowspan="1" colspan="1">DQ663281</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. castaneicolor</italic> A.H. Sm. 1944</td><td align="left" rowspan="1" colspan="1">SMI38</td><td align="left" rowspan="1" colspan="1">UBC</td><td align="left" rowspan="1" colspan="1">Canada, BC</td><td align="left" rowspan="1" colspan="1">FJ157126</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. chromataphilus</italic> Rob. Henry 1989</td><td align="left" rowspan="1" colspan="1">TUB 011862</td><td align="left" rowspan="1" colspan="1">TUB</td><td align="left" rowspan="1" colspan="1">Germany</td><td align="left" rowspan="1" colspan="1">AY669550</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. citriolens</italic> Ammirati & M.M. Moser 1999</td><td align="left" rowspan="1" colspan="1">19970154</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">USA, WY</td><td align="left" rowspan="1" colspan="1">AF325607</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. claricolor</italic> (Fr.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">TN07-138</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, Ks, Kuusamo</td><td align="left" rowspan="1" colspan="1">KF732616</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. claricolor</italic></td><td align="left" rowspan="1" colspan="1">TUB 011852</td><td align="left" rowspan="1" colspan="1">TUB</td><td align="left" rowspan="1" colspan="1">Germany</td><td align="left" rowspan="1" colspan="1">AY669522</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cobaltinus </italic>Kytöv., Liimat. & Niskanen 2013</td><td align="left" rowspan="1" colspan="1">TSJ2006068</td><td align="left" rowspan="1" colspan="1">C</td><td align="left" rowspan="1" colspan="1">Norway, Oppl, Jevnaker</td><td align="left" rowspan="1" colspan="1">KF673471</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cobaltinus </italic></td><td align="left" rowspan="1" colspan="1">TF2006-103</td><td align="left" rowspan="1" colspan="1">C</td><td align="left" rowspan="1" colspan="1">Norway, Oppl, Jevnaker</td><td align="left" rowspan="1" colspan="1">KF673472</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cremeiamarescens </italic>Kytöv., Liimat. & Niskanen sp. nov.</td><td align="left" rowspan="1" colspan="1">IK00-027</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, U, Siuntio</td><td align="left" rowspan="1" colspan="1">KF732494</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cremeiamarescens</italic></td><td align="left" rowspan="1" colspan="1">TN06-273</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, V, Parainen</td><td align="left" rowspan="1" colspan="1">KF732495</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cremeiamarescens</italic></td><td align="left" rowspan="1" colspan="1">OCS153F</td><td align="left" rowspan="1" colspan="1">UBC</td><td align="left" rowspan="1" colspan="1">Canada, BC</td><td align="left" rowspan="1" colspan="1">EF218754</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cremeiamarescens</italic></td><td align="left" rowspan="1" colspan="1">clone AlASOILE08</td><td rowspan="1" colspan="1"></td><td align="left" rowspan="1" colspan="1">USA, AK</td><td align="left" rowspan="1" colspan="1">JN889840</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cruentipellis </italic>Kytöv., Liimat., Niskanen & Dima sp. nov.</td><td align="left" rowspan="1" colspan="1">IK01-055</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Estonia, Hiiumaa, Pühalepa</td><td align="left" rowspan="1" colspan="1">KF732540</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cruentipellis</italic></td><td align="left" rowspan="1" colspan="1">IK98-2503</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Sweden, Öl, Algutsrum</td><td align="left" rowspan="1" colspan="1">KF732541</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cruentipellis</italic></td><td align="left" rowspan="1" colspan="1">IK11-008</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Norway, Akh, Asker</td><td align="left" rowspan="1" colspan="1">KF732542</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cumatilis</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">IK92-2927</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, V, Lohja</td><td align="left" rowspan="1" colspan="1">KF732517</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cumatilis</italic></td><td align="left" rowspan="1" colspan="1">H6016455</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, U, Espoo</td><td align="left" rowspan="1" colspan="1">KF732643</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cupreorufus</italic> Brandrud 1994</td><td align="left" rowspan="1" colspan="1">TN02-700</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, Ks</td><td align="left" rowspan="1" colspan="1">KF732548</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cupreorufus</italic></td><td align="left" rowspan="1" colspan="1">TN07-378</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">USA, WA, Olympic Peninsula</td><td align="left" rowspan="1" colspan="1">KF732549</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cupreorufus</italic></td><td align="left" rowspan="1" colspan="1">TN11-129</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">USA, AK, Fairbanks</td><td align="left" rowspan="1" colspan="1">KF732550</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cyanites</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">IK98-1476</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, U, Espoo</td><td align="left" rowspan="1" colspan="1">KF732502</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cyanites</italic></td><td align="left" rowspan="1" colspan="1">AT2000154</td><td align="left" rowspan="1" colspan="1">UPS</td><td align="left" rowspan="1" colspan="1">Sweden, Upl, Uppsala</td><td align="left" rowspan="1" colspan="1">KF732503</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cyanites*</italic></td><td align="left" rowspan="1" colspan="1">AT2004139</td><td align="left" rowspan="1" colspan="1">UPS</td><td align="left" rowspan="1" colspan="1">Sweden, Upl, Uppsala</td><td align="left" rowspan="1" colspan="1">KF732391</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. cyanites</italic></td><td align="left" rowspan="1" colspan="1">AT2004089</td><td align="left" rowspan="1" colspan="1">UPS</td><td align="left" rowspan="1" colspan="1">Sweden, Upl, Uppsala</td><td align="left" rowspan="1" colspan="1">UDB001154</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. dionysae </italic>Rob. Henry 1933 s. auct</td><td align="left" rowspan="1" colspan="1">TUB011450</td><td align="left" rowspan="1" colspan="1">TUB</td><td align="left" rowspan="1" colspan="1">Germany</td><td align="left" rowspan="1" colspan="1">AY174813</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. dionysae </italic>s. auct</td><td align="left" rowspan="1" colspan="1">TSJ2000-102</td><td align="left" rowspan="1" colspan="1">C</td><td align="left" rowspan="1" colspan="1">Germany, Bayern</td><td align="left" rowspan="1" colspan="1">DQ083782</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. dionysae</italic> var. <italic>avellanus</italic> Rob. Henry ex Bidaud & Carteret 2008</td><td align="left" rowspan="1" colspan="1">IK94-1745a</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">France, Ain, Oyonnax SE</td><td align="left" rowspan="1" colspan="1">KF732606</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. eliae</italic> Bidaud, Moënne-Locc. & Reumaux 1996</td><td align="left" rowspan="1" colspan="1">41098</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France</td><td align="left" rowspan="1" colspan="1">KF732617</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. elotoides</italic> M.M. Moser & McKnight 1995</td><td align="left" rowspan="1" colspan="1">CFP503</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Vg</td><td align="left" rowspan="1" colspan="1">DQ663395</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. evosmus</italic> Joachim ex Bidaud & Reumaux 2006</td><td align="left" rowspan="1" colspan="1">TSJ2004-014</td><td align="left" rowspan="1" colspan="1">C</td><td align="left" rowspan="1" colspan="1">Denmark</td><td align="left" rowspan="1" colspan="1">DQ663368</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. evosmus</italic></td><td align="left" rowspan="1" colspan="1">TUB011399</td><td align="left" rowspan="1" colspan="1">TUB</td><td align="left" rowspan="1" colspan="1">Germany</td><td align="left" rowspan="1" colspan="1">AY174815</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. flavipallens </italic>Kytöv., Liimat. & Niskanen sp. nov.</td><td align="left" rowspan="1" colspan="1">H6032393</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, LK, Parikkala</td><td align="left" rowspan="1" colspan="1">KF732555</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. flavipallens</italic></td><td align="left" rowspan="1" colspan="1">TK368</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, PeP, Tervola</td><td align="left" rowspan="1" colspan="1">KF732556</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. </italic>aff. <italic>flavipallens</italic></td><td align="left" rowspan="1" colspan="1">SMIA06</td><td align="left" rowspan="1" colspan="1">UBC</td><td align="left" rowspan="1" colspan="1">Canada, BC</td><td align="left" rowspan="1" colspan="1">FJ039640</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. flavobulbus</italic> Ammirati & M.M. Moser 1997</td><td align="left" rowspan="1" colspan="1">JFA11826</td><td align="left" rowspan="1" colspan="1">WTU</td><td align="left" rowspan="1" colspan="1">USA, CA, Del Norte</td><td align="left" rowspan="1" colspan="1">EU057017</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. fraudulosus</italic> Britzelm. 1885</td><td align="left" rowspan="1" colspan="1">TU106876</td><td align="left" rowspan="1" colspan="1">TU</td><td align="left" rowspan="1" colspan="1">Estonia, Saare, Kihelkonna</td><td align="left" rowspan="1" colspan="1">UDB011906</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. fraudulosus</italic></td><td align="left" rowspan="1" colspan="1">19960696</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">Italy, Bozen</td><td align="left" rowspan="1" colspan="1">UDB001036</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. fuligineofolius</italic> (M.M. Moser) M.M. Moser & Peintner 2002</td><td align="left" rowspan="1" colspan="1">19910576</td><td align="left" rowspan="1" colspan="1">IB</td><td rowspan="1" colspan="1"></td><td align="left" rowspan="1" colspan="1">AF478578</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. fuligineofolius</italic></td><td align="left" rowspan="1" colspan="1">19910682</td><td align="left" rowspan="1" colspan="1">IB</td><td rowspan="1" colspan="1"></td><td align="left" rowspan="1" colspan="1">AF478577</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. gentianeus</italic> Bidaud 1993</td><td align="left" rowspan="1" colspan="1">IK11-015</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Sweden, Gtl, Alskog and När parish</td><td align="left" rowspan="1" colspan="1">KF732496</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. gentianeus</italic></td><td align="left" rowspan="1" colspan="1">IK97-1378</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, Kn, Suomussalmi</td><td align="left" rowspan="1" colspan="1">KF732497</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. gentianeus</italic></td><td align="left" rowspan="1" colspan="1">IK09-1525</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, ES, Kerimäki</td><td align="left" rowspan="1" colspan="1">KF732498</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. gentianeus</italic></td><td align="left" rowspan="1" colspan="1">CFP775</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Dlr, Rättvik</td><td align="left" rowspan="1" colspan="1">KF732499</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. gentianeus</italic></td><td align="left" rowspan="1" colspan="1">TUB011845</td><td align="left" rowspan="1" colspan="1">TUB</td><td align="left" rowspan="1" colspan="1">Germany</td><td align="left" rowspan="1" colspan="1">AY669519</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. georgiolens</italic> Rob. Henry 1986</td><td align="left" rowspan="1" colspan="1">IK98-2504</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Sweden, Öl, Algutsrum</td><td align="left" rowspan="1" colspan="1">KF732618</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. glaucocephalus</italic> M.M. Moser, Ammirati & Halling 2000</td><td align="left" rowspan="1" colspan="1">F19524</td><td align="left" rowspan="1" colspan="1">UBC</td><td align="left" rowspan="1" colspan="1">Canada, BC</td><td align="left" rowspan="1" colspan="1">HQ604682</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. glaucopus</italic> (Schaeff.: Fr.) Gray 1821</td><td align="left" rowspan="1" colspan="1">IK92-1105a</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, SoL, Pelkosenniemi</td><td align="left" rowspan="1" colspan="1">KF732619</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. glaucopus</italic></td><td align="left" rowspan="1" colspan="1">SMIA20</td><td align="left" rowspan="1" colspan="1">UBC</td><td align="left" rowspan="1" colspan="1">Canada, BC</td><td align="left" rowspan="1" colspan="1">FJ039616</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. glaucopus</italic></td><td align="left" rowspan="1" colspan="1">clone NWBRO20</td><td rowspan="1" colspan="1"></td><td align="left" rowspan="1" colspan="1">Canada, BC</td><td align="left" rowspan="1" colspan="1">EU645632</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. glaucopus</italic></td><td align="left" rowspan="1" colspan="1">SMI293</td><td align="left" rowspan="1" colspan="1">UBC</td><td align="left" rowspan="1" colspan="1">Canada, BC</td><td align="left" rowspan="1" colspan="1">FJ039615</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. gracilior</italic> (Jul. Schäff. ex M.M. Moser) M.M. Moser 1967</td><td align="left" rowspan="1" colspan="1">JV19538</td><td align="left" rowspan="1" colspan="1">H, TUR</td><td align="left" rowspan="1" colspan="1">Italy, Veneto, Belluno</td><td align="left" rowspan="1" colspan="1">KF732492</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. herpeticus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">IK92-593</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Norway, Troms, Storfjord</td><td align="left" rowspan="1" colspan="1">KF732507</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. herpeticus</italic></td><td align="left" rowspan="1" colspan="1">DB2142</td><td align="left" rowspan="1" colspan="1">BP</td><td align="left" rowspan="1" colspan="1">Hungary, Vas, Ispánk</td><td align="left" rowspan="1" colspan="1">KF732508</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. herpeticus</italic></td><td align="left" rowspan="1" colspan="1">TUB011456</td><td align="left" rowspan="1" colspan="1">TUB</td><td align="left" rowspan="1" colspan="1">Germany</td><td align="left" rowspan="1" colspan="1">AY174808</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. herpeticus</italic></td><td align="left" rowspan="1" colspan="1">9204</td><td rowspan="1" colspan="1"></td><td align="left" rowspan="1" colspan="1">Italy</td><td align="left" rowspan="1" colspan="1">JF907950</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. immixtus</italic> Kauffman 1932</td><td align="left" rowspan="1" colspan="1">F17145OC73</td><td align="left" rowspan="1" colspan="1">UBC</td><td align="left" rowspan="1" colspan="1">Canada, BC</td><td align="left" rowspan="1" colspan="1">GQ159888</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. infractiflavus </italic>(M.M. Moser) Kytöv., Niskanen, Liimat., Bojantchev & Ammirati stat. nov. & nom. nov.</td><td align="left" rowspan="1" colspan="1">DBB19634</td><td align="left" rowspan="1" colspan="1">UC</td><td align="left" rowspan="1" colspan="1">Bulgaria, Pirin Mts</td><td align="left" rowspan="1" colspan="1">KF732534</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. infractiflavus</italic></td><td align="left" rowspan="1" colspan="1">IK92-1109</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, SoL, Pelkosenniemi</td><td align="left" rowspan="1" colspan="1">KF732535</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. infractiflavus</italic></td><td align="left" rowspan="1" colspan="1">SMI286</td><td align="left" rowspan="1" colspan="1">UBC</td><td align="left" rowspan="1" colspan="1">Canada, BC</td><td align="left" rowspan="1" colspan="1">FJ039612</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. infractus</italic> (Pers.: Fr.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">TN02-832</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, Ks, Oulanka</td><td align="left" rowspan="1" colspan="1">KF732523</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. infractus </italic></td><td align="left" rowspan="1" colspan="1">IK93-223</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, EH, Pälkäne</td><td align="left" rowspan="1" colspan="1">KF732524</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. juxtadibaphus</italic> Rob. Henry 1983</td><td align="left" rowspan="1" colspan="1">CFP1108</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">France, Ain</td><td align="left" rowspan="1" colspan="1">DQ663286</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. largus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">CFP1085</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">France, Ain, Ordonnaz, Penant</td><td align="left" rowspan="1" colspan="1">KF732333</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. leonicolor</italic> Reumaux 2001</td><td align="left" rowspan="1" colspan="1">CFP852</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Belgium, Brabant, Tervuren</td><td align="left" rowspan="1" colspan="1">KF732490</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. luteicolor </italic>(A.H. Sm.) Ammirati, Bojantchev, Niskanen & Liimat. stat. nov. & nom. nov.</td><td align="left" rowspan="1" colspan="1">DBB46740</td><td align="left" rowspan="1" colspan="1">UC</td><td align="left" rowspan="1" colspan="1">USA, CA, Yosemite Nat’l Park</td><td align="left" rowspan="1" colspan="1">KF732546</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. luteicolor</italic></td><td align="left" rowspan="1" colspan="1">DBB38211</td><td align="left" rowspan="1" colspan="1">UC</td><td align="left" rowspan="1" colspan="1">USA, CA, Yosemite Nat’l Park</td><td align="left" rowspan="1" colspan="1">KF732547</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. luteicolor</italic></td><td align="left" rowspan="1" colspan="1">JMB10-06-2007-03</td><td rowspan="1" colspan="1"></td><td align="left" rowspan="1" colspan="1">USA</td><td align="left" rowspan="1" colspan="1">HM068562</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. luteicolor</italic></td><td align="left" rowspan="1" colspan="1">VMS13</td><td rowspan="1" colspan="1"></td><td align="left" rowspan="1" colspan="1">Canada, BC</td><td align="left" rowspan="1" colspan="1">FJ717511</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. luteicolor</italic></td><td align="left" rowspan="1" colspan="1">JFA11701</td><td align="left" rowspan="1" colspan="1">WTU</td><td align="left" rowspan="1" colspan="1">USA, OR, Clackamus</td><td align="left" rowspan="1" colspan="1">EU057024</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. luteobrunnescens</italic> A.H. Sm. 1944</td><td align="left" rowspan="1" colspan="1">IK97-2298</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, V, Lohja</td><td align="left" rowspan="1" colspan="1">KF732620</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. maculatocaespitosus</italic> Bidaud 2009</td><td align="left" rowspan="1" colspan="1">TUB011396</td><td align="left" rowspan="1" colspan="1">TUB</td><td rowspan="1" colspan="1"></td><td align="left" rowspan="1" colspan="1">AY174780</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. meinhardii </italic>Bon 1986 s. auct.</td><td align="left" rowspan="1" colspan="1">TN05-168</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, Ks, Kuusamo</td><td align="left" rowspan="1" colspan="1">KF732551</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. meinhardii </italic>s. auct.</td><td align="left" rowspan="1" colspan="1">TUB011443</td><td align="left" rowspan="1" colspan="1">TUB</td><td align="left" rowspan="1" colspan="1">Germany</td><td align="left" rowspan="1" colspan="1">AY174840</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. melleicarneus </italic>Kytöv., Liimat., Niskanen & Brandrud sp. nov.</td><td align="left" rowspan="1" colspan="1">O-125960</td><td align="left" rowspan="1" colspan="1">O</td><td align="left" rowspan="1" colspan="1">Norway, AA, Grimstad</td><td align="left" rowspan="1" colspan="1">AY669533</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. mendax</italic> Bidaud, Mahiques & Reumaux 2011</td><td align="left" rowspan="1" colspan="1">TN06-291</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, A, Sund</td><td align="left" rowspan="1" colspan="1">KF732515</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. mendax</italic></td><td align="left" rowspan="1" colspan="1">TN06-157</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, PK, Kitee</td><td align="left" rowspan="1" colspan="1">KF732516</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. metarius</italic> Kauffman 1921</td><td align="left" rowspan="1" colspan="1">PML5204</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France, Ain</td><td align="left" rowspan="1" colspan="1">DQ663236</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. misermontii</italic> Chevassut & Rob. Henry 1986</td><td align="left" rowspan="1" colspan="1">IK98-2417</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Sweden, Öl, Högsrum</td><td align="left" rowspan="1" colspan="1">KF732621</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. misermontii</italic></td><td align="left" rowspan="1" colspan="1">IK872172</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Spain, Catalonia, Girona</td><td align="left" rowspan="1" colspan="1">KF732622</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. montanus</italic> Kauffman 1932</td><td align="left" rowspan="1" colspan="1">OSC1064150</td><td align="left" rowspan="1" colspan="1">OSC</td><td align="left" rowspan="1" colspan="1">USA, OR</td><td align="left" rowspan="1" colspan="1">EU525972</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. montanus</italic></td><td align="left" rowspan="1" colspan="1">DBB00174</td><td align="left" rowspan="1" colspan="1">UC</td><td align="left" rowspan="1" colspan="1">USA, OR</td><td align="left" rowspan="1" colspan="1">JF795378</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. multiformis</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">IK08-1857</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, PS, Sonkajärvi</td><td align="left" rowspan="1" colspan="1">KF732623</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. multiformis</italic></td><td align="left" rowspan="1" colspan="1">TN06-139</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, PK, Kitee</td><td align="left" rowspan="1" colspan="1">KF732624</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. multiformis</italic></td><td align="left" rowspan="1" colspan="1">IK98-1401</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, EH Vilppula</td><td align="left" rowspan="1" colspan="1">KF732625</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. multiformis</italic></td><td align="left" rowspan="1" colspan="1">TN05-247</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Norway, Hord, Voss</td><td align="left" rowspan="1" colspan="1">KF732626</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. multiformis</italic></td><td align="left" rowspan="1" colspan="1">19940224</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">Sweden, Sm</td><td align="left" rowspan="1" colspan="1">UDB001004</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. multiformis</italic></td><td align="left" rowspan="1" colspan="1">TAAM128778</td><td align="left" rowspan="1" colspan="1">TAAM</td><td align="left" rowspan="1" colspan="1">Estonia, Tartu</td><td align="left" rowspan="1" colspan="1">UDB016114</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. multiformis</italic></td><td align="left" rowspan="1" colspan="1">TU105180</td><td align="left" rowspan="1" colspan="1">TU</td><td align="left" rowspan="1" colspan="1">Estonia, Voru</td><td align="left" rowspan="1" colspan="1">UDB016130</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. multiformis</italic></td><td align="left" rowspan="1" colspan="1">AT2004187</td><td align="left" rowspan="1" colspan="1">UPS</td><td align="left" rowspan="1" colspan="1">Sweden, Jmtl</td><td align="left" rowspan="1" colspan="1">UDB002163</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. multiformis</italic></td><td align="left" rowspan="1" colspan="1">PK4471</td><td align="left" rowspan="1" colspan="1">UBC</td><td align="left" rowspan="1" colspan="1">Canada, BC</td><td align="left" rowspan="1" colspan="1">FJ039634</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. multiformis</italic></td><td align="left" rowspan="1" colspan="1">F16414</td><td align="left" rowspan="1" colspan="1">UBC</td><td align="left" rowspan="1" colspan="1">Canada, BC</td><td align="left" rowspan="1" colspan="1">FJ039635</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. myrtilliphilus</italic> Kytöv., Liimat., Niskanen & Brandrud sp. nov.</td><td align="left" rowspan="1" colspan="1">O-125949</td><td align="left" rowspan="1" colspan="1">O</td><td align="left" rowspan="1" colspan="1">Norway, Oppl, Ringebu</td><td align="left" rowspan="1" colspan="1">AY669518</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. neotriumphans</italic> Bidaud, Moënne-Locc. & Reumaux 2000</td><td align="left" rowspan="1" colspan="1">TN05-232</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Norway, Hord, Ulvik</td><td align="left" rowspan="1" colspan="1">KF732607</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. neotriumphans</italic></td><td align="left" rowspan="1" colspan="1">CFP475</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Ång, Säbrå</td><td align="left" rowspan="1" colspan="1">KF732608</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. norrlandicus</italic></td><td align="left" rowspan="1" colspan="1">IK99-711</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, PK, Juuka</td><td align="left" rowspan="1" colspan="1">KF732627</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. olivaceodionysae</italic> A. Ortega, Vila & Fdez.-Brime</td><td align="left" rowspan="1" colspan="1">TN06-311</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, SF, A, Jomala</td><td align="left" rowspan="1" colspan="1">KF732480</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. olivaceodionysae</italic></td><td align="left" rowspan="1" colspan="1">IK94-1899</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">France, Ain</td><td align="left" rowspan="1" colspan="1">KF732481</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. olivaceodionysae</italic></td><td align="left" rowspan="1" colspan="1">TN06-306</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, A, Finström</td><td align="left" rowspan="1" colspan="1">KF732482</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. olivaceodionysae</italic></td><td align="left" rowspan="1" colspan="1">IK11-013</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Sweden, Gtl</td><td align="left" rowspan="1" colspan="1">KF732483</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. olivaceodionysae</italic></td><td align="left" rowspan="1" colspan="1">MK2540</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, A, Lemland</td><td align="left" rowspan="1" colspan="1">KF732484</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. olivaceodionysae</italic></td><td align="left" rowspan="1" colspan="1">IK12-001</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Estonia, Hiiumaa, Raplamaa</td><td align="left" rowspan="1" colspan="1">KF732485</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. olivaceodionysae</italic></td><td align="left" rowspan="1" colspan="1">IK07-1417</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Sweden, Gstr</td><td align="left" rowspan="1" colspan="1">KF732486</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. olivaceodionysae</italic></td><td align="left" rowspan="1" colspan="1">IK11-012</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Sweden, Gtl</td><td align="left" rowspan="1" colspan="1">KF732487</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. olivaceodionysae</italic></td><td align="left" rowspan="1" colspan="1">TUB011856</td><td align="left" rowspan="1" colspan="1">TUB</td><td align="left" rowspan="1" colspan="1">Germany</td><td align="left" rowspan="1" colspan="1">AY669523</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. oliveopetasatus</italic> M.M. Moser 2000</td><td align="left" rowspan="1" colspan="1">F14280</td><td align="left" rowspan="1" colspan="1">UBC</td><td align="left" rowspan="1" colspan="1">Canada, BC</td><td align="left" rowspan="1" colspan="1">FJ157042</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. olympianus</italic> A.H. Sm. 1939</td><td align="left" rowspan="1" colspan="1">IK94-1225</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, ES, Kerimäki</td><td align="left" rowspan="1" colspan="1">KF732553</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. ophiopus</italic> Peck 1878</td><td align="left" rowspan="1" colspan="1">IK01-050</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, U, Helsinki</td><td align="left" rowspan="1" colspan="1">KF732609</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. ophiopus</italic></td><td align="left" rowspan="1" colspan="1">AT2004256</td><td align="left" rowspan="1" colspan="1">UPS</td><td align="left" rowspan="1" colspan="1">Sweden, Upl, Norrtälje</td><td align="left" rowspan="1" colspan="1">UDB000729</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. oregonensis</italic> A.H. Sm. 1939</td><td align="left" rowspan="1" colspan="1">F15822</td><td align="left" rowspan="1" colspan="1">UBC</td><td align="left" rowspan="1" colspan="1">Canada, BC</td><td align="left" rowspan="1" colspan="1">FJ157035</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. orichalceus</italic> var. <italic>xanthocephalus</italic> A.H. Sm. 1944</td><td align="left" rowspan="1" colspan="1">CFP769</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Dlr, Rättvik</td><td align="left" rowspan="1" colspan="1">KF732543</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. orichalceus </italic>var. <italic>xanthocephalus</italic></td><td align="left" rowspan="1" colspan="1">TN04-874</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, V, Lohja</td><td align="left" rowspan="1" colspan="1">KF732544</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. orichalceus </italic> var. <italic> xanthocephalus</italic></td><td align="left" rowspan="1" colspan="1">IK08-1482</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, Kn, Paltamo</td><td align="left" rowspan="1" colspan="1">KF732545</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. palazonianus</italic> Vila, A. Ortega & Fdez.-Brime</td><td align="left" rowspan="1" colspan="1">TN04-1106 (H7017897)</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Italy, Sardinia, Nuoro, Gavoi, Lago di Gusana</td><td align="left" rowspan="1" colspan="1">KF732531</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. palazonianus</italic></td><td align="left" rowspan="1" colspan="1">clone PS01-02</td><td rowspan="1" colspan="1"></td><td align="left" rowspan="1" colspan="1">Spain</td><td align="left" rowspan="1" colspan="1">FJ946938</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pallidirimosus </italic>Kytöv., Liimat. & Niskanen sp. nov.</td><td align="left" rowspan="1" colspan="1">IK07-692</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, PeP, Tervola</td><td align="left" rowspan="1" colspan="1">KF732579</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pallidirimosus </italic></td><td align="left" rowspan="1" colspan="1">TN04-470</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, PeP, Rovaniemi</td><td align="left" rowspan="1" colspan="1">KF732580</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pallidirimosus </italic></td><td align="left" rowspan="1" colspan="1">IK92-966</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, SoL, Sodankylä</td><td align="left" rowspan="1" colspan="1">KF732581</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pallidirimosus </italic></td><td align="left" rowspan="1" colspan="1">IK98-711</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Norway, Troms, Storfjord</td><td align="left" rowspan="1" colspan="1">KF732582</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pallidirimosus </italic></td><td align="left" rowspan="1" colspan="1">19990590</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">Russia, Sakha</td><td align="left" rowspan="1" colspan="1">UDB001073</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pallidirimosus </italic></td><td align="left" rowspan="1" colspan="1">clone 8-73M8</td><td rowspan="1" colspan="1"></td><td align="left" rowspan="1" colspan="1">USA, OR</td><td align="left" rowspan="1" colspan="1">JQ393042</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pansa</italic> (Fr.) Sacc. 1887</td><td align="left" rowspan="1" colspan="1">IK97-1914</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, ES, Mäntyharju</td><td align="left" rowspan="1" colspan="1">KF732521</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pansa</italic></td><td rowspan="1" colspan="1"></td><td rowspan="1" colspan="1"></td><td align="left" rowspan="1" colspan="1">UK</td><td align="left" rowspan="1" colspan="1">AM084701</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pansa</italic></td><td align="left" rowspan="1" colspan="1">UP21</td><td rowspan="1" colspan="1"></td><td align="left" rowspan="1" colspan="1">Sweden</td><td align="left" rowspan="1" colspan="1">DQ179120</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. papulosus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">TN06-319</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, A, Jomala</td><td align="left" rowspan="1" colspan="1">KF732629</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. papulosus</italic></td><td align="left" rowspan="1" colspan="1">IK90-1822</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, V, Kemiö</td><td align="left" rowspan="1" colspan="1">KF732630</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. parafulmineus</italic> Rob. Henry 1993</td><td align="left" rowspan="1" colspan="1">Arangu-Cort-03101201</td><td rowspan="1" colspan="1"></td><td align="left" rowspan="1" colspan="1">Spain, Roncal Navarra</td><td align="left" rowspan="1" colspan="1">EF014269</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pardinus</italic> Reumaux 1995</td><td align="left" rowspan="1" colspan="1">RH70356</td><td align="left" rowspan="1" colspan="1">PC</td><td align="left" rowspan="1" colspan="1">France</td><td align="left" rowspan="1" colspan="1">KF732461</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. patibilis</italic> Brandrud & Melot 1983</td><td align="left" rowspan="1" colspan="1">IK97-086</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, V, Karkkila</td><td align="left" rowspan="1" colspan="1">KF732631</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. patibilis</italic></td><td align="left" rowspan="1" colspan="1">IK97-087, H6032748</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, V, Karkkila</td><td align="left" rowspan="1" colspan="1">KF732632</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. patrickensis </italic>(M.M. Moser) Niskanen, Liimat., Kytöv., Bojantchev & Ammirati comb. nov.*</td><td align="left" rowspan="1" colspan="1">DBB39406</td><td align="left" rowspan="1" colspan="1">UC</td><td align="left" rowspan="1" colspan="1">USA, CA, Humboldt</td><td align="left" rowspan="1" colspan="1">KF732532</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. patrickensis* </italic></td><td align="left" rowspan="1" colspan="1">IK95-1400, H7019584</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Sweden, Mpd, Borgsjö</td><td align="left" rowspan="1" colspan="1">KF732533</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. percomis</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">CFP1104</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">France, Ain, Brenod</td><td align="left" rowspan="1" colspan="1">KF732520</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. percomis</italic></td><td align="left" rowspan="1" colspan="1">AT2004243</td><td align="left" rowspan="1" colspan="1">UPS</td><td align="left" rowspan="1" colspan="1">Sweden, Upl</td><td align="left" rowspan="1" colspan="1">UDB000726</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. percomis</italic></td><td align="left" rowspan="1" colspan="1">TU105232</td><td align="left" rowspan="1" colspan="1">TU</td><td align="left" rowspan="1" colspan="1">Sweden, Gtl</td><td align="left" rowspan="1" colspan="1">UDB015914</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pini</italic> Brandrud 1996</td><td align="left" rowspan="1" colspan="1">IK90-2288</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, V, Parainen</td><td align="left" rowspan="1" colspan="1">KF732633</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. porphyropus</italic> (Alb. & Schwein.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">TN10-004</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Canada, QC, Riviere-á-Pierre</td><td align="left" rowspan="1" colspan="1">KF732513</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. porphyropus</italic></td><td align="left" rowspan="1" colspan="1">TN06-151</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, PK, Kitee</td><td align="left" rowspan="1" colspan="1">KF732514</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. praestans</italic> (Cordier) Gillet 1874</td><td align="left" rowspan="1" colspan="1">TAAM128528</td><td align="left" rowspan="1" colspan="1">TAAM</td><td align="left" rowspan="1" colspan="1">Estonia, Saare</td><td align="left" rowspan="1" colspan="1">UDB015946</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. praestans</italic></td><td align="left" rowspan="1" colspan="1">CFP482</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Upl, Börstil, Marieberg</td><td align="left" rowspan="1" colspan="1">KF732267</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. praestans</italic></td><td align="left" rowspan="1" colspan="1">TUB011460</td><td align="left" rowspan="1" colspan="1">TUB</td><td align="left" rowspan="1" colspan="1">Germany</td><td align="left" rowspan="1" colspan="1">AY174804</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pseudocephalixus</italic> Bidaud & Moënne-Locc. 2000</td><td align="left" rowspan="1" colspan="1">IK98-1842</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Sweden, Ög, V. Tolstad</td><td align="left" rowspan="1" colspan="1">KF732634</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pseudocephalixus</italic></td><td align="left" rowspan="1" colspan="1">TUB011444</td><td align="left" rowspan="1" colspan="1">TUB</td><td align="left" rowspan="1" colspan="1">Germany</td><td align="left" rowspan="1" colspan="1">AY174784</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. pseudonaevosus</italic> Rob. Henry 1957</td><td align="left" rowspan="1" colspan="1">CFP1175</td><td align="left" rowspan="1" colspan="1">S</td><td rowspan="1" colspan="1"></td><td align="left" rowspan="1" colspan="1">KF732635</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. purpurascens</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">IK87-1174</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, ES, Joutseno</td><td align="left" rowspan="1" colspan="1">KF732511</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. purpurascens</italic></td><td align="left" rowspan="1" colspan="1">IK09-1510</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, LK, Parikkala</td><td align="left" rowspan="1" colspan="1">KF732644</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. purpurascens</italic></td><td align="left" rowspan="1" colspan="1">TUB011401</td><td align="left" rowspan="1" colspan="1">TUB</td><td align="left" rowspan="1" colspan="1">Germany</td><td align="left" rowspan="1" colspan="1">AY174858</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. purpurascens</italic></td><td align="left" rowspan="1" colspan="1">TAAM128795</td><td align="left" rowspan="1" colspan="1">TAAM</td><td align="left" rowspan="1" colspan="1">Estonia, Lääne-Viru</td><td align="left" rowspan="1" colspan="1">UDB016117</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. rapaceoides</italic> Bidaud, G. Riousset & Riousset 2000</td><td align="left" rowspan="1" colspan="1">TUB012692</td><td align="left" rowspan="1" colspan="1">TUB</td><td align="left" rowspan="1" colspan="1">Italy, Laina</td><td align="left" rowspan="1" colspan="1">EU057049</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C.</italic> cf. <italic>rhizophorus</italic> Bidaud & Cons. 2012</td><td align="left" rowspan="1" colspan="1">IK95-1973</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Germany, Baden-Württemberg, Freudenstadt</td><td align="left" rowspan="1" colspan="1">KF732569</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. </italic>cf. <italic>rhizophorus</italic></td><td align="left" rowspan="1" colspan="1">IK98-2451</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Sweden, Öl, Borgholm</td><td align="left" rowspan="1" colspan="1">KF732570</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. </italic>cf. <italic>rhizophorus</italic></td><td align="left" rowspan="1" colspan="1">TUB011860</td><td align="left" rowspan="1" colspan="1">TUB</td><td align="left" rowspan="1" colspan="1">Germany</td><td align="left" rowspan="1" colspan="1">AY669527</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. </italic>cf. <italic>rhizophorus</italic></td><td align="left" rowspan="1" colspan="1">JV01-574</td><td align="left" rowspan="1" colspan="1">C</td><td align="left" rowspan="1" colspan="1">Denmark, Jylland</td><td align="left" rowspan="1" colspan="1">DQ083813</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. rosargutus</italic> Chevassut & Rob. Henry 1978</td><td align="left" rowspan="1" colspan="1">IK96-1279</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Germany, Baden-Württemberg Freiburg, Biederbach</td><td align="left" rowspan="1" colspan="1">KF732636</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. rosargutus</italic></td><td align="left" rowspan="1" colspan="1">IK07-242</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, OP, Kiiminki</td><td align="left" rowspan="1" colspan="1">KF732637</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. rosargutus</italic></td><td align="left" rowspan="1" colspan="1">IK08-565</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, Ks, Taivalkoski</td><td align="left" rowspan="1" colspan="1">KF732638</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. rufoallutus</italic> Rob. Henry ex Bidaud & Reumaux 2006</td><td align="left" rowspan="1" colspan="1">AV010997</td><td rowspan="1" colspan="1"></td><td align="left" rowspan="1" colspan="1">Sweden, Jmt (Borgsjö Congress)</td><td align="left" rowspan="1" colspan="1">KF732639</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. rufoallutus</italic></td><td align="left" rowspan="1" colspan="1">DBB00242</td><td rowspan="1" colspan="1"></td><td align="left" rowspan="1" colspan="1">USA, OR</td><td align="left" rowspan="1" colspan="1">JF750423</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. rufolatus </italic>Moënne-Locc. 1996</td><td align="left" rowspan="1" colspan="1">CFP1112</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">France, Ain, Brenod</td><td align="left" rowspan="1" colspan="1">KF732640</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. russus</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">CFP923</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Ång, Säbrå</td><td align="left" rowspan="1" colspan="1">KF732519</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. sannio</italic> M.M. Moser 1999</td><td align="left" rowspan="1" colspan="1">IK98-891</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Sweden, Nb, Pajala</td><td align="left" rowspan="1" colspan="1">KF732536</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. sannio </italic></td><td align="left" rowspan="1" colspan="1">IK88-1160</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, KiL, Kittilä</td><td align="left" rowspan="1" colspan="1">KF732537</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. saxamontanus</italic> Fogel 1995</td><td align="left" rowspan="1" colspan="1">MTS-97-166-11</td><td align="left" rowspan="1" colspan="1">WTU</td><td align="left" rowspan="1" colspan="1">USA, WA, Kittitas</td><td align="left" rowspan="1" colspan="1">EU057026</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. scaurocaninus </italic>Chevassut & Rob. Henry 1982</td><td align="left" rowspan="1" colspan="1">HS031095</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Germany, Baden-Württemberg Karlsruhe</td><td align="left" rowspan="1" colspan="1">KF732641</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. scaurus</italic> (Fr.: Fr.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">TN10-053</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Canada, QC</td><td align="left" rowspan="1" colspan="1">KF732509</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. scaurus</italic></td><td align="left" rowspan="1" colspan="1">TN03-1698</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Slovakia, Liptovská kotlina basin, Važec</td><td align="left" rowspan="1" colspan="1">KF732510</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. scaurus</italic></td><td align="left" rowspan="1" colspan="1">19940243</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">Sweden, Femsjö</td><td align="left" rowspan="1" colspan="1">UDB001068</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. scaurus</italic></td><td align="left" rowspan="1" colspan="1">156946</td><td align="left" rowspan="1" colspan="1">TRTC</td><td align="left" rowspan="1" colspan="1">Canada, ON</td><td align="left" rowspan="1" colspan="1">JN021010</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. scaurus</italic></td><td align="left" rowspan="1" colspan="1">19980175</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">Sweden</td><td align="left" rowspan="1" colspan="1">AF325562</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. scaurus</italic></td><td align="left" rowspan="1" colspan="1">19960092</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">Italy, Sudtirol</td><td align="left" rowspan="1" colspan="1">UDB001069</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. scaurus</italic></td><td align="left" rowspan="1" colspan="1">AF-D35</td><td rowspan="1" colspan="1"></td><td align="left" rowspan="1" colspan="1">Great Britain, Scotland, Banffshire</td><td align="left" rowspan="1" colspan="1">UDB002441</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. serarius</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">TN04-923</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, U, Kirkkonummi</td><td align="left" rowspan="1" colspan="1">KF732558</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. sobrius</italic> P. Karst. 1890</td><td align="left" rowspan="1" colspan="1">IK04-045</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, U, Helsinki</td><td align="left" rowspan="1" colspan="1">KF732559</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>Cortinarius</italic> sp.</td><td align="left" rowspan="1" colspan="1">IK03-005</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Sweden</td><td align="left" rowspan="1" colspan="1">KF732525</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>Cortinarius </italic>sp.</td><td align="left" rowspan="1" colspan="1">IK03-004</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Sweden, Öl, Nötbrunnskärret</td><td align="left" rowspan="1" colspan="1">KF732526</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>Cortinarius </italic>sp.<italic>* </italic></td><td align="left" rowspan="1" colspan="1">IK11-010</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Norway, Busk, Röyken</td><td align="left" rowspan="1" colspan="1">KF732527</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. spectabilis </italic>M.M. Moser 1952 s. auct.</td><td align="left" rowspan="1" colspan="1">TEB594-04</td><td align="left" rowspan="1" colspan="1">O</td><td align="left" rowspan="1" colspan="1">Norway, Oppl</td><td align="left" rowspan="1" colspan="1">DQ663425</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. spectabilis</italic> s. auct.</td><td align="left" rowspan="1" colspan="1">TEB595-04</td><td align="left" rowspan="1" colspan="1">O</td><td align="left" rowspan="1" colspan="1">Norway, Oppl</td><td align="left" rowspan="1" colspan="1">DQ663426</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subdecolorans</italic> M. Langl. & Reumaux 2000</td><td align="left" rowspan="1" colspan="1">IK11-011</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Norway</td><td align="left" rowspan="1" colspan="1">KF732491</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subfoetens</italic> M.M. Moser & McKnight 1995</td><td align="left" rowspan="1" colspan="1">IK08-2010</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, V, Lohja</td><td align="left" rowspan="1" colspan="1">KF732560</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subfoetens</italic></td><td align="left" rowspan="1" colspan="1">F17229OC157</td><td align="left" rowspan="1" colspan="1">UBC</td><td align="left" rowspan="1" colspan="1">Canada, BC</td><td align="left" rowspan="1" colspan="1">GQ159817</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subfraudulosus </italic>Kytöv., Liimat. & Niskanen sp. nov.</td><td align="left" rowspan="1" colspan="1">CFP481</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Upl, Börstils sn</td><td align="left" rowspan="1" colspan="1">KF732565</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subfraudulosus</italic></td><td align="left" rowspan="1" colspan="1">IK98-2245</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Sweden, Srm, Mörkö parish</td><td align="left" rowspan="1" colspan="1">KF732566</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subfraudulosus</italic></td><td align="left" rowspan="1" colspan="1">IK88-2016</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Sweden, Ög, Väversunda parish</td><td align="left" rowspan="1" colspan="1">KF732567</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subfraudulosus</italic></td><td align="left" rowspan="1" colspan="1">TU106607</td><td align="left" rowspan="1" colspan="1">TU</td><td align="left" rowspan="1" colspan="1">Estonia, Saare, Kihelkonna</td><td align="left" rowspan="1" colspan="1">UDB011261</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. sublilacinopes </italic>Bidaud, Moënne-Locc. & Reumaux 2001</td><td align="left" rowspan="1" colspan="1">TSJ2002-043</td><td align="left" rowspan="1" colspan="1">C</td><td align="left" rowspan="1" colspan="1">Czech Republic</td><td align="left" rowspan="1" colspan="1">DQ663434</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subolivascens</italic> A.H. Sm. 1944</td><td align="left" rowspan="1" colspan="1">C1-EC172</td><td align="left" rowspan="1" colspan="1">WTU</td><td align="left" rowspan="1" colspan="1">USA, WA</td><td align="left" rowspan="1" colspan="1">AY356323</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subpurpurascens</italic> (Batsch) Fr. 1838</td><td align="left" rowspan="1" colspan="1">AT2004275</td><td align="left" rowspan="1" colspan="1">UPS</td><td align="left" rowspan="1" colspan="1">Sweden, Upl</td><td align="left" rowspan="1" colspan="1">UDB000736</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subpurpurascens</italic></td><td align="left" rowspan="1" colspan="1">14615</td><td rowspan="1" colspan="1"></td><td align="left" rowspan="1" colspan="1">Italy</td><td align="left" rowspan="1" colspan="1">JF907905</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subpurpureophyllus</italic> A.H. Sm. 1939</td><td align="left" rowspan="1" colspan="1">TN04-855</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, U, Vantaa</td><td align="left" rowspan="1" colspan="1">KF732557</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subpurpureophyllus</italic></td><td align="left" rowspan="1" colspan="1">AT2005152</td><td align="left" rowspan="1" colspan="1">UPS</td><td align="left" rowspan="1" colspan="1">Sweden, Upl</td><td align="left" rowspan="1" colspan="1">UDB002241</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subpurpureophyllus</italic></td><td align="left" rowspan="1" colspan="1">19890242</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">USA, Wyoming, Yellowstone National Park</td><td align="left" rowspan="1" colspan="1">GU363492</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subrubrovelatus</italic> (Bidaud) Kytöv., Liimat., Niskanen & Dima comb. nov.</td><td align="left" rowspan="1" colspan="1">TUB011414</td><td align="left" rowspan="1" colspan="1">TUB</td><td align="left" rowspan="1" colspan="1">Germany</td><td align="left" rowspan="1" colspan="1">AY174787</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subrubrovelatus </italic></td><td align="left" rowspan="1" colspan="1">TU106663</td><td align="left" rowspan="1" colspan="1">TU</td><td align="left" rowspan="1" colspan="1">Estonia, Saare</td><td align="left" rowspan="1" colspan="1">UDB011262</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subrugulosus</italic> Bidaud & Armada 2006</td><td align="left" rowspan="1" colspan="1">IK98-2578</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Sweden, Öl, Böda</td><td align="left" rowspan="1" colspan="1">KF732463</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subtortus</italic> (Pers.: Fr.) Fr. 1838</td><td align="left" rowspan="1" colspan="1">TN05-021</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, ES, Joutsa</td><td align="left" rowspan="1" colspan="1">KF732645</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subtortus</italic></td><td align="left" rowspan="1" colspan="1">19760289</td><td align="left" rowspan="1" colspan="1">IB</td><td align="left" rowspan="1" colspan="1">Sweden, Femsjö</td><td align="left" rowspan="1" colspan="1">UDB001087</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. subtortus</italic></td><td align="left" rowspan="1" colspan="1">JMB07-08-2007-05</td><td rowspan="1" colspan="1"></td><td align="left" rowspan="1" colspan="1">Canada, BC</td><td align="left" rowspan="1" colspan="1">FJ717556</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. sulphurinus </italic>Quél. 1883 s. auct.</td><td align="left" rowspan="1" colspan="1">CFP506</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Vg, Medelplana</td><td align="left" rowspan="1" colspan="1">DQ663437</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. sulphurinus </italic> var. <italic> fageticola </italic>Brandrud 1998</td><td align="left" rowspan="1" colspan="1">CFP783</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Sk, Degeberga</td><td align="left" rowspan="1" colspan="1">DQ663439</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. superbus</italic> A.H. Sm. 1944</td><td align="left" rowspan="1" colspan="1">SMI45</td><td rowspan="1" colspan="1"></td><td align="left" rowspan="1" colspan="1">Canada, BC</td><td align="left" rowspan="1" colspan="1">FJ157114</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. talimultiformis </italic>Kytöv., Liimat., Niskanen, A.F.S. Taylor & Sesli sp. nov.</td><td align="left" rowspan="1" colspan="1">IK300866 (H6032747)</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, PH, Virrat</td><td align="left" rowspan="1" colspan="1">KF732584</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. talimultiformis </italic></td><td align="left" rowspan="1" colspan="1">SES2741</td><td rowspan="1" colspan="1"></td><td align="left" rowspan="1" colspan="1">Turkey, Trabzon, Macka</td><td align="left" rowspan="1" colspan="1">KF732585</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. talimultiformis </italic></td><td align="left" rowspan="1" colspan="1">TUB0118410</td><td align="left" rowspan="1" colspan="1">TUB</td><td align="left" rowspan="1" colspan="1">Germany</td><td align="left" rowspan="1" colspan="1">AY669532</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. talimultiformis </italic></td><td align="left" rowspan="1" colspan="1">TAAM128693</td><td align="left" rowspan="1" colspan="1">TAAM</td><td align="left" rowspan="1" colspan="1">Estonia, Tartu</td><td align="left" rowspan="1" colspan="1">UDB015959</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. tiliae</italic> Brandrud 1996</td><td align="left" rowspan="1" colspan="1">O-63407</td><td align="left" rowspan="1" colspan="1">O</td><td align="left" rowspan="1" colspan="1">Norway</td><td align="left" rowspan="1" colspan="1">AY669556</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. triumphalis</italic> Bidaud, Moënne-Locc. & Reumaux 2000</td><td align="left" rowspan="1" colspan="1">CFP781</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Sk, Degeberga</td><td align="left" rowspan="1" colspan="1">KF732642</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. turmalis</italic> Fr. 1838</td><td align="left" rowspan="1" colspan="1">IK92-1133</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, SoL, Pelkosenniemi</td><td align="left" rowspan="1" colspan="1">KF732436</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. variegatus </italic>Bres. 1884 s. auct.</td><td align="left" rowspan="1" colspan="1">CFP525</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Upl, Vattholma</td><td align="left" rowspan="1" colspan="1">KF732376</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. variegatus</italic> s. auct.</td><td align="left" rowspan="1" colspan="1">F16442</td><td align="left" rowspan="1" colspan="1">UBC</td><td align="left" rowspan="1" colspan="1">Canada, BC</td><td align="left" rowspan="1" colspan="1">FJ039663</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. variipes</italic> Rob. Henry 1977</td><td align="left" rowspan="1" colspan="1">CFP981</td><td align="left" rowspan="1" colspan="1">S</td><td align="left" rowspan="1" colspan="1">Sweden, Ång, Häggdånger</td><td align="left" rowspan="1" colspan="1">KF732472</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. variosimilis</italic> M.M. Moser & Ammirati 1999</td><td align="left" rowspan="1" colspan="1">VMS26</td><td align="left" rowspan="1" colspan="1">UBC</td><td align="left" rowspan="1" colspan="1">Canada, BC</td><td align="left" rowspan="1" colspan="1">FJ717596</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. violaceonitens</italic> (Rob. Henry) Moënne-Locc. 2009</td><td align="left" rowspan="1" colspan="1">TN06-170</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, PK, Kitee</td><td align="left" rowspan="1" colspan="1">KF732505</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. violaceonitens</italic></td><td align="left" rowspan="1" colspan="1">TN00-661</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, V, Kisko</td><td align="left" rowspan="1" colspan="1">KF732506</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. violaceorubens</italic> Moënne-Locc. & Reumaux 1990</td><td align="left" rowspan="1" colspan="1">TN07-062</td><td align="left" rowspan="1" colspan="1">H</td><td align="left" rowspan="1" colspan="1">Finland, V, Kisko</td><td align="left" rowspan="1" colspan="1">KF732504</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. violaceorubens</italic></td><td align="left" rowspan="1" colspan="1">TUB011885</td><td align="left" rowspan="1" colspan="1">TUB</td><td align="left" rowspan="1" colspan="1">Germany</td><td align="left" rowspan="1" colspan="1">AY669647</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. viridirubescens</italic> M.M. Moser & Ammirati 1997</td><td align="left" rowspan="1" colspan="1">JFA11817</td><td align="left" rowspan="1" colspan="1">WTU</td><td align="left" rowspan="1" colspan="1">USA, CA, Mendocino</td><td align="left" rowspan="1" colspan="1">EU057007</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>C. wiebeae</italic> Thiers & A.H. Sm. 1969</td><td align="left" rowspan="1" colspan="1">clone NHPY20</td><td rowspan="1" colspan="1"></td><td align="left" rowspan="1" colspan="1">USA, OR/CA</td><td align="left" rowspan="1" colspan="1">FJ440879</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>Hebeloma fastibile</italic> (Pers.) P. Kumm.</td><td align="left" rowspan="1" colspan="1">19940036</td><td align="left" rowspan="1" colspan="1">IB</td><td rowspan="1" colspan="1"></td><td align="left" rowspan="1" colspan="1">AF325643</td></tr><tr><td align="left" rowspan="1" colspan="1"><italic>H. mesophaeum</italic> (Pers.) Quél.</td><td align="left" rowspan="1" colspan="1">GLM31004</td><td rowspan="1" colspan="1"></td><td rowspan="1" colspan="1"></td><td align="left" rowspan="1" colspan="1">AF126100</td></tr></tbody></table></table-wrap><fig id="F1" orientation="portrait" position="float"><label>Fig. 1</label><caption><p>The Bayesian 50 % majority-rule consensus tree inferred from ITS regions. PP > 0.50 are indicated above branches. In blue colour and <bold>bold</bold> are new species described in this study as well as new combinations made and in orange and <bold>bold</bold> neo- and epitypes designated in this study. In pink are species described by French authors, in red species described by other European authors, in green species described by North American authors and in black other specimens included in this study.</p></caption><graphic xlink:href="per-33-098-g001a"></graphic><graphic xlink:href="per-33-098-g001b"></graphic><graphic xlink:href="per-33-098-g001c"></graphic><graphic xlink:href="per-33-098-g001d"></graphic><graphic xlink:href="per-33-098-g001e"></graphic><graphic xlink:href="per-33-098-g001f"></graphic></fig><fig id="F2" orientation="portrait" position="float"><label>Fig. 2</label><caption><p>Photo of: a. <italic>C. cremeiamarescens</italic> TN06-273; b. <italic>C. flavivelatus</italic> IK98-885; c. <italic>C. kytoevuorii</italic> TN05-158; d. <italic>C. flavipallens</italic> T. Kekki 368; e. <italic>C. boreidionysae</italic> IK97-1220; f. <italic>C. cruentipellis</italic> IK01-053; g. <italic>C. caesiophylloides</italic> TEB277-09. — Photos: a, c. Kare Liimatainen; b, e, f. I. Kytövuori; d. T. Kekki; g. T.E. Brandrud.</p></caption><graphic xlink:href="per-33-098-g002"></graphic></fig><fig id="F3" orientation="portrait" position="float"><label>Fig. 3</label><caption><p>Spores of: a. <italic>C. cremeiamarescens</italic> TN04-768; b. <italic>C. flavivelatus</italic> IK98-885; c. <italic>C. kytoevuorii</italic> TN05-158; d. <italic>C. ochribubalinus</italic> IK93-641; e. <italic>C. subfraudulosus</italic> IK11-006, in Melzer’s reagent. — Drawn by I. Kytövuori and T. Niskanen. — Scale bar = 10 μm.</p></caption><graphic xlink:href="per-33-098-g003"></graphic></fig><fig id="F4" orientation="portrait" position="float"><label>Fig. 4</label><caption><p>Spores of: a. <italic>C. flavipallens</italic> IK08-1729; b. <italic>C. boreicyanites</italic> CFP931; c. <italic>C. boreidionysae</italic> IK97-1220; d.<italic> C. cruentipellis</italic> IK98-2503; e. <italic>C. luteiaureus</italic> IK07-247a, in Melzer’s reagent. — Drawn by I. Kytövuori and T. Niskanen. — Scale bar = 10 μm.</p></caption><graphic xlink:href="per-33-098-g004"></graphic></fig><fig id="F5" orientation="portrait" position="float"><label>Fig. 5</label><caption><p>Spores of: a. <italic>C. caesiophylloides</italic> TN05-016; b. <italic>C. melleicarneus</italic> IK01-053; c. <italic>C. pallidirimosus</italic> IK95-585; d. <italic>C.talimultiformis</italic> AT2004096, in Melzer’s reagent. — Drawn by I. Kytövuori and T. Niskanen. — Scale bar = 10 μm.</p></caption><graphic xlink:href="per-33-098-g005"></graphic></fig><fig id="F6" orientation="portrait" position="float"><label>Fig. 6</label><caption><p>Photo of: a. <italic>C. melleicameus</italic> IK01-053; b. <italic>C.pallidirimosus</italic> IK95-585; c. <italic>C. tallmultiformis</italic> AT2004096; d. <italic>C.balteatialutaceus</italic> IK09-751; e. <italic>C. balteatibulbosus</italic> IK98-1624; f. <italic>C. balteaticlavatus</italic> IK95-382; g. <italic>C. brunneiaurantius</italic> JV17979; h. <italic>C. caesiocolor</italic> IK00-029. — Photos: a, b, d–f, h. I. Kytövuori; c. A. Taylor; g. J. Vauras.</p></caption><graphic xlink:href="per-33-098-g006"></graphic></fig><fig id="F7" orientation="portrait" position="float"><label>Fig. 7</label><caption><p>Spores of: a. <italic>C. balteatialutaceus</italic> IK09-751; b. <italic>C. balteatibulbosus</italic> IK98-1624; c. <italic>C. balteaticlavatus</italic> IK96-595; d. <italic>C. brunneiaurantius</italic> JV17979; e. <italic>C. caesiocolor</italic> IK00-029; f. <italic>C. myrtilliphilus</italic> IK 97-1469, in Melzer’s reagent. — Drawn by I. Kytövuori and T. Niskanen. — Scale bar = 10 μm.</p></caption><graphic xlink:href="per-33-098-g007"></graphic></fig></floats-group></pmc></record>Pour manipuler ce document sous Unix (Dilib)
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